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Blattulidae

Insecta - Blattodea - Blattulidae

Synonyms
Synonymy list
YearName and author
1982Blattulidae Vishnyakova p. 68
2005Blattulidae Martins-Neto et al. p. 17
2005Blattulidae Vršanský p. 115
2007Blattulidae Martins-Neto and Gallego p. 203
2007Blattulidae Vršanský and Ansorge p. 114
2007Blattulidae Wang et al.
2008Blattulidae Vršanský p. 40
2010Araripeblattidae Martins-Neto et al. p. 10
2016Blattulidae Lee p. 437
2019Blattulidae Qiu et al. p. 282
2020Blattulidae Vršanský p. 50
2021Blattulidae Vršanský p. 23

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—Ecdysozoa
—Panarthropoda
phylumArthropodaLatreille 1829
superclassHexapodaLatreille 1825
classInsecta
—Dicondylia
—Paranotalia
RankNameAuthor
subclassPterygota()
—NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
—PolyneopteraMartynov 1938
—Holopandictyoptera
superorderDictyopteraLatreille 1829
orderBlattodea(Latreille 1810)
—Solumblattodea
superfamilyCorydioidea
familyBlattulidae
familyBlattulidae

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Fm. †Blattulidae Vishnyakova 1982
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G. †Anablatta Martins-Neto and Gallego 2007
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†Anablatta compacta Martins-Neto and Gallego 2007
G. †Argentinoblattula Martins-Neto and Gallego 2005
+
†Argentinoblattula revelata Martins-Neto and Gallego 2005
G. †Batola Vršanský 2009
+
†Batola nikolai Vršanský 2009
G. †Blattula Handlirsch 1906
+
†Blattula aberrans Vishnyakova 1982
†Blattula anuniversala Vršanský 2020
†Blattula apicifurca Lin 1986
†Blattula bacharensis Vršanský 2020
†Blattula brevicaudata Vishnyakova 1968
†Blattula choutinensis Vršanský 2008
†Blattula colorata Vishnyakova 1982
†Blattula ctinoida Lin 1986
†Blattula cubitalis Vishnyakova 1982
†Blattula curvula Ren 1995
†Blattula delicatula Ren 1995
†Blattula discors Barna 2014
†Blattula disjuncta Scudder 1886
†Blattula dubia Handlirsch 1939
†Blattula elongata Martynov 1937
†Blattula extensa Vishnyakova 1982
†Blattula flamma Vršanský 2020
†Blattula geinitzi Handlirsch 1906
†Blattula hymena Lin 1986
†Blattula iensis Vishnyakova 1982
†Blattula incompleta Giebel 1856
†Blattula intercalata Handlirsch 1939
†Blattula isotoma Vishnyakova 1982
†Blattula kellos Zhang 1986
†Blattula kiensis Martynov 1937
†Blattula lanceolata Vishnyakova 1982
†Blattula langfeldti Geinitz 1880
+
Invalid names: Blattina (Mesoblattina) dobbertinensis Geinitz 1884 [synonym], Blattina incerta Geinitz 1884 [synonym], Blattina nana Geinitz 1883 [synonym], Blattula acutipennis Handlirsch 1939 [synonym], Blattula ancilla Handlirsch 1906 [synonym], Blattula brunneri Handlirsch 1939 [synonym], Blattula debilis Handlirsch 1906 [synonym], Blattula hattorfensis Bode 1953 [synonym], Blattula pusillima Handlirsch 1906 [synonym], Blattula riparia Handlirsch 1939 [synonym], Blattula vicina Handlirsch 1939 [synonym], Chiloblattula longipennis Handlirsch 1939 [synonym], Chiloblattula simplex Handlirsch 1939 [synonym], Chiloblattula subcostalis Handlirsch 1939 [synonym], Dipluroblattina scudderi Geinitz 1888 [synonym], Ectinoblattula medialis Handlirsch 1939 [synonym], Englyptoblatta elegantula Bode 1953 [synonym], Mesoblattula ala Handlirsch 1939 [synonym], Mesoblattula dobbertiniana Handlirsch 1906 [synonym], Mesoblattula geinitziana Handlirsch 1906 [synonym], Metablattula lipomena Handlirsch 1939 [synonym], Parablattula reticulata Handlirsch 1939 [synonym], Parablattula simplicissima Handlirsch 1939 [synonym], Peloblattula oligoneura Handlirsch 1939 [synonym]
†Blattula mikro Vršanský 2020
†Blattula mini Vršanský 2020
†Blattula mirta Zhang 1986
†Blattula mongolica Vršanský 2004
†Blattula piveteaui Laurentiaux 1946
†Blattula platypa Ren 1995
†Blattula prestwichii Scudder 1886
†Blattula rudis Ren 1995
†Blattula shurabica Martynov 1937
†Blattula sibirica Vishnyakova 1985
†Blattula similis Vishnyakova 1982
†Blattula sincera Lin 1985
†Blattula tchakrasensis Laurentiaux 1946
†Blattula tuodianensis Zhang and Hong 2003
†Blattula turanica Martynov 1937
†Blattula udensis Vishnyakova 1982
†Blattula universala Vršanský 2020
†Blattula velika Vršanský 2020
†Blattula vidlickai Vršanský 2004
†Blattula vulgara Vršanský 2020
†Blattula willmotti Martin 2010
