PBDB Taxon

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Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Macronaria

Reptilia

Taxonomy

Macronaria was named by Sereno (1997). It is not extant.

It was synonymized subjectively with Camarasauromorpha by Moser et al. (2006).

It was assigned to Neosauropoda by Sereno (1997), Sereno (1998), Wilson and Sereno (1998), Sereno (1999), Sereno et al. (1999), Wilson (2002), Upchurch et al. (2004), Gallina and Apesteguía (2005), Taylor and Naish (2005), Royo-Torres et al. (2006), Sánchez-Hernández et al. (2007), Allain and Aquesbi (2008), Wilson and Upchurch (2009), Suteethorn et al. (2010), D'Emic (2012), Xing et al. (2015), Mannion et al. (2019) and Apesteguía et al. (2021).

Subtaxa

Apatosaurus minimus, Aragosaurus, Astrodon (syn. Astrodontonius), Atlasaurus, Bothriospondylus robustus, Camarasauromorpha (syn. Brachiosauria), Dashanpusaurus, Dinodocus, Dinodocus mackesoni, Galveosaurus, Titanomorpha, Titanosauriformes, Yuzhoulong

Sister taxa

Camarasauromorpha (syn. Brachiosauria), Titanomorpha, Titanosauriformes

Synonyms

Astrodon was named by Johnston (1859). Its type is Astrodon johnstoni. It is the type genus of Astrodontidae.

It was replaced with Astrodonius by Kuhn (1964); it was synonymized subjectively with Pleurocoelus by McIntosh (1990) and Dong (1990); it was considered a nomen dubium by Upchurch et al. (2004); it was considered a nomen dubium by Weishampel (2006), Mannion et al. (2013) and D'Emic (2013).

It was assigned to Sauria by Leidy (1865); to Orthopoda by Cope (1870); to Sauropoda by Zittel (1890); to Pleurocoelidae by Marsh (1896) and Marsh (1898); to Atlantosauridae by Nopcsa (1901); to Sauropoda by Huene (1908), Huene (1909), von Zittel (1911) and Lull (1911); to Astrodontidae by Lapparent (1960); to Morosauridae by Hay (1902), Lull (1911) and Charig (1967); to Brachiosaurinae by Romer (1956) and Steel (1970); to Brachyosauridae by Langston (1974); to Titanosauridae by Madsen and Miller (1979); to Camarasauridae by Hay (1930), Buscalioni and Sanz (1987) and Sanz et al. (1987); to Brachiosauridae by Swinton (1936), Tatarinov (1964), Romer (1966), Russell (1984) and Carroll (1988); and to Titanosauriformes by Carpenter and Tidwell (2005).
Dinodocus was named by Owen (1884).

It was synonymized subjectively with Titanosaurus by Nopcsa (1901); it was synonymized subjectively with Pelorosaurus by Huene (1927), Huene (1927), Romer (1956), Tatarinov (1964), Romer (1966), Steel (1970) and McIntosh (1990); it was considered a nomen dubium by Blows (1998), Upchurch et al. (2004), Naish and Martill (2007) and Mannion et al. (2013).

It was assigned to Cetiosauridae by Lydekker (1888); to Sauropoda by Zittel (1890); to Sauropoda by Marsh (1889), Woodward (1908), Huene (1909) and von Zittel (1911); and to Brachiosauridae by Nopcsa (1928).
Dinodocus mackesoni was named by Owen (1884). Its type specimen is BMNH 14695 and is a 3D body fossil. Its type locality is Hythe, Kent (BMNH), which is in an Aptian marine sandstone in the Atherfield Clay Formation of the United Kingdom.

It was recombined as Titanosaurus makesoni by Nopcsa (1901); it was recombined as Pelorosaurus mackesoni by Huene (1927), Charig (1967), Steel (1970) and McIntosh (1990); it was considered a nomen dubium by Blows (1998), Upchurch et al. (2004), Naish and Martill (2007) and Mannion et al. (2013).
Bothriospondylus robustus was named by Owen (1875). Its type specimen is BMNH R.22428 and is a 3D body fossil. Its type locality is Bradford-on-Avon, Wiltshire (BMNH), which is in a Bathonian terrestrial horizon in the Forest Marble Formation of the United Kingdom.

