PBDB Taxon

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Scorpiones (scorpion)

Arachnida - Scorpiones

Taxonomy

Scorpiones was named by Latreille (1817). It is extant.

It was reranked as the class Scorpionida by Jeram (1994); it was corrected as Scorpionida by Sepkoski (2002).

It was assigned to Arthropoda by Jeram (1994); to Arachnida by Fritsch (1904) and Sepkoski (2002); and to Stomothecata by Shultz (2007).

Synonyms

Archaeoctonoidea was named by Petrunkevitch (1949). Its type is Archaeoctonus.

It was considered an invalid subgroup of Scorpiones by Dunlop et al. (2013).

It was assigned to Holosternina by Kjellesvig-Waering (1986); and to Palaeoscorpiones by Jeram (1994).
Allopalaeophonoidea was named by Kjellesvig-Waering (1986). Its type is Allopalaeophonus.

It was considered an invalid subgroup of Scorpiones by Dunlop and Garwood (2023).

It was assigned to Holosternina by Kjellesvig-Waering (1986); and to Holosternina by Dunlop et al. (2013).
Branchioscorpionina was named by Kjellesvig-Waering (1986).

It was considered an invalid subgroup of Scorpiones by Dunlop et al. (2013).

It was assigned to Scorpionida by Kjellesvig-Waering (1986).
Liassoscorpionidae was named by Kjellesvig-Waering (1986).

It was considered an invalid subgroup of Scorpiones by Dunlop et al. (2013).

It was assigned to Mesophonoidea by Kjellesvig-Waering (1986).
Palaeophonus lightbodyi was named by Kjellesvig-Waering (1954). Its type specimen is BGS 89424, a claw, and it is a compression fossil. Its type locality is Ludford Lane, Ludlow (BGS collection), which is in a Pridoli marine shale in the United Kingdom.

It was considered a nomen dubium by Dunlop and Garwood (2023).

Synonymy list

Year	Name and author
1817	Scorpiones Latreille
1904	Scorpiones Fritsch p. 63
1949	Archaeoctonoidea Petrunkevitch
1954	Palaeophonus lightbodyi Kjellesvig-Waering p. 485 fig. 1
1986	Branchioscorpionina Kjellesvig-Waering p. 38
1986	Allopalaeophonoidea Kjellesvig-Waering p. 56
1986	Archaeoctonoidea Kjellesvig-Waering p. 66
1986	Liassoscorpionidae Kjellesvig-Waering p. 90
1994	Archaeoctonoidea Jeram p. 287
1994	Scorpionida Jeram p. 287
2002	Scorpionida Sepkoski
2007	Scorpiones Shultz
2013	Allopalaeophonoidea Dunlop et al. p. 30
2013	Palaeophonus lightbodyi Dunlop et al. p. 35

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Ecdysozoa
—	Panarthropoda
phylum	Arthropoda	Latreille 1829
—	Deuteropoda
subphylum	Chelicerata	Heymons 1901

Rank	Name	Author
—	Euchelicerata	Weygoldt and Paulus 1979
—	Prosomapoda
—	Planaterga
—	Dekatriata
—	Sclerophorata	Kamenz et al. 2011
class	Arachnida	Lamarck 1801
—	Stomothecata
order	Scorpiones	Latreille 1817
order	Scorpiones	Latreille 1817

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Or. Scorpiones Latreille 1817 [scorpion]

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Subor. †Bilobosternina Kjellesvig-Waering 1986

Superfm. †Branchioscorpionoidea Kjellesvig-Waering 1986

Fm. †Branchioscorpionidae Kjellesvig-Waering 1986

G. †Branchioscorpio Kjellesvig-Waering 1986

†Branchioscorpio richardsoni Kjellesvig-Waering 1986

Fm. †Dolichophoniidae Petrunkevitch 1953

G. †Dolichophonus Petrunkevitch 1949

†Dolichophonus loudonensis Laurie 1899

Invalid names: Dolichophonidae Petrunkevitch 1953 [empty]