Invalid names: Chiloblattula Handlirsch 1939 [synonym], Ectinoblattula Handlirsch 1939 [synonym], Englyptoblatta Bode 1953 [synonym], Mesoblattula Handlirsch 1906 [synonym], Metablattula Handlirsch 1939 [synonym], Parablattula Handlirsch 1939 [synonym], Peloblattula Handlirsch 1939 [synonym]
G. †Blattulites Vishnyakova 1982
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†Blattulites bilineatus Vishnyakova 1982
G. †Clypeblattula Zhang et al. 2024
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†Clypeblattula panda Zhang et al. 2024
G. †Elisama Giebel 1856
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†Elisama algeriaensis Vršanský 2021
†Elisama brevis Mendes 2000
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Invalid names: Araripeblatta beckeri Mendes 2000 [synonym], Araripeblatta cesae Martins-Neto et al. 2010 [synonym], Elisama americana Vršanský 2002 [synonym]
†Elisama cuboides Wang et al. 2007
†Elisama elongata Giebel 1856
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Invalid names: Mesoblattina bucklandi Scudder 1886 [objective synonym]
†Elisama extenuata Ren 1995
†Elisama fragmentaria Vršanský 2005
†Elisama globosa Vršanský et al. 2023
†Elisama grandis Vršanský 2003
†Elisama hindwingnii Lee 2016
†Elisama incerta Vršanský 2003
†Elisama kneri Giebel 1856
†Elisama minor Giebel 1856
†Elisama parallela Vršanský 2003
†Elisama pterostigmata Vršanský 2004
†Elisama scudderi Handlirsch 1906
†Elisama tsaganica Vršanský 1999
Invalid names: Araripeblatta Mendes 2000 [synonym], Ctenoblattina Scudder 1886 [synonym]
G. †Globula Vršanský 2009
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†Globula lake Vršanský 2009
G. †Habroblattula Wang et al. 2007
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†Habroblattula drepanoides Wang et al. 2007
†Habroblattula laiyangensis Chen et al. 2019
G. †Huablattula Qiu et al. 2019
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†Huablattula hui Qiu et al. 2019
†Huablattula jiewenae Qiu et al. 2019
†Huablattula vrsanskyi Zhang et al. 2023
G. †Kridla Vršanský 2005
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†Kridla stastia Vršanský 2005
G. †Macaroblattula Wang et al. 2007
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†Macaroblattula ellipsoides Wang et al. 2007
G. †Nannoblattula Handlirsch
G. †Nula Vršanský 2008
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†Nula sis Vršanský 2008
G. †Ocelloblattula Anisyutkin and Gorochov 2008
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†Ocelloblattula margarita Koubová and Vršanský 2021
†Ocelloblattula ponomarenkoi Anisyutkin and Gorochov 2008
†Ocelloblattula striata Káčerová and Azar 2022
G. †Orbablattula Martínez-Delclòs 1993
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†Orbablattula infrequens Martínez-Delclòs 1993
G. †Polliciblattula Vršanský 2020
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†Polliciblattula analis Vršanský 2020
†Polliciblattula tatosanerata Vršanský 2020
†Polliciblattula vana Vršanský 2020
G. †Pravdupovediac Sendi et al. 2023
+
†Pravdupovediac maaloufi Sendi et al. 2023
†Pravdupovediac neklam Sendi and Vršanský 2023
G. †Pseudomantina Sendi and Vršanský 2021
+
†Pseudomantina baijuyii Liang et al. 2023
†Pseudomantina nigroalba Vršanský 2021
†Pseudomantina occisor Sendi 2021
G. †Svabula Vršanský 2005
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†Svabula vzorkovana Vršanský 2005
G. †Truhla Vršanský 2020
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†Truhla vekov Vršanský 2020
G. †Vrtula Vršanský 2008
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†Vrtula sama Vršanský 2008
G. †Xonpepetla Cifuentes-Ruiz and Vršanský 2006
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†Xonpepetla rinconensis Cifuentes-Ruiz and Vršanský 2006
Invalid names: Araripeblattidae Martins-Neto et al. 2010 [synonym]
Diagnosis
ReferenceDiagnosis
R. G. Martins-Neto et al. 2005Apart from the characteristics Triassic blattopteran insects from Argentina 719 discussed by Vischnyakova (1982), tegmen with RP and MA secondary branches restricted to the apical area, above the apex. RA and CuP distal extremity perpendicularly aligned to both costal and anal margin and MA origin after this alignment (lcm = aml < oMA ). Two-branched MA
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Kiessling 2004, Bush and Bambach 2015