It was considered a nomen dubium by Upchurch et al. (2004), Naish and Martill (2007) and Mannion (2010).
Astrodontonius was named by Kuhn (1961). It is not extant.

It was synonymized objectively with Astrodon by Kuhn (1966) and Steel (1970).
Apatosaurus minimus was named by Mook (1917). Its type specimen is AMNH 675, a set of postcrania (Sacrum and partial pelvis), and it is a 3D body fossil. Its type locality is Bone Cabin Quarry (AMNH), which is in a Kimmeridgian/Tithonian coarse channel fill sandstone in the Morrison Formation of Wyoming.

It was recombined as Atlantosaurus minimus by Steel (1970); it was considered a nomen dubium by Chure et al. (2006), Mannion et al. (2013) and Tschopp et al. (2015).

Synonymy list

Year	Name and author
1859	Astrodon Johnston p. 341
1865	Astrodon Leidy p. 119
1870	Astrodon Cope p. 99
1875	Bothriospondylus robustus Owen pp. 21-22 figs. Pl. 6
1876	Bothriospondylus robustus Sauvage p. 439
1880	Bothriospondylus robustus Sauvage p. 523
1882	Bothriospondylus robustus Etheridge p. 202
1884	Dinodocus Owen p. ix
1884	Dinodocus mackesoni Owen p. ix
1888	Dinodocus Lydekker p. 136
1888	Dinodocus mackesoni Lydekker p. 136
1888	Bothriospondylus robustus Lydekker p. 171
1889	Dinodocus Marsh p. 206
1890	Bothriospondylus robustus Woodward and Sherborn p. 212
1890	Dinodocus mackesoni Woodward and Sherborn p. 226
1890	Dinodocus Zittel p. 716
1890	Astrodon Zittel p. 717
1895	Bothriospondylus robustus Lydekker p. 335
1896	Astrodon Marsh p. 242
1898	Astrodon Marsh p. 158
1900	Bothriospondylus robustus Sauvage p. 504
1900	Dinodocus mackesoni Van Den Broeck p. 101
1901	Astrodon Nopcsa p. 204
1901	Titanosaurus makesoni Nopcsa p. 207
1902	Astrodon Hay p. 484
1908	Astrodon Huene p. 348
1908	Dinodocus Woodward p. 206
1909	Astrodon Huene p. 15
1909	Dinodocus Huene p. 15
1911	Astrodon Lull p. 177
1911	Astrodon von Zittel p. 288
1911	Dinodocus von Zittel p. 288
1917	Apatosaurus minimus Mook
1927	Bothriospondylus robustus Huene p. 446
1927	Pelorosaurus mackesoni Huene p. 456
1928	Dinodocus Nopcsa p. 184
1930	Astrodon Hay p. 193
1930	Apatosaurus minimus Hay p. 195
1936	Astrodon Swinton p. 210
1956	Astrodon Romer p. 621
1960	Astrodon Lapparent p. 39
1961	Astrodontonius Kuhn
1964	Astrodon Tatarinov p. 549
1966	Astrodon Romer p. 370
1967	Astrodon Charig p. 713
1967	Pelorosaurus mackesoni Charig p. 713
1970	Astrodon Steel p. 67
1970	Bothriospondylus robustus Steel p. 67
1970	Pelorosaurus mackesoni Steel p. 70
1970	Atlantosaurus minimus Steel p. 79
1973	Bothriospondylus robustus Delair p. 4
1974	Astrodon Langston p. 82
1979	Astrodon Madsen and Miller p. 5
1984	Astrodon Russell p. 23
1987	Astrodon Buscalioni and Sanz p. 66
1987	Astrodon Sanz et al. p. 48
1988	Astrodon Carroll
1990	Bothriospondylus robustus McIntosh p. 348
1990	Pelorosaurus mackesoni McIntosh p. 348
1997	Macronaria Sereno p. 444
1998	Macronaria Sereno p. 63
1998	Macronaria Wilson and Sereno p. 49
1999	Macronaria Sereno p. 2138 fig. 2
1999	Macronaria Sereno et al. p. 1346 fig. 4
2002	Macronaria Wilson p. 240 fig. 13
2004	Macronaria Upchurch et al. p. 265
2005	Astrodon Carpenter and Tidwell p. 85
2005	Macronaria Gallina and Apesteguía p. 159 fig. 5
2005	Macronaria Taylor and Naish p. 2 fig. 1
2006	Apatosaurus minimus Mateus p. 231
2006	Macronaria Royo-Torres et al. p. 1926 fig. 3
2007	Macronaria Sánchez-Hernández et al. p. 197
2008	Macronaria Allain and Aquesbi p. 403
2009	Macronaria Wilson and Upchurch p. 229 fig. 27
2010	Macronaria Suteethorn et al.
2012	Macronaria D'Emic
2015	Macronaria Xing et al.
2019	Macronaria Mannion et al.
2021	Macronaria Apesteguía et al.