G. †Brontoscorpio Kjellesvig-Waering 1972

†Brontoscorpio anglicus Kjellesvig-Waering 1972

Fm. †Chaerilobuthidae Lourenço and Beigel 2011

G. †Chaerilobuthus Lourenço and Beigel 2011

†Chaerilobuthus birmanicus Lourenço 2015

†Chaerilobuthus brandti Lourenço 2022

†Chaerilobuthus bruckschi Lourenço 2015

†Chaerilobuthus complexus Lourenço and Beigel 2011

†Chaerilobuthus gigantosternum Lourenço 2016

†Chaerilobuthus longiaculeus Lourenço 2013

†Chaerilobuthus meggeri Lourenço 2021

†Chaerilobuthus petersi Lourenço 2024

†Chaerilobuthus schwarzi Lourenço 2015

†Chaerilobuthus serratus Lourenço 2016

†Chaerilobuthus staxi Lourenço 2024

G. †Eramoscorpius Waddington et al. 2015

†Eramoscorpius brucensis Waddington et al. 2015

G. †Gondwanascorpio Gess 2013

†Gondwanascorpio emzantsiensis Gess 2013

G. †Gymnoscorpius Jeram 1994

†Gymnoscorpius mutillidigitus Jeram 1994

Subor. †Holosternina Kjellesvig-Waering 1986

Superfm. †Acanthoscorpionoidea Kjellesvig-Waering 1986

Fm. †Acanthoscorpionidae Kjellesvig-Waering 1986

G. †Acanthoscorpio Kjellesvig-Waering 1986

†Acanthoscorpio mucronatus Kjellesvig-Waering 1986

Fm. †Stenoscorpionidae Kjellesvig-Waering 1986

G. †Stenoscorpio Kjellesvig-Waering 1986

†Stenoscorpio gracilis Wills 1910

†Stenoscorpio pseudogracilis Wills 1947

Superfm. †Eoctonoidea Kjellesvig-Waering 1986

Fm. †Allobuthiscorpiidae Kjellesvig-Waering 1986

G. †Aspiscorpio Kjellesvig-Waering 1986

†Aspiscorpio eagari Kjellesvig-Waering 1986

Fm. †Anthracoscorpionidae Fritsch 1904

G. †Allobuthus Kjellesvig-Waering 1986

†Allobuthus pescei Vachon and Heyler 1985

G. †Anthracoscorpio Kušta 1889

†Anthracoscorpio dunlopi Pocock 1911

†Anthracoscorpio juvenis Kušta 1889

Fm. †Buthiscorpiidae Kjellesvig-Waering 1986

G. †Buthiscorpius Petrunkevitch 1953

†Buthiscorpius lemayi Kjellesvig-Waering 1986

Fm. †Eoctonidae Kjellesvig-Waering 1986

G. †Eoctonus Petrunkevitch 1913

†Eoctonus miniatus Petrunkevitch 1913

Fm. †Garnettiidae Dubinin 1962

G. †Garnettius Petrunkevitch 1953

†Garnettius hungerfordi Elias 1937

Superfm. †Gigantoscorpionoidea Kjellesvig-Waering 1986

Fm. †Gigantoscorpionidae Kjellesvig-Waering 1986

G. †Gigantoscorpio Stormer 1963

†Gigantoscorpio willsi Stormer 1963

G. †Petaloscorpio Kjellesvig-Waering 1986

†Petaloscorpio bureaui Kjellesvig-Waering 1986

Invalid names: Petaloscorpionidae Kjellesvig-Waering 1986 [synonym]

Superfm. †Mesophonoidea Wills 1910

Fm. †Centromachidae Petrunkevitch 1953

G. †Acanthracochaerilus Kjellesvig-Waering 1986

†Acanthracochaerilus palustris Kjellesvig-Waering 1986

G. †Anthracochaerilus Kjellesvig-Waering 1986

†Anthracochaerilus palustris Kjellesvig-Waering 1986

G. †Centromachus Thorell and Lindström 1885

†Centromachus euglyptus Peach 1883

G. †Opsieobuthus Kjellesvig-Waering 1986

†Opsieobuthus pottsvillensis Moore 1923

†Opsieobuthus tungeri Dunlop et al. 2016

G. †Phoxiscorpio Kjellesvig-Waering 1986

†Phoxiscorpio peachi Kjellesvig-Waering 1986

G. †Pulmonoscorpius Jeram 1994

†Pulmonoscorpius kirktonensis Jeram 1994

Invalid names: Anthracochaerilidae Kjellesvig-Waering 1986 [synonym], Phoxiscorpionidae Kjellesvig-Waering 1986 [synonym]

Fm. †Gallioscorpionidae Lourenço and Gall 2004

G. †Gallioscorpio Lourenço and Gall 2004

†Gallioscorpio voltzi Lourenço and Gall 2004

Fm. †Heloscorpionidae Kjellesvig-Waering 1986

G. †Heloscorpio Kjellesvig-Waering 1986

†Heloscorpio sutcliffei Woodward 1907

Fm. †Mazoniidae Petrunkevitch 1913

G. †Mazonia Meek and Worthen 1868

†Mazonia acadica Scudder 1895

†Mazonia wardingleyi Woodward 1907

†Mazonia woodiana Meek and Worthen 1868

Fm. †Mesophonidae Wills 1910

G. †Mesophonus Wills 1910

†Mesophonus perornatus Wills 1910

Invalid names: Mesophonus opisthophthalmus Wills 1947 [synonym]

†Mesophonus pulcherrimus Wills 1910

Fm. †Willsiscorpionidae Kjellesvig-Waering 1986

G. †Willsiscorpio Kjellesvig-Waering 1986

†Willsiscorpio bromsgroviensis Wills 1910

Superfm. †Palaeoscorpoidea Lehmann 1944

Fm. †Palaeoscorpionidae Lehmann 1944

G. †Palaeoscorpius Lehmann 1944

†Palaeoscorpius devonicus Lehmann 1944

Superfm. †Spongiophonoidea Kjellesvig-Waering 1986

Fm. †Praearcturidae Kjellesvig-Waering 1986

G. †Praearcturus Woodward 1871

†Praearcturus gigas Woodward 1871

Fm. †Spongiophonidae Kjellesvig-Waering 1986

G. †Spongiophonus Wills 1947

†Spongiophonus pustulosus Wills 1947

Invalid names: Stoermeroscorpionoidea Kjellesvig-Waering 1986 [invalid subgroup]

G. †Hubeiscorpio Walossek et al. 1990

†Hubeiscorpio gracilitarsus Walossek et al. 1990

G. †Liassoscorpionides Bode 1951

†Liassoscorpionides schmidti Bode 1951

Subor. †Lobosternina Pocock 1911

Superfm. †Isobuthoidea Petrunkevitch 1913

Fm. †Eobuthidae Kjellesvig-Waering 1986

G. †Eobuthus Fritsch 1904

†Eobuthus cordai Kjellesvig-Waering 1986

†Eobuthus holti Pocock 1911

†Eobuthus rakovnicensis Fritsch 1904

Fm. †Eoscorpiidae Scudder 1884

G. †Eoscorpius Meek and Worthen 1868

†Eoscorpius bornaensis Sterzel 1918

†Eoscorpius carbonarius Meek and Worthen 1868

Invalid names: Eoscorpius granulosus Petrunkevitch 1913 [synonym], Eoscorpius typicus Petrunkevitch 1913 [synonym], Palaeopisthacanthus mazonensis Petrunkevitch 1913 [synonym], Trigonoscorpio americanus Petrunkevitch 1913 [synonym]

