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Unicalcarida

Insecta - Hymenoptera

Synonyms
Synonymy list
YearName and author
1980Orussomorpha Rasnitsyn p. 64
2002Unicalcarida Schulmeister et al.
2010Orussomorpha Rasnitsyn and Zhang p. 839
2011Unicalcarida Heraty et al.
2012Unicalcarida Krogmann and Nel p. 221

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—EcdysozoaAguinaldo et al. 1997
—Panarthropoda
phylumArthropodaGravenhorst 1843
—Deuteropoda
—Mandibulata
—PancrustaceaZrzavý and Štys 1997
—AltocrustaceaRegier et al. 2010
superclassAllotriocarida
—Athalassocarida
—LabiocaridaSchwentner et al. 2017
RankNameAuthor
superclassHexapodaLatreille 1825
—Cercofilata
classInsecta
—Dicondylia
—Paranotalia
subclassPterygota()
—NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
—Holometabola
—Hymenopterida
orderHymenoptera
—Unicalcarida
—Unicalcarida

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Unr. Unicalcarida Schulmeister et al. 2002
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Subor. Apocrita Gerstaecker 1867 [wasp]
+
Unr. Aculeata Latreille 1802
+
Superfm. Apoidea Latreille 1802 [apoid wasp]
+
Fm. †Allommationidae Rosa and Melo 2021
+
G. †Allommation Rosa and Melo 2021
Fm. Ampulicidae Shuckard 1840 [cockroach wasp]
+
G. †Cariridris Brandão and Martins-Neto 1990
Tr. †Cretampulicini Antropov 2000
Subfm. Dolichurinae Lepeletier 1845
G. †Gallosphex Schlüter 1978
G. †Longivenula Zhang 1989
Tr. †Mendampulicini Antropov 2000
Invalid names: Ampulicini Shuckard 1840 [empty]
Fm. †Angarosphecidae Rasnitsyn 1975
+
G. †Angarosphex Rasnitsyn 1975
+
Invalid names: Calobaissodes Zhang 1992 [synonym], Mataeosphex Zhang 1985 [synonym], Palaeapis Hong 1984 [synonym], Shandongodes Zhang 1985 [synonym]
G. †Cretobestiola Pulawski and Rasnitsyn 2000
+
Invalid names: Bestiola Pulawski and Rasnitsyn 1999 [replaced]
G. †Cretosphecium Pulawski and Rasnitsyn 2000
G. †Eosphecium Pulawski and Rasnitsyn 2000
G. †Ilerdosphex Rasnitsyn 2000
G. †Mesorhopalosoma Darling 1990
G. †Montsecosphex Rasnitsyn and Martínez-Delclòs 2000
G. †Paleorhopalosoma Nel et al. 2010
G. †Pompilopterus Rasnitsyn 1975
G. †Vitimosphex Rasnitsyn 1975
Invalid names: Baissodinae Rasnitsyn 1975 [synonym]
Unr. Anthophila Latreille 1804 [bee]
+
Subfm. Andreninae Latreille 1802
Fm. Apidae Latreille 1802
Fm. Colletidae Lepeletier 1841 [plasterer bee]
Fm. Halictidae Thomson 1869 [sweat bee]
Fm. Megachilidae Latreille 1802 [mason bee]
Fm. Melittidae Kawall 1857
Fm. †Paleomelittidae Engel 2001
Invalid names: Stenotritidae [empty]
Fm. Astatidae Lepeletier 1845
+
Tr. †Cretastatini Rosa and Melo 2021
Tr. Eremiaspheciini Menke 1967
Fm. †Burmasphecidae Rosa and Melo 2023
+
G. †Burmasphex Melo and Rosa 2018
G. †Callisphex Rosa and Melo 2023
G. †Decasphex Zheng et al. 2020
G. †Simplisphex Rosa and Melo 2023
Fm. †Cirrosphecidae Antropov 2000
+
Subfm. †Cirrosphecinae Antropov 2000
Subfm. †Glenocephalinae Rosa and Melo 2021
Fm. Crabronidae Latreille 1802
+
Subfm. Bembicinae Latreille 1802
Subfm. †Burmastatinae Antropov 2000
Subfm. Crabroninae Latreille 1802
Tr. †Exallopterini Rosa and Melo 2022
Subfm. Mellininae Latreille 1802
Fm. Dasypodaidae Sagemehl 1882
+
Subfm. Dasypodainae Sagemehl 1882
Fm. Heterogynaidae Nagy 1969
+
Subfm. †Ptilocosminae Rosa and Melo 2021
Fm. Pemphredonidae Dahlbom 1835 [aphid wasp]
+
Tr. †Palangini Antropov 2011
Tr. Pemphredonini Dahlbom 1835
Tr. †Rasnitsynapini Antropov 2011
Tr. Spilomenini Menke 1989
Fm. Philanthidae Latreille 1802
+
Tr. Cercerini Lepeletier de Saint Fargeau 1845
Tr. Philanthini Latreille 1802
Fm. Psenidae Costa 1858
+
G. †Eomimesa Budrys 1993
Subtr. †Helosericina Rosa and Melo 2021
Subtr. †Odontosericina Rosa and Melo 2021
Subtr. Psenina Costa 1858
G. †Tytthopsen Antropov 2014
Fm. Sphecidae Latreille 1802 [digger wasp]
+
Subfm. Ammophilinae André 1886
G. †Larrophanes Handlirsch 1907
Subfm. Sceliphrinae Ashmead 1899
Subfm. Sphecinae Latreille 1802
Invalid names: Spheginae Latreille 1802 [empty]
Fm. †Spheciellidae Rosa and Melo 2021
+
G. †Spheciellus Rosa and Melo 2021
Fm. †Temnogynidae Rosa and Melo 2024
+
G. †Rhapidogyna Rosa and Melo 2024
+
Invalid names: Rhabdogyna Rosa and Melo 2024 [replaced]
G. †Temnogyna Rosa and Melo 2024
G. †Trigampulex Antropov 2000
+
†Trigampulex pervetus Cockerell 1917
Invalid names: Ammoplanidae Evans 1959 [empty], Bembicidae Latreille 1802 [empty], Sphecoidea Latreille 1802 [synonym]
Superfm. †Bethylonymoidea Rasnitsyn 1975
+
Fm. †Bethylonymidae Rasnitsyn 1975
+
G. †Allogaster Ren 1995
G. †Bethylonymellus Rasnitsyn 1975
G. †Bethylonymus Rasnitsyn 1975
G. †Meiagaster Rasnitsyn and Ansorge 2000
Fm. †Trifionychidae Lepeco and Melo 2022
+
G. †Mirabythus Cai et al. 2012
G. †Prionaspidion Lepeco and Melo 2022
G. †Trifionyx Lepeco and Melo 2022
G. †Trifionyximus Lepeco and Melo 2022
Superfm. Chrysidoidea Latreille 1802
+
Fm. Bethylidae Forster 1856 [flat wasp]
+
Subfm. Bethylinae Halliday 1840
G. †Boreobythus Engel and Grimaldi 2007
Subfm. †Cretabythinae Brazidec et al. 2023
G. †Cursoribythus Cockx and McKellar 2016
Subfm. †Elektroepyrinae Colombo and Azevedo 