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Deuterostomia
phylum	Chordata	Haeckel 1874
subphylum	Vertebrata
superclass	Gnathostomata
—	Osteichthyes	()
subclass	Sarcopterygii	()
subclass	Dipnotetrapodomorpha	(Nelson 2006)
subclass	Tetrapodomorpha	()
—	Tetrapoda
—	Reptiliomorpha
—	Anthracosauria
subclass	Amphibiosauria	Kuhn 1967
—	Cotylosauria	()
—	Amniota
—	Sauropsida
class	Reptilia
subclass	Eureptilia	()

Rank	Name	Author
—	Romeriida
—	Diapsida	()
—	Archosauromorpha	(Huene 1946)
—	Crocopoda
—	Archosauriformes	Gauthier 1986
—	Eucrocopoda
—	Archosauria	()
informal	Avemetatarsalia
—	Ornithodira
—	Dinosauromorpha
—	Dinosauriformes
—	Dinosauria	()
—	Saurischia	()
—	Sauropoda	()
—	Gravisauria
—	Eusauropoda
—	Neosauropoda
—	Macronaria
—	Macronaria

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Unr. †Macronaria Sereno 1997

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G. †Aragosaurus Sanz et al. 1987

†Aragosaurus ischiaticus Sanz et al. 1987

G. †Atlasaurus Monbaron et al. 1999

†Atlasaurus imelakei Monbaron et al. 1999

Unr. †Camarasauromorpha Salgado et al. 1997

Fm. †Camarasauridae Cope 1877 [camarasaurid]

G. †Camarasaurus Cope 1877

†Camarasaurus grandis Marsh 1877

Invalid names: Morosaurus impar Marsh 1878 [synonym], Morosaurus robustus Marsh 1878 [synonym], Pleurocoelus montanus Marsh 1896 [synonym]

†Camarasaurus lentus Marsh 1889

Invalid names: Camarasaurus annae Ellinger 1950 [synonym], Uintasaurus douglassi Holland 1919 [synonym]

†Camarasaurus lewisi Jensen 1988

†Camarasaurus supremus Cope 1877

Invalid names: Amphicoelias latus Cope 1877 [synonym], Camarasaurus leptodirus Cope 1879 [synonym], Caulodon diversidens Cope 1877 [synonym], Caulodon leptoganus Cope 1878 [synonym]

Invalid names: Cathetosaurus Jensen 1988 [synonym], Caulodon Cope 1877 [synonym], Morosaurus Marsh 1878 [synonym], Uintasaurus Holland 1919 [synonym]

G. †Lourinhasaurus Dantas et al. 1998

†Lourinhasaurus alenquerensis Lapparent and Zbyszewski 1957

Invalid names: Helopodinae Huene 1932 [empty]

G. †Europasaurus Sander et al. 2006

†Europasaurus holgeri Sander et al. 2006

G. †Tehuelchesaurus Rich et al. 1999

†Tehuelchesaurus benitezii Rich et al. 1999

Invalid names: Brachiosauria Upchurch 1997 [synonym]

G. †Dashanpusaurus Peng et al. 2005

†Dashanpusaurus dongi Peng et al. 2005

G. †Galveosaurus Sánchez-Hernández 2005

Unr. †Titanosauriformes Salgado et al. 1997

Fm. †Brachiosauridae Riggs 1904 [brachiosaurid]