†Eoscorpius casei Kjellesvig-Waering 1986

†Eoscorpius distinctus Petrunkevitch 1949

†Eoscorpius mucronatus Kjellesvig-Waering 1986

Invalid names: Europhthalmus longimanus Petrunkevitch 1949 [synonym]

†Eoscorpius primaevus Jordan and Gilbert 1919

†Eoscorpius pulcher Petrunkevitch 1949

†Eoscorpius sparthensis Baldwin and Sutcliffe 1904

Invalid names: Alloscorpius Petrunkevitch 1949 [synonym], Europhthalmus Petrunkevitch 1949 [synonym], Lichnophthalmus Petrunkevitch 1949 [synonym], Trigonoscorpio Petrunkevitch 1913 [synonym], Typhlopisthacanthus Petrunkevitch 1949 [synonym], Typhloscorpius Petrunkevitch 1949 [synonym]

G. †Eskiscorpio Kjellesvig-Waering 1986

†Eskiscorpio parvus Kjellesvig-Waering 1986

G. †Trachyscorpio Kjellesvig-Waering 1986

†Trachyscorpio squarrosus Kjellesvig-Waering 1986

Fm. †Isobuthidae Petrunkevitch 1913

G. †Boreoscorpio Kjellesvig-Waering 1986

†Boreoscorpio copelandi Kjellesvig-Waering 1986

G. †Bromsgroviscorpio Kjellesvig-Waering 1986

†Bromsgroviscorpio willsi Kjellesvig-Waering 1986

G. †Feistmantelia Fritsch 1904

†Feistmantelia ornata Fritsch 1904

G. †Isobuthus Fritsch 1904

†Isobuthus kralupensis Thorell and Lindström 1885

†Isobuthus nyranensis Fritsch 1904

†Isobuthus pottsvillensis Moore 1923

Fm. †Kronoscorpionidae Kjellesvig-Waering 1986

G. †Kronoscorpio Kjellesvig-Waering 1986

†Kronoscorpio danielsi Petrunkevitch 1913

Fm. †Pareobuthidae Kjellesvig-Waering 1986

G. †Pareobuthus Wills 1959

†Pareobuthus salopiensis Wills 1959

Superfm. †Loboarchaeoctonoidea Kjellesvig-Waering 1986

Fm. †Loboarchaeoctonidae Kjellesvig-Waering 1986

G. †Loboarchaeoctonus Kjellesvig-Waering 1986

†Loboarchaeoctonus squamosus Kjellesvig-Waering 1986

Fm. †Waterstoniidae Kjellesvig-Waering 1986

G. †Waterstonia Kjellesvig-Waering 1986

†Waterstonia aidriensis Kjellesvig-Waering 1986

†Waterstonia brachistodactyla Kjellesvig-Waering 1986

Superfm. †Paraisobuthoidea Kjellesvig-Waering 1986

Fm. †Paraisobuthidae Kjellesvig-Waering 1986

G. †Paraisobuthus Kjellesvig-Waering 1986

†Paraisobuthus duobicarinatus Kjellesvig-Waering 1986

†Paraisobuthus frici Kjellesvig-Waering 1986

†Paraisobuthus prantli Kjellesvig-Waering 1986

†Paraisobuthus virginiae Kjellesvig-Waering 1986

Fm. †Scoloposcorpionidae Kjellesvig-Waering 1986

G. †Benniescorpio Wills 1960

†Benniescorpio tuberculatus Peach 1883

G. †Scoloposcorpio Kjellesvig-Waering 1986

†Scoloposcorpio cramondensis Kjellesvig-Waering 1986

Fm. †Telmatoscorpionidae Kjellesvig-Waering 1986

G. †Telmatoscorpio Kjellesvig-Waering 1986

†Telmatoscorpio brevipectus Kjellesvig-Waering 1986

Invalid names: Opsieobuthidae Kjellesvig-Waering 1986 [empty]

Superfm. †Pseudobuthiscorpioidea Kjellesvig-Waering 1986

Invalid names: Pseudobuthiscorpiidae Kjellesvig-Waering 1986 [empty]

Invalid names: Anthracochaeriloidea Kjellesvig-Waering 1986 [empty], Palaeophonoidea Thorell and Lindstroem 1884 [empty]

Subor. †Meristosternina Kjellesvig-Waering 1986

Superfm. †Cyclophthalmoidea Thorell and Lindstroem 1885

Fm. †Cyclophthalmidae Thorell and Lindström 1885

G. †Cyclophthalmus Corda 1835

†Cyclophthalmus euglyptus Peach 1883

†Cyclophthalmus robustus Kjellesvig-Waering 1986

†Cyclophthalmus senior Corda 1835

†Cyclophthalmus sibiricus Novojilov and Størmer 1963

Fm. †Microlabiidae Kjellesvig-Waering 1986

G. †Microlabis Corda 1839

†Microlabis sternbergii Corda 1839

Superfm. †Palaeobuthoidea Kjellesvig-Waering 1986

Fm. †Palaeobuthidae Kjellesvig-Waering 1986

G. †Palaeobuthus Petrunkevitch 1913

†Palaeobuthus distinctus Petrunkevitch 1913

Invalid names: Mazoniscorpio mazonensis Wills 1960 [synonym]