Age range: base of the Carnian to the top of the Late/Upper Campanian or 237.00000 to 70.60000 Ma

Collections (116 total)


Time interval Ma Country or state Original ID and collection number
Longobardian242.0 - 235.0Argentina (Mendoza) Anablatta compacta (115104)
Carnian237.0 - 228.0Argentina (La Rioja) Argentinoblattula revelata (122987)
Rhaetian208.5 - 201.3Turkey Blattula tchakrasensis, Mesoblattula piveteaui (152783)
Rhaetian208.5 - 201.3United Kingdom (England) Insecta indet. (113961)
Hettangian201.3 - 199.3Poland (Swietokrzyskie) Blattulidae indet. (123446)
Hettangian201.3 - 199.3China (Hunan) Mesoblattula apicifurca (155338)
Sinemurian199.3 - 190.8China (Hunan) Blattula zaoshangensis (155353 155356 155358) Mesoblattula kiensis (155357)
Pliensbachian190.8 - 182.7Kyrgyzstan Blattula sp., Mesoblattula kiensis, Blattula turanica, Blattula elongata (114701)
Early/Lower Toarcian183.0 - 182.0Germany Blattina langfeldti, Blattina nana, Blattina dobbertinensis, Blattina incerta, Dipluroblattina scudderi, Blattula ancilla, Blattula geinitzi, Blattula debilis, Blattula pusillima, Parablattula reticulata, Mesoblattula ala, Blattula riparia, Blattula intercalata, Blattula brunneri, Blattula vicina, Peloblattula oligoneura, Parablattula simplicissima, Metablattula lipomena, Chiloblattula simplex, Chiloblattula subcostalis, Chiloblattula longipennis, Ectinoblattula medialis, Chiloblattula dubia, Mesoblattina dobbertinensis, Mesoblattina dobbertinensis (123987) Blattula acutipennis (122986) Blattula langfeldti, Blattula dubia (114991)
Early/Lower Toarcian183.0 - 182.0Germany (Mecklenburg-Vorpommern) Blattula langfeldti, Blattula dubia (114993)
Early/Lower Toarcian183.0 - 182.0Germany (Niedersachsen) Blattula hattorfensis, Englyptoblatta elegantula (107552) Blattula langfeldti (117795)
Falciferum183.0 - 182.0Germany Blattula langfeldti (151355)
Toarcian182.7 - 174.1Russian Federation (Irkutsk) Blattula extensa, Blattula iensis, Parablattula cubitalis (104946) Blattula similis, Parablattula isotoma (104996) Mesoblattula colorata, Blattulites bilineatus (104995)
Toarcian182.7 - 174.1Australia (Western Australia) Blattula willmotti (114239)
Toarcian182.7 - 174.1China (Guangxi) Blattula pachohymena, Blattula hymena, Mesoblattula sincera (151540)
Toarcian182.7 - 174.1Kyrgyzstan Mesoblattula shurabica (114706)
Toarcian - Aalenian182.7 - 170.3China (Hunan) Blattula ctinoida (155351)
Bajocian - Bathonian170.3 - 166.1Russian Federation (Krasnoyarsk) Blattula aberrans (104947) Chiloblattula sibirica (118010)
Bajocian - Bathonian170.3 - 166.1Mongolia (Bayan-Khongor) Blattula flamma (215956) Blattula flamma, Blattula universala, Truhla vekov, Polliciblattula analis (121709) Blattula mini (215951) Blattula velika, Blattula vulgara, Blattula mini, Blattula mikro, Blattula anuniversala, Polliciblattula vana (124340) Polliciblattula tatosanerata (124339) Truhla vekov (122313)
Bajocian - Bathonian170.3 - 166.1China (Anhui) Blattula pachohymena, Mesoblattula kiensis (106695)
Callovian - Oxfordian166.1 - 157.3China (Nei Mongol) Pseudomantina baijuyii (225253)
Callovian - Oxfordian166.1 - 157.3China (Hebei) Mesoblattula chengdeensis (154837) Mesoblattula mirta, Blattula kellos (157564) Parablattula curvula (107000)
Callovian - Oxfordian166.1 - 157.3Kazakhstan (Chimkent) Blattula brevicaudata (106101) Pseudomantina nigroalba (105426 105970 113439)
Callovian - Oxfordian166.1 - 157.3China (Liaoning) Blattula liaoningensis (110855) Ctenoblattina dignata (157657 157658)
Oxfordian163.5 - 157.3Russian Federation (Buryat) Blattula lanceolata (104949)