G. †Abydosaurus Chure et al. 2010

†Abydosaurus mcintoshi Chure et al. 2010

G. †Brachiosaurus Riggs 1903

Subg. †Brachiosaurus (Brachiosaurus) Paul 1988

†Brachiosaurus altithorax Riggs 1903

G. †Cedarosaurus Tidwell et al. 1999

†Cedarosaurus weiskopfae Tidwell et al. 1999

G. †Galvesaurus Barco et al. 2005 [Galve lizard]

†Galvesaurus herreroi Barco et al. 2005

Invalid names: Galveosaurus herreroi Sánchez-Hernández 2005 [objective synonym]

G. †Giraffatitan Paul 1988

†Brachiosaurus (Giraffatitan) brancai Janensch 1914

†Giraffatitan brancai Janensch 1914

Invalid names: Brachiosaurus fraasi Janensch 1914 [synonym]

G. †Lusotitan Antunes and Mateus 2003

†Lusotitan atalaiensis Lapparent and Zbyszewski 1957

G. †Sonorasaurus Ratkevich 1998

†Sonorasaurus thompsoni Ratkevich 1998

G. †Soriatitan Royo-Torres et al. 2017

†Soriatitan golmayensis Royo-Torres et al. 2017

G. †Venenosaurus Tidwell et al. 2001

†Venenosaurus dicrocei Tidwell et al. 2001

G. †Vouivria Mannion et al. 2017

†Vouivria damparisensis Mannion et al. 2017

Invalid names: Astrodontidae Huene 1932 [synonym], Bellusaurinae Dong 1990 [empty], Brachiosaurinae Huene 1927 [empty], Camarasaurinae Nopcsa 1928 [empty], Euhelopodinae Romer 1956 [empty], Klamelisaurinae Zhao 1993 [empty], Ornithopsis eucamerotus Hulke 1882 [nomen dubium], Pleurocoelidae Marsh 1888 [synonym]

G. †Duriatitan Barrett et al. 2010

†Duriatitan humerocristatus Lydekker 1888

G. †Fukuititan Azuma and Shibata 2010

†Fukuititan nipponensis Azuma and Shibata 2010

G. †Fushanosaurus Wang et al. 2019

†Fushanosaurus qitaiensis Wang et al. 2019

G. †Fusuisaurus Mo et al. 2006

†Fusuisaurus zhaoi Mo et al. 2006

G. †Liaoningotitan Zhou et al. 2018

†Liaoningotitan sinensis Zhou et al. 2018

G. †Occitanopodus Moreau et al. 2020

†Occitanopodus gandi Moreau et al. 2020

G. †Oceanotitan Mocho et al. 2019

†Oceanotitan dantasi Mocho et al. 2019

G. †Ornithopsis Seeley 1870

†Ornithopsis hulkei Seeley 1870

Invalid names: Bothriospondylus magnus Owen 1875 [objective synonym]

G. †Pelorosaurus Mantell 1850

†Pelorosaurus conybeari Melville 1849

†Pelorosaurus praecursor Sauvage 1876

Unr. †Somphospondyli Wilson and Sereno 1998

G. †Angolatitan Mateus et al. 2011

†Angolatitan adamastor Mateus et al. 2011

G. †Astrophocaudia D'Emic 2013

†Astrophocaudia slaughteri D'Emic 2013

G. †Australodocus Remes 2007

†Australodocus bohetii Remes 2007

G. †Austrosaurus Longman 1933

†Austrosaurus mckillopi Longman 1933

G. †Brontomerus Taylor et al. 2011

†Brontomerus mcintoshi Taylor et al. 2011

G. †Chubutisaurus del Corro 1975

†Chubutisaurus insignis del Corro 1975

Unr. †Diamantinasauria Poropat et al. 2021

G. †Diamantinasaurus Hocknull et al. 2009

†Diamantinasaurus matildae Hocknull et al. 2009

G. †Mansourasaurus Sallam et al. 2018

†Mansourasaurus shahinae Sallam et al. 2018

G. †Sarmientosaurus Martínez et al. 2016

†Sarmientosaurus musacchioi Martínez et al. 2016

G. †Savannasaurus Poropat et al. 2016

†Savannasaurus elliottorum Poropat et al. 2016

G. †Wintonotitan Hocknull et al. 2009

†Wintonotitan wattsi Hocknull et al. 2009

Invalid names: Australotitan cooperensis Hocknull et al. 2021 [nomen dubium]