Invalid names: Mazoniscorpio Wills 1960 [synonym]

Subor. Orthosternina Pocock 1911

Superfm. Buthoidea Koch 1837

Fm. †Archaeobuthidae Lourenço 2001

G. †Archaeobuthus Lourenço 2001

†Archaeobuthus estephani Lourenço 2001

Fm. Buthidae Koch 1837

G. Centruroides Marx 1890

Centruroides nitidus Thorell 1876

Invalid names: Centruroides beynai Schawaller 1979 [synonym]

Invalid names: Centrurus Ehrenberg 1829 [replaced]

G. †Cretaceousbuthus Lourenço 2022

†Cretaceousbuthus fraaijeorum Lourenço 2022

†Cretaceousbuthus petersi Lourenço 2023

G. Microcharmus Lourenco 1995

†Microcharmus henderickxi Lourenço 2009

G. Microtityus Kjellesvig-Waering 1966

†Microtityus ambarensis Schawaller 1982

G. †Palaeoakentrobuthus Lourenço and Weitschat 2000

†Palaeoakentrobuthus knodeli Lourenço and Weitschat 2000

G. †Palaeoananteris Lourenço and Weitschat 2001

†Palaeoananteris ribnitiodamgartensis Lourenço and Weitschat 2001

†Palaeoananteris ukrainensis Lourenço and Weitschat 2009

†Palaeoananteris wunderlichi Lourenço 2004

G. †Palaeogrosphus Lourenço 2000

†Palaeogrosphus copalensis Lourenço 1996

†Palaeogrosphus jacquesi Lourenço and Henderickx 2012

G. †Palaeoisometrus Lourenço and Weitschat 2005

†Palaeoisometrus elegans Lourenço and Weitschat 2005

G. †Palaeolychas Lourenço and Weitschat 1996

†Palaeolychas balticus Lourenço and Weitschat 1996

†Palaeolychas weitschati Lourenço 2012

G. †Palaeoprotobuthus Lourenço and Weitschat 2000

†Palaeoprotobuthus pusillus Lourenço and Weitschat 2000

G. †Palaeospinobuthus Lourenço et al. 2005

†Palaeospinobuthus cenozoicus Lourenço et al. 2005

G. †Palaeotityobuthus Lourenço and Weitschat 2000

†Palaeotityobuthus longiaculeus Lourenço and Weitschat 2000

G. Tityus Koch 1836

Subg. Tityus (Brazilotityus) Lourenco 2006

†Tityus apozonalli Riquelme et al. 2015

†Tityus geratus Santiago-Blay and Poinar 1988

Invalid names: Tityus (Brazilotityus) hartkorni Lourenço 2009 [synonym], Tityus azari Lourenço 2013 [synonym]

G. †Uintascorpio Perry 1995

†Uintascorpio halandrasorum Perry 1995

Invalid names: Microcharmidae Lourenco 1996 [synonym], Scorpio schweiggeri Holl 1829 [nomen dubium], Tityus eogenus Menge 1869 [nomen dubium]

Fm. †Palaeoburmesebuthidae Lourenço 2015

G. †Betaburmesebuthus Lourenço 2015

†Betaburmesebuthus bellus Lourenço 2016

†Betaburmesebuthus fleissneri Lourenço and Velten 2016

†Betaburmesebuthus fuscus Xuan et al. 2023

†Betaburmesebuthus joergi Lourenço and Rossi 2017

†Betaburmesebuthus knodelorum Lourenço 2021

†Betaburmesebuthus kobberti Lourenço 2015

†Betaburmesebuthus larafleissnerae Lourenço and Velten 2016

†Betaburmesebuthus muelleri Lourenço 2015

†Betaburmesebuthus pohli Lourenço and Velten 2017

†Betaburmesebuthus spinipedis Xuan et al. 2022

†Betaburmesebuthus villosus Xuan et al. 2023

Invalid names: Spinoburmesebuthus Lourenço and Velten 2017 [synonym]

G. †Palaeoburmesebuthus Lourenço 2002

†Palaeoburmesebuthus grimaldii Lourenço 2002

†Palaeoburmesebuthus ohlhoffi Lourenço 2015

G. †Paranotaburmesebuthus Lourenço 2024

†Paranotaburmesebuthus schmidti Lourenço 2024

Fm. †Protobuthidae Lourenço and Gall 2004

G. †Protobuthus Lourenço and Gall 2004

†Protobuthus elegans Lourenço and Gall 2004

†Protobuthus ziliolii Viaretti et al. 2023

Superfm. Chactoidea Pocock 1893

Fm. Chactidae Pocock 1893

G. †Araripescorpius Campos 1986

†Araripescorpius ligabuei Campos 1986

G. Chactas Gervais 1844

†Chactas pleistocenicus Lourenço and Weitschat 2005

Fm. Euscorpiidae Laurie 1896

G. †Eoeuscorpius Kühl and Lourenço 2017

†Eoeuscorpius ceratoi Kühl and Lourenço 2017

Fm. Hemiscorpiidae Pocock 1893

Invalid names: Hormuriinae Laurie 1896 [empty]