Oxfordian - Kimmeridgian163.5 - 152.1China (Yunnan) Mesoblattula sp., Mesoblattula tuodianensis (164858)
Oxfordian - Kimmeridgian163.5 - 152.1Russian Federation (Buryat) Parablattula udensis (104950)
Tithonian152.1 - 145.0Mongolia (Dund-Gov’ Aymag) Blattula choutinensis (117697 117698)
Tithonian152.1 - 145.0Mongolia (Bayan-Khongor) Blattula bacharensis (122322)
Tithonian152.1 - 145.0Mongolia Blattula mongolica, Blattula vidlickai, Elisama pterostigmata (104748 121393 121394 125981) Blattula vidlickai (118966 165034) Blattula vidlickai, Elisama pterostigmata (152133) Elisama pterostigmata (165035)
Late/Upper Berriasian145.0 - 140.2United Kingdom (England) Blattidae indet. (117342) Ctenoblattina arcta, Nannoblattina prestwichii, Mesoblattina bucklandi, Elisama scudderi (133535)
Middle Berriasian145.0 - 140.2United Kingdom (England) Blattidae indet. 1, Blattidae indet. 3 (104008)
Berriasian145.0 - 139.8United Kingdom (England) Rithma disjuncta (133536)
Hauterivian132.9 - 129.4Algeria Elisama algeriaensis (123148)
Early/Lower Barremian130.0 - 125.45Spain Orbablattula infrequens (104052)
Early/Lower Barremian130.0 - 125.45Lebanon Blattulidae indet. (122359) Blattulidae indet., Ocelloblattula ponomarenkoi, Pseudomantina occisor, Elisama globosa, Pravdupovediac neklam (122471) Elisama sp. (123017) Ocelloblattula striata (227088) Pravdupovediac maaloufi (123416)
Late/Upper Barremian130.0 - 125.45China (Hebei) Blattula extenuata, Blattula platypa, Blattula delicatula (107001)
Late/Upper Barremian130.0 - 125.45China (Liaoning) Elisama extenuata, Elisama cuboides, Macaroblattula ellipsoides, Habroblattula drepanoides (111710) Habroblattula drepanoides (152003)
Barremian129.4 - 125.0Russian Federation (Chita) Blattula discors (105968)
Barremian129.4 - 125.0Mongolia (Selenginskij) Elisama fragmentaria (118911) Elisama fragmentaria, Svabula vzorkovana (117954)
Barremian129.4 - 125.0Mongolia Elisama incerta (124343)
Barremian - Aptian129.4 - 113.0North Korea Habroblattula drepanoides (200272)
Aptian125.0 - 113.0China (Shandong) Clypeblattula panda (161254) Habroblattula laiyangensis (201905)
Aptian125.0 - 113.0Mongolia Ctenoblattina tsaganica, Elisama parallela (113355) Elisama grandis (155546) Elisama parallela (113358 125186 130003 155553) Elisama parallela, Elisama grandis (155547) Elisama tsaganica (121714 125194 155549) Elisama tsaganica, Elisama parallela (113359 121710 121713 129999 138924 146608 155551) Elisama tsaganica, Elisama parallela, Elisama grandis (113356) Vrtula sama (103655)
Late/Upper Aptian122.46 - 113.0China (Beijing Shi) Blattula rudis (106965)
Late/Upper Aptian122.46 - 113.0Brazil (Ceará) Araripeblatta brevis, Blattulopsis beckeri, Araripeblatta oliveirai, Araripeblatta bolzoni, Araripeblatta toledoi, Araripeblatta dornellesi, Araripeblatta simplex (123014) Araripeblatta cesae (161261) Elisama americana (122352) Elisama americana, Elisama brevis, Elisama hindwingnii, Ocelloblattula santanensis (122557)
Albian113.0 - 100.5South Korea Blattula hymena, Blattula exetenuata (184784) Blattula sp. (184785) Mesoblattula sp. (184788)
Albian113.0 - 100.5Russian Federation (Khabarovsk) Kridla stastia (118912)
Late/Upper Albian105.3 - 99.6France (Poitou-Charentes) Batola nikolai, Globula lake (99114)
Cenomanian100.5 - 93.9France (Alpes-de-Haute- Provence) Nula sis (123441)
Early/Lower Cenomanian99.6 - 93.5Myanmar Huablattula hui (111688) Huablattula sp., Huablattula hui, Huablattula jiewenae (199518) Huablattula vrsanskyi (161726) Ocelloblattula margarita (184414)
Late/Upper Santonian85.8 - 83.5Russian Federation Blattulidae indet. (122590)
Late/Upper Campanian83.5 - 70.6Mexico (Coahuila) Xonpepetla rinconensis (119500)