G. †Dongbeititan Wang et al. 2007

†Dongbeititan dongi Wang et al. 2007

Unr. †Euhelopodidae Romer 1956 [euhelopodid]

G. †Daxiatitan You et al. 2008

†Daxiatitan binglingi You et al. 2008

G. †Erketu Ksepka and Norell 2006

†Erketu ellisoni Ksepka and Norell 2006

G. †Euhelopus Romer 1956

†Euhelopus zdanskyi Wiman 1929

Invalid names: Helopus Wiman 1929 [replaced]

G. †Gobititan You et al. 2003

†Gobititan shenzhouensis You et al. 2003

G. †Huabeisaurus Pang and Cheng 2000

†Huabeisaurus allocotus Pang and Cheng 2000

G. †Phuwiangosaurus Martin et al. 1994

†Phuwiangosaurus sirindhornae Martin et al. 1994

G. †Qiaowanlong You and Li 2009

†Qiaowanlong kangxii You and Li 2009

G. †Ruixinia Mo et al. 2023

†Ruixinia zhangi Mo et al. 2023

G. †Ruyangosaurus Lü et al. 2009

†Ruyangosaurus giganteus Lü et al. 2009

G. †Silutitan Wang et al. 2021

†Silutitan sinensis Wang et al. 2021

G. †Tambatitanis Saegusa and Ikeda 2014

†Tambatitanis amicitiae Saegusa and Ikeda 2014

G. †Tangvayosaurus Allain et al. 1999

†Tangvayosaurus hoffeti Allain et al. 1999

G. †Xianshanosaurus Lü et al. 2009

†Xianshanosaurus shijiagouensis Lü et al. 2009

G. †Yongjinglong Li et al. 2014

†Yongjinglong datangi Li et al. 2014

G. †Europatitan Torcida Fernández-Baldor et al. 2017

†Europatitan eastwoodi Torcida Fernández-Baldor et al. 2017

G. †Gannansaurus Lü et al. 2013

†Gannansaurus sinensis Lü et al. 2013

G. †Garumbatitan Mocho et al. 2024

†Garumbatitan morellensis Mocho et al. 2024

G. †Huanghetitan You et al. 2006

†Huanghetitan liujiaxiaensis You et al. 2006

G. †Jiangshanosaurus Tang et al. 2001

†Jiangshanosaurus lixianensis Tang et al. 2001

G. †Jiangxititan Mo et al. 2023

†Jiangxititan ganzhouensis Mo et al. 2023

G. †Ligabuesaurus Bonaparte et al. 2006

†Ligabuesaurus leanzai Bonaparte et al. 2006

G. †Liubangosaurus Mo et al. 2010

†Liubangosaurus hei Mo et al. 2010

G. †Padillasaurus Carballido et al. 2015

†Padillasaurus leivaensis Carballido et al. 2015

G. †Sauroposeidon Wedel et al. 2000

†Sauroposeidon proteles Wedel et al. 2000

Invalid names: Paluxysaurus jonesi Rose 2007 [synonym]

Invalid names: Paluxysaurus Rose 2007 [synonym]

G. †Sibirotitan Averianov et al. 2018

†Sibirotitan astrosacralis Averianov et al. 2018

G. †Sonidosaurus Xu et al. 2006

†Sonidosaurus saihangaobiensis Xu et al. 2006

G. †Tastavinsaurus Canudo et al. 2008

†Tastavinsaurus sanzi Canudo et al. 2008

G. †Triunfosaurus Carvalho et al. 2017

†Triunfosaurus leonardii Carvalho et al. 2017

G. †Yunmenglong Lü et al. 2013

†Yunmenglong ruyangensis Lü et al. 2013

Invalid names: Arkharavia Alifanov and Bolotsky 2010 [nomen dubium], Arkharavia heterocoelica Alifanov and Bolotsky 2010 [nomen dubium], Australotitan Hocknull et al. 2021 [nomen dubium], Huanghetitan ruyangensis Lü et al. 2007 [nomen dubium]