Fm. †Palaeoeuscorpiidae Lourenço 2003

Subfm. †Archaeoscorpiopinae Lourenço 2015

G. †Archaeoscorpiops Lourenço 2015

G. †Burmesescorpiops Lourenço 2016

G. †Palaeoeuscorpius Lourenço 2003

†Palaeoeuscorpius gallicus Lourenço 2003

Fm. †Protochactidae Lourenço et al. 2022

G. †Protochactas Lourenço et al. 2022

†Protochactas furreri Lourenço et al. 2022

Superfm. Chaeriloidea Pocock 1893

Fm. Chaerilidae Pocock 1893

G. Chaerilus Simon 1877

Subfm. †Electrochaerilinae Santiago-Blay et al. 2004

G. †Electrochaerilus Santiago-Blay et al. 2004

G. †Corniops Jeram 1994

†Corniops mapesii Jeram 1994

Superfm. Scorpionoidea Latreille 1802

G. †Compsoscorpius Petrunkevitch 1949

†Compsoscorpius buthiformis Pocock 1911

Invalid names: Allobuthus macrostethus Kjellesvig-Waering 1986 [synonym], Buthiscorpius major Wills 1960 [synonym], Compsoscorpius elegans Petrunkevitch 1949 [synonym], Compsoscorpius elongatus Petrunkevitch 1949 [synonym], Coseleyscorpio lanceolatus Kjellesvig-Waering 1986 [synonym], Leioscorpio pseudobuthiformis Kjellesvig-Waering 1986 [synonym], Lichnoscorpius minutus Petrunkevitch 1949 [synonym], Pseudobuthiscorpius labiosus Kjellesvig-Waering 1986 [synonym], Typhlopisthacanthus anglicus Petrunkevitch 1949 [synonym]

Invalid names: Allobuthiscorpius Kjellesvig-Waering 1986 [synonym], Coseleyscorpio Kjellesvig-Waering 1986 [synonym], Leioscorpio Kjellesvig-Waering 1986 [synonym], Lichnoscorpius Petrunkevitch 1949 [synonym], Pseudobuthiscorpius Kjellesvig-Waering 1986 [synonym]

Fm. †Palaeopisthacanthidae Kjellesvig-Waering 1986

G. †Cryptoscorpius Jeram 1994

†Cryptoscorpius americanus Jeram 1994

G. †Palaeopisthacanthus Petrunkevitch 1913

†Palaeopisthacanthus schucherti Petrunkevitch 1913

†Palaeopisthacanthus vogelandurdeni Jeram 1994

Fm. †Protoischnuridae de Carvalho and Lourenço 2001

G. †Cretaceoushormiops Lourenço 2018

†Cretaceoushormiops elegans Xuan et al. 2024

†Cretaceoushormiops staxi Lourenço 2022

G. †Cretaceousopisthacanthus Lourenço 2021

†Cretaceousopisthacanthus smeelei Lourenço 2021

G. †Protoischnurus de Carvalho and Lourenço 2001

†Protoischnurus axelrodorum de Carvalho and Lourenço 2001

Fm. Scorpionidae Latreille 1802

G. †Mioscorpio Kjellesvig-Waering 1986

†Mioscorpio zeuneri Hadzi 1931

G. Scorpio Linnaeus 1758

G. †Sinoscorpius Hong 1983

†Sinoscorpius shandongensis Hong 1983

G. †Palaeomachus Pocock 1911

†Palaeomachus anglicus Woodward 1876

Fm. †Palaeotrilineatidae Lourenço 2012

G. †Palaeotrilineatus Lourenço 2012

†Palaeotrilineatus ellenbergeri Lourenço 2012

G. †Permomatveevia Dammann 2017

†Permomatveevia perneri Dammann 2017

Fm. †Proscorpiidae Scudder 1885

G. †Archaeoctonus Pocock 1911

†Archaeoctonus glaber Peach 1883

G. †Hydroscorpius Kjellesvig-Waering 1986

†Hydroscorpius denisoni Kjellesvig-Waering 1986

G. †Proscorpius

†Proscorpius osborni Whitfield 1885

Invalid names: Archaeophonus eurypteroides Kjellesvig-Waering 1966 [synonym], Stoermeroscorpio delicatus Kjellesvig-Waering 1986 [synonym]

Invalid names: Archaeophonus Kjellesvig-Waering 1966 [synonym], Stoermeroscorpio Kjellesvig-Waering 1986 [synonym]

G. †Pseudarchaeoctonus Kjellesvig-Waering 1986

†Pseudarchaeoctonus denticulatus Kjellesvig-Waering 1986

G. †Waeringoscorpio Størmer 1970

†Labriscorpio alliedensis Leary 1980

†Waeringoscorpio hefteri Størmer 1970

†Waeringoscorpio westerwaldensis Poschmann et al. 2008

Invalid names: Archaeoctonidae Petrunkevitch 1949 [synonym], Hydroscorpionidae Kjellesvig-Waering 1986 [synonym], Labriscorpionidae Kjellesvig-Waering 1986 [synonym], Stoermeroscorpioniidae Kjellesvig-Waering 1986 [synonym], Waeringoscorpionidae Størmer 1970 [synonym]

G. †Titanoscorpio Kjellesvig-Waering 1986

†Titanoscorpio douglassi Kjellesvig-Waering 1986

G. †Wattisonia Wills 1960

†Wattisonia coseleyensis Wills 1960

Invalid names: Allopalaeophonoidea Kjellesvig-Waering 1986 [invalid subgroup], Archaeoctonoidea Petrunkevitch 1949 [invalid subgroup], Branchioscorpionina Kjellesvig-Waering 1986 [invalid subgroup], Liassoscorpionidae Kjellesvig-Waering 1986 [invalid subgroup], Mesoscorpionina Stockwell 1989 [empty], Palaeophonus lightbodyi Kjellesvig-Waering 1954 [nomen dubium]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

chitinc

Grouping:

solitaryc

Environment:

terrestrialc

Locomotion:

actively mobilec

Created:

2017-04-17 09:55:58

Modified:

2017-04-17 09:55:58

Source: c = class

Reference: Bush and Bambach 2015

Age range

Maximum range based only on fossils: base of the Telychian to the top of the Holocene or 438.50000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 433.4 Ma

Collections (187 total)

Time interval	Ma	Country or state	Original ID and collection number
Telychian	438.5 - 433.4	United Kingdom (Scotland)	Palaeophonus loudonensis (153581)
Telychian - Sheinwoodian	438.5 - 430.5	United Kingdom (South Lanarkshire)	Palaeophonus caledonicus (153592)
Sheinwoodian	433.4 - 430.5	Sweden	Palaeophonus nuncius (153548)
Wenlock	433.4 - 427.4	Canada (Ontario)	Eramoscorpius brucensis (165189 165190)
Ludfordian	425.6 - 423.0	United Kingdom (Wales)	Scorpiones indet. (153611)
Pridoli	423.0 - 419.2	USA (New York)	Palaeophonus osborni (153533) Proscorpius osborni (491 153530 153531) Proscorpius osborni, Archaeophonus eurypteroides (153532) Proscorpius osborni, Archaeophonus eurypteroides, Stoermeroscorpio delicatus (153529)
Pridoli	423.0 - 419.2	United Kingdom (Shropshire)	Palaeophonus lightbodyi (153549)
Lochkovian	419.2 - 410.8	United Kingdom (England)	Praearcturus gigas (153544)
Lochkovian	419.2 - 410.8	USA (Wyoming)	Praearcturus sp., Hydroscorpius denisoni, Acanthoscorpio mucronatus, Branchioscorpio richardsoni (153534)
Lochkovian	419.2 - 410.8	United Kingdom (Herefordshire)	Praearcturus gigas (201736)
Lochkovian	419.2 - 410.8	United Kingdom (Worcestershire)	Brontoscorpio anglicus (153545)
Pragian	410.8 - 407.6	Germany	Waeringoscorpio westerwaldensis (155760 155761)
Early/Lower Emsian	409.1 - 402.5	Germany	Palaeoscorpius devonicus (26904) Waeringoscorpio hefteri (155757 155758 155759)
Early/Lower Emsian	409.1 - 402.5	Germany (Alken)	Waeringoscorpio hefteri (27049)
Frasnian	382.7 - 372.2	China (Hubei)	Hubeiscorpio gracilitarsus (155762)
Frasnian	382.7 - 372.2	Canada (Quebec)	Petaloscorpio bureaui (153566 179176)
Famennian	372.2 - 358.9	South Africa (Eastern Cape)	Gondwanascorpio emzantsiensis (68049)
Famennian	372.2 - 358.9	United Kingdom (Somerset)	Praearcturus sp. (232123)
Ivorian	353.8 - 346.7	United Kingdom (Scotland)	Gigantoscorpio willsi (216790) Trachyscorpio squarrosus (153557)
Arundian	346.7 - 340.0	United Kingdom (Scotland)	Gigantoscorpio willsi, Pseudarchaeoctonus denticulatus (90682) Trachyscorpio sp., Anthracoscorpio juvenis, Archaeoctonus tuberculatus, Centromachus euglyptus (153558)
Arundian - Holkerian	346.7 - 340.0	United Kingdom	Eoscorpius sp. (153537)
Chadian	346.7 - 340.0	United Kingdom	Eoscorpius tuberculatus (153561)
Chadian	346.7 - 340.0	United Kingdom (Scotland)	Eoscorpius glaber, Eoscorpius sp. (153535)
Holkerian	346.7 - 340.0	Russian Federation (Novgorod)	Scorpiones indet. (156763)
Visean	346.7 - 330.9	United Kingdom (Scotland)	Centromachus euglyptus, Acanthracochaerilus palustris, Gigantoscorpio willsi, Eskiscorpio parvus (34471)
Asbian	340.0 - 336.0	United Kingdom	Eoscorpius glaber, Eoscorpius tuberculatus (153590)
Brigantian	336.0 - 330.9	Germany	Eoscorpius bornaensis (153594)
Brigantian	336.0 - 330.9	United Kingdom (Scotland)	Archaeoctonoidea indet., Pulmonoscorpius kirktonensis, Orthosterni indet. (131671) Pulmonoscorpius sp. A, Pulmonoscorpius sp. B, Pulmonoscorpius kirktonensis, Orthosterni indet. (131672) Scorpiones indet. (22725)
Pendleian	330.9 - 327.0	United Kingdom (Scotland)	Waterstonia brachistodactyla (153579)
Bashkirian	323.2 - 315.2	USA (Illinois)	Labriscorpio alliedensis (153591)
Westphalian A	318.7 - 316.9	United Kingdom (Durham)	Compsoscorpius elegans (153598)
Westphalian A	318.7 - 316.9	United Kingdom (Scotland)	Anthracoscorpio dunlopi, Garnettius sp. (34904) Eoscorpius tuberculatus (153562)
Westphalian A	318.7 - 316.9	United Kingdom (South Lanarkshire)	Scorpiones indet. (153653)
Westphalian A	318.7 - 316.9	United Kingdom (Lancashire)	Compsoscorpius elegans (153597) Compsoscorpius elegans, Gymnoscorpius mutillidigitus (153599)
Westphalian A	318.7 - 316.9	United Kingdom (England)	Anthracoscorpio buthiformis, Eoscorpius wardingleyi, Geralinura sutcliffei (34905) Compsoscorpius elegans (146853) Eoscorpius sp. (124367) Lichnophthalmus pulcher (153554) Scorpiones indet., Eobuthus holti, Eoscorpius sparthensis, Aspiscorpio eagari (127287)