G. †Sousatitanosauripus Gomes et al. 2024

†Sousatitanosauripus robsoni Gomes et al. 2024

Unr. †Titanosauria Bonaparte and Coria 1993

G. †Abdarainurus Averianov and Lopatin 2020

†Abdarainurus barsboldi Averianov and Lopatin 2020

G. †Aegyptosaurus Stromer 1932

†Aegyptosaurus baharijensis Stromer 1932

G. †Andesaurus Calvo and Bonaparte 1991

†Andesaurus delgadoi Calvo and Bonaparte 1991

G. †Argyrosaurus Lydekker 1893

†Argyrosaurus superbus Lydekker 1893

G. †Baalsaurus Calvo and Gonzalez Riga 2019

†Baalsaurus mansillai Calvo and Gonzalez Riga 2019

Fm. †Balochisauridae Malkani 2003

Subfm. †Balochisaurinae Malkani 2010

G. †Balochisaurus Malkani 2004

G. †Qaikshaheen Malkani 2023

Subfm. †Marisaurinae Malkani 2010

G. †Marisaurus Malkani 2003

G. †Baotianmansaurus Zhang et al. 2009

†Baotianmansaurus henanensis Zhang et al. 2009

G. †Barrosasaurus Salgado and Coria 2009

†Barrosasaurus casamiquelai Salgado and Coria 2009

G. †Borealosaurus You et al. 2004

†Borealosaurus wimani You et al. 2004

G. †Choconsaurus Simón et al. 2018

†Choconsaurus baileywillisi Simón et al. 2018

Unr. †Colossosauria González Riga et al. 2019

Unr. †Aeolosaurini Franco-Rosas et al. 2004

G. †Aeolosaurus Powell 1987

G. †Baurutitan Kellner et al. 2005

Invalid names: Trigonosaurus Campos et al. 2005 [synonym]