Westphalian A	318.7 - 316.9	United Kingdom (Lanarkshire)	Waterstonia aidriensis (153574)
Westphalian A	318.7 - 316.9	Canada (Nova Scotia)	Eoscorpius casei (153556) Mazonia sp., Mazonia acadica (28069)
Westphalian A - Westphalian B	318.7 - 314.6	Netherlands	Eobuthus sp. (153552)
Westphalian A - Westphalian B	318.7 - 314.6	USA (Indiana)	Eobuthus pottsvillensis (153563)
Westphalian	318.7 - 307.5	United Kingdom (Nottinghamshire)	Eoscorpius anglicus (153582)
Westphalian B	316.9 - 314.6	United Kingdom (England)	Compsoscorpius elongatus, Anthracoscorpio buthiformis (125238) Europhthalmus sp. (127289) Lichnophthalmus pulcher, Europhthalmus longimanus (127286) Scorpiones indet., Eobuthus holti (127218)
Westphalian B	316.9 - 314.6	United Kingdom	Eoscorpius anglicus (153652)
Westphalian B	316.9 - 314.6	United Kingdom (Staffordshire)	Anthracoscorpio buthiformis (34906 34908) Eoscorpius distinctus (153555) Lichnoscorpius minutus, Typhlopisthacanthus anglicus, Anthracoscorpio buthiformis (34907) Palaeomachus sp. (153546) Typhloscorpius distinctus (34935) Wattisonia coseleyensis, Buthiscorpius buthiformis (153540)
Westphalian B	316.9 - 314.6	United Kingdom (Yorkshire)	Compsoscorpius elegans (153600 153604) Compsoscorpius elegans, Gymnoscorpius mutillidigitus (153601 153602 153603)
Westphalian B	316.9 - 314.6	United Kingdom (Nottinghamshire)	Scorpiones indet., Buthiscorpius major (153539)
Westphalian B	316.9 - 314.6	United Kingdom (Shropshire)	Pareobuthus salopiensis (153559)
Moscovian	315.2 - 307.0	Czech Republic (Pilsen)	Isobuthus nyranensis (128574)
Moscovian	315.2 - 307.0	Czech Republic (Central Bohemian)	Anthracoscorpio juvenis, Cyclophthalmus senior (126152)
Westphalian C	314.6 - 309.8	Czech Republic	Cyclophthalmus senior, Microlabis sternbergii (146851) Eobuthus cordai, Paraisobuthus frici, Cyclophthalmus senior (153550) Eobuthus rakovnicensis (153560)
Westphalian C	314.6 - 309.8	Germany	Aspiscorpio sp. (153593)
Westphalian C - Westphalian D	314.6 - 307.5	Germany (Lower Saxony)	Eoscorpius carbonarius (216437)
Westphalian C - Westphalian D	314.6 - 307.5	Canada (Nova Scotia)	Eoscorpius sp. (153551)
Westphalian D	309.8 - 307.5	USA (Illinois)	Anthracoscorpio sp., Eoscorpius granulosus, Palaeopisthacanthus schucherti, Eoctonus miniatus, Palaeobuthus distinctus, Paraisobuthus virginiae, Telmatoscorpio brevipectus, Eoscorpius typicus (112630) Buthiscorpius lemayi (153538) Eoctonus miniatus (151857) Eoscorpius carbonarius (132455 153553) Eoscorpius carbonarius, Mazonia woodiana (152106) Eoscorpius danielsi, Trigonoscorpio americanus (146857) Eoscorpius typicus, Palaeopisthacanthus mazonensis (21981) Kronoscorpio danielsi (122538) Mazonia woodiana, Eoctonus miniatus (122493) Mazoniscorpio mazonensis (133093) Palaeopisthacanthus schucherti (153605) Titanoscorpio douglassi (151855 153536)
Stephanian	307.5 - 300.4	France (Saône-et-Loire)	Allobuthus pescei (236920)
Kasimovian	307.0 - 303.7	USA (Kansas)	Mazonia hungerfordi (122510) Palaeopisthacanthus vogelandurdeni, Cryptoscorpius americanus, Corniops mapesii (153595)
Stephanian A	305.1 - 301.5	Russian Federation (Kemerovo)	Cyclophthalmus sibiricus (153547)
Gzhelian	303.7 - 298.9	Czech Republic	Feistmantelia ornata (146852)
Stephanian B	301.5 - 301.0	France	Allobuthus pescei (122490)
Stephanian C	301.0 - 300.4	France	Compsoscorpius sp. (177541) Eoscorpius sp. (177540)
Asselian	298.9 - 295.5	China (Nei Mongol)	Eoscorpius sp. (211361)
Asselian	298.9 - 295.5	France	Lichnophthalmus sp. (147002)
Sakmarian - Artinskian	293.52 - 279.3	Germany	Opsieobuthus tungeri (153497)
Kungurian	279.3 - 272.3	Russian Federation	Permomatveevia perneri (196229)
Wuchiapingian	259.9 - 254.17	Russian Federation (Vologda)	Scorpiones indet. (104754)
Changhsingian	254.17 - 252.17	Russian Federation (Vologda)	Scorpiones indet. (153588)
Aegean	247.2 - 242.0	France	Protobuthus elegans, Gallioscorpio voltzi (107683)
Aegean	247.2 - 242.0	France (Bas-Rhin)	Eoscorpiidae indet. (140639 206491 206495)