G. †Caieiria Silva Junior et al. 2022

G. †Gondwanatitan Kellner and Azevedo 1999

G. †Panamericansaurus Calvo and Porfiri 2010

G. †Chucarosaurus Agnolin et al. 2023

†Chucarosaurus diripienda Agnolin et al. 2023

Unr. †Lognkosauria Calvo et al. 2007

G. †Austroposeidon Bandeira et al. 2016

G. †Drusilasaura Navarrete et al. 2011

G. †Futalognkosaurus Calvo et al. 2007

G. †Mendozasaurus González-Riga 2003

G. †Quetecsaurus González Riga and Ortiz David 2014

G. †Traukutitan Juárez Valieri and Calvo 2011

G. †Notocolossus González Riga et al. 2016

†Notocolossus gonzalezparejasi González Riga et al. 2016

G. †Puertasaurus Novas et al. 2005

†Puertasaurus reuili Novas et al. 2005

Unr. †Rinconsauria Calvo et al. 2007

G. †Adamantisaurus Santucci and Bertini 2006

G. †Arrudatitan Silva Junior et al. 2022

G. †Brasilotitan Machado et al. 2013

G. †Bravasaurus Hechenleitner et al. 2020

G. †Maxakalisaurus Kellner et al. 2006

G. †Muyelensaurus Calvo et al. 2007

G. †Overosaurus Coria et al. 2013

G. †Pitekunsaurus Filippi and Garrido 2008

G. †Punatitan Hechenleitner et al. 2020

G. †Rinconsaurus Calvo and González Riga 2003

G. †Uberabatitan Salgado and Carvalho 2008

G. †Dgkhansaurus Malkani 2019

†Dgkhansaurus maarri Malkani 2019

G. †Dongyangosaurus Lü et al. 2008

†Dongyangosaurus sinensis Lü et al. 2008

G. †Dzharatitanis Averianov and Sues 2021

†Dzharatitanis kingi Averianov and Sues 2021

Unr. †Eutitanosauria Sanz et al. 1999

G. †Argentinosaurus Bonaparte and Coria 1993

†Argentinosaurus huinculensis Bonaparte and Coria 1993

G. †Bonitasaura Apesteguía 2004

†Bonitasaura salgadoi Apesteguía 2004

G. †Dreadnoughtus Lacovara et al. 2014

†Dreadnoughtus schrani Lacovara et al. 2014

G. †Inawentu Filippi et al. 2024

†Inawentu oslatus Filippi et al. 2024

G. †Menucocelsior Rolando et al. 2022

†Menucocelsior arriagadai Rolando et al. 2022

G. †Patagotitan Carballido et al. 2017

†Patagotitan mayorum Carballido et al. 2017

G. †Gandititan Han et al. 2024

†Gandititan cavocaudatus Han et al. 2024

G. †Hamititan Wang et al. 2021

†Hamititan xinjiangensis Wang et al. 2021

G. †Karongasaurus Gomani 2005

†Karongasaurus gittelmani Gomani 2005

G. †Laplatasaurus Huene 1927

†Laplatasaurus araukanicus Huene 1927

Unr. †Lithostrotia Wilson and Upchurch 2003

G. †Antarctosaurus Huene 1927

†Antarctosaurus wichmannianus Huene 1927

Invalid names: Antarctosaurus brasiliensis Arid and Vizotto 1971 [nomen dubium], Antarctosaurus giganteus Huene 1927 [nomen dubium]

G. †Arackar Rubilar-Rogers et al. 2021

†Arackar licanantay Rubilar-Rogers et al. 2021

G. †Atacamatitan Kellner et al. 2011

†Atacamatitan chilensis Kellner et al. 2011

G. †Bustingorrytitan Simón and Salgado 2023

†Bustingorrytitan shiva Simón and Salgado 2023

G. †Elaltitan Mannion and Otero 2012

†Elaltitan lilloi Mannion and Otero 2012

G. †Epachthosaurus Powell 1990

†Epachthosaurus sciuttoi Powell 1990

G. †Jainosaurus Hunt et al. 1995

†Jainosaurus septentrionalis Huene and Matley 1933

G. †Kaijutitan Filippi et al. 2019

†Kaijutitan maui Filippi et al. 2019

G. †Magyarosaurus Huene 1932

†Magyarosaurus dacus Nopcsa 1915

Invalid names: Magyarosaurus hungaricus Huene 1932 [synonym], Magyarosaurus transsylvanicus Huene 1932 [synonym]

G. †Malawisaurus Jacobs et al. 1993

†Malawisaurus dixeyi Haughton 1928

Fm. †Nemegtosauridae Upchurch 1995

G. †Nemegtosaurus Nowinski 1971

G. †Quaesitosaurus Kurzanov and Bannikov 1983

G. †Normanniasaurus Le Loeuff et al. 2013

†Normanniasaurus genceyi Le Loeuff et al. 2013

G. †Nullotitan Novas et al. 2019

†Nullotitan glaciaris Novas et al. 2019

G. †Paludititan Csiki et al. 2010

†Paludititan nalatzensis Csiki et al. 2010

G. †Paralititan Smith et al. 2001

†Paralititan stromeri Smith et al. 2001

G. †Pellegrinisaurus Salgado 1996

†Pellegrinisaurus powelli Salgado 1996

Superfm. †Saltasauroidea Powell 1992

G. †Isisaurus Wilson and Upchurch 2003

Unr. †Lohuecosauria Mocho et al. 2024

G. †Rapetosaurus Curry Rogers and Forster 2001

G. †Tapuiasaurus Zaher et al. 2011

G. †Titanomachya Pérez-Moreno et al. 2024

G. †Udelartitan Soto et al. 2024

G. †Shingopana Gorscak et al. 2017

†Shingopana songwensis Gorscak et al. 2017

G. †Tengrisaurus Averianov and Skutschas 2017

†Tengrisaurus starkovi Averianov and Skutschas 2017

G. †Vahiny Curry Rogers and Wilson 2014

†Vahiny depereti Curry Rogers and Wilson 2014

G. †Volgatitan Averianov and Efimov 2018

†Volgatitan simbirskiensis Averianov and Efimov 2018

Invalid names: Antarctosaurus jaxarticus Riabinin 1938 [nomen dubium]

G. †Malakhelisaurus Malkani 2008 [Malakhel lizard]

†Malakhelisaurus mianwali Malkani 2007

Invalid names: Malasaurus Malkani 2007 [replaced]

G. †Malarguesaurus González Riga et al. 2009 [Malargüe lizard]

†Malarguesaurus florenciae González Riga et al. 2009

G. †Mnyamawamtuka Gorscak and O'Connor 2019

†Mnyamawamtuka moyowamkia Gorscak and O'Connor 2019

G. †Mongolosaurus Gilmore 1933

†Mongolosaurus haplodon Gilmore 1933

G. †Narambuenatitan Filippi et al. 2011

†Narambuenatitan palomoi Filippi et al. 2011

G. †Ninjatitan Gallina et al. 2021

†Ninjatitan zapatai Gallina et al. 2021

G. †Pashtosaurus Malkani 2014

†Pashtosaurus zhobi Malkani 2014

G. †Petrobrasaurus Filippi et al. 2011

†Petrobrasaurus puestohernandezi Filippi et al. 2011

Unr. †Poripuchia Malkani 2020

Fm. †Gspsauridae Malkani 2015

Subfm. †Gspsaurinae Malkani 2015

Subfm. †Saraikimasoominae Malkani 2015

Fm. †Pakisauridae Malkani 2003

Subfm. †Isisaurinae Malkani 2010

G. †Khanazeem Malkani 2023

Subfm. †Khetranisaurinae Malkani 2010

G. †Pakisaurus Malkani 2004

Invalid names: Pakisaurinae Malkani 2010 [empty]

G. †Pseudomegaloolithus Vianey-Liaud and Garcia 2003

†Pseudomegaloolithus atlasi Vianey-Liaud and Garcia 2003

G. †Rukwatitan Gorscak et al. 2014

†Rukwatitan bisepultus Gorscak et al. 2014

G. †Teratopodus Tomaselli et al. 2022

†Teratopodus malarguensis Tomaselli et al. 2022

G. †Tiamat Pereira et al. 2024

†Tiamat valdecii Pereira et al. 2024

G. †Titanopodus González Riga and Calvo 2009

†Titanopodus mendozensis González Riga and Calvo 2009

Fm. †Titanosauridae Lydekker 1885 [titanosaurid]

Unr. †Megaloolithidae Zhao 1979

G. †Megaloolithus Mikhailov 1991

Invalid names: Antarctosauridae Olshevsky 1978 [synonym], Antarctosaurinae Powell 1986 [empty], Hypselosaurus Matheron 1869 [nomen dubium], Titanosaurinae Lydekker 1893 [empty]

Invalid names: Andesauridae Calvo and Bonaparte 1991 [empty], Andesauroidea Salgado 2003 [empty], Argyrosaurinae Powell 1986 [invalid subgroup], Titanosaurus Lydekker 1877 [nomen dubium], Titanosaurus indicus Lydekker 1877 [nomen dubium]

G. †Titanosaurimanus Dalla Vecchia and Tarlao 2000

†Titanosaurimanus nana Dalla Vecchia and Tarlao 2000

Invalid names: Brohisaurus Malkani 2003 [nomen dubium], Brohisaurus kirthari Malkani 2003 [nomen dubium], Eucamerotus Hulke 1871 [nomen dubium], Huanghetitanidae Lü et al. 2007 [empty], Iuticosaurus Le Loeuff 1993 [nomen dubium], Iuticosaurus valdensis Huene 1929 [nomen dubium], Jiutaisaurus Wu et al. 2006 [nomen dubium], Jiutaisaurus xidiensis Wu et al. 2006 [nomen dubium], Laurasiformes Royo-Torres 2009 [empty], Macrurosaurus Seeley 1869 [nomen dubium], Macrurosaurus semnus Seeley 1869 [nomen dubium], Pleurocoelus Marsh 1888 [nomen dubium], Pukyongosaurus Dong et al. 2001 [nomen dubium], Pukyongosaurus millenniumi Dong et al. 2001 [nomen dubium], Rugocaudia Woodruff 2012 [nomen dubium], Rugocaudia cooneyi Woodruff 2012 [nomen dubium]

G. †Yuzhoulong Dai et al. 2022

†Yuzhoulong qurenensis Dai et al. 2022

Invalid names: Apatosaurus minimus Mook 1917 [nomen dubium], Astrodon Johnston 1859 [nomen dubium], Bothriospondylus robustus Owen 1875 [nomen dubium], Dinodocus Owen 1884 [nomen dubium], Dinodocus mackesoni Owen 1884 [nomen dubium], Titanomorpha Sereno 1997 [empty]

Diagnosis

No diagnoses are available