Aegean	247.2 - 242.0	France (Moselle)	Eoscorpiidae indet. (206489 206490)
Anisian	247.2 - 242.0	Australia (New South Wales)	Scorpiones indet. (106483)
Anisian	247.2 - 242.0	United Kingdom	Willsiscorpio bromsgroviensis, Spongiophonus pustulosus (153541)
Anisian	247.2 - 242.0	France (Grand Est)	Eoscorpiidae indet. (206487)
Anisian	247.2 - 242.0	United Kingdom (Worcestershire)	Mesophonus gracilis, Mesophonus pseudogracilis, Mesophonus perornatus, Mesophonus opisthophthalmus, Mesophonus pulcherrimus, Mesophonus bromsgroviensis, Spongiophonus pustulosus, Bromsgroviscorpio willsi (153542)
Illyrian - Fassanian	247.2 - 235.0	Italy	Protobuthus ziliolii (117255)
Fassanian	242.0 - 235.0	Switzerland	Protochactas furreri (230229)
Early/Lower Toarcian	183.0 - 182.0	Germany (Niedersachsen)	Liassoscorpionides schmidti (105002)
Early/Lower Barremian	130.0 - 125.45	Lebanon	Archaeobuthus estephani (153585)
Late/Upper Aptian	122.46 - 113.0	Brazil (Ceará)	Araripescorpius ligabuei (124526) Protoischnurus axelrodorum (122352 123013 153606 228902) Protoischnurus axelrodorum, Araripescorpius ligabuei (122557 122922)
Late/Upper Aptian	122.46 - 113.0	Brazil (Ceara)	Protoischnurus axelrodorum (34900 34901)
Late/Upper Albian	105.3 - 99.6	France (Poitou-Charentes)	Palaeoeuscorpius gallicus (99114)
Early/Lower Cenomanian	99.6 - 93.5	Myanmar (Kachin)	Electrochaerilus buckleyi (123429)
Early/Lower Cenomanian	99.6 - 93.5	Myanmar	Betaburmesebuthus bellus, Burmesescorpiops groehni, Chaerilobuthus gigantosternum, Chaerilobuthus serratus (172900) Betaburmesebuthus kobberti (196549) Betaburmesebuthus sp., Betaburmesebuthus bellus, Betaburmesebuthus kobberti, Betaburmesebuthus spinipedis, Betaburmesebuthus fleissneri, Betaburmesebuthus larafleissnerae, Betaburmesebuthus muelleri, Betaburmesebuthus villosus, Betaburmesebuthus fuscus, Chaerilus sp. 1, Chaerilus sp. 2, Cretaceoushormiops sp., Cretaceoushormiops elegans (161726) Chaerilobuthus brandti (230228) Chaerilobuthus bruckschi (196548) Chaerilobuthus complexus (153586) Chaerilobuthus meggeri (230227) Cretaceousbuthus fraaijeorum (230230) Cretaceousbuthus petersi, Chaerilobuthus petersi (230231) Cretaceoushormiops staxi, Chaerilobuthus staxi (230232) Cretaceousopisthacanthus smeelei (220288) Palaeoburmesebuthus grimaldii (111688 123020) Palaeoburmesebuthus ohlhoffi (196547) Palaeoburmesebuthus sp., Chaerilobuthus longiaculeus, Archaeoscorpiops cretacicus, Chaerilobuthus birmanicus (144466) Palaeotrilineatus ellenbergeri (153583) Paranotaburmesebuthus schmidti (234345) Spinoburmesebuthus knodelorum (230226)
Ypresian	56.0 - 47.8	Italy	Eoeuscorpius ceratoi (188969)
Bridgerian	50.3 - 46.2	USA (Colorado)	Uintascorpio halandrasi (153589)
Priabonian	38.0 - 33.9	Russian Federation (Kaliningrad)	Palaeoakentrobuthus knodeli (155731) Palaeoananteris ribnitiodamgartensis (155982) Palaeoananteris wunderlichi (124318) Palaeoisometrus elegans (155981) Palaeolychas balticus (123760) Palaeolychas weitschati, Palaeoprotobuthus pusillus (123764) Palaeospinobuthus cenozoicus (154709) Palaeotityobuthus longiaculeus (152042)
Priabonian	38.0 - 33.9	Poland	Tityus eogenus (120811)
Priabonian	38.0 - 33.9	Ukraine	Palaeoananteris ukrainensis (153587)
Rupelian	33.9 - 28.1	Germany (Baden-Württemberg)	Scorpiones indet. (68021)
Early/Lower Miocene	23.03 - 15.97	Mexico (Chiapas)	Tityus apozonalli (217223)
Early/Lower Miocene	23.03 - 15.97	Mexico	Centruroides sp. (153609)
Miocene	23.03 - 5.333	USA (California)	Eoscorpius primaevus (186018)
Burdigalian	20.44 - 15.97	China (Shandong)	Sinoscorpius shandongensis (130412)
Burdigalian - Langhian	20.44 - 13.82	Dominican Republic	Scorpiones indet., Centruroides beynai, Tityus ambarensis (122474) Tityus azari (146557) Tityus geratus (124475) Tityus hartkorni (153584)
Sarmatian	12.7 - 11.608	Germany (Wurttemberg)	Scorpio zeuneri (113752)
Holocene	0.0117 - 0.0	Madagascar (Antseranana)	Microcharmus henderickxi (153607) Palaeogrosphus jacquesi (153608)
Holocene	0.0117 - 0.0	Colombia (Santander)	Chactas pleistocenicus (155980)
Holocene	0.0117 - 0.0	Tanzania	Scorpio schweiggeri (147705)
Holocene	0.0117 - 0.0	Madagascar	Tityobuthus copalensis (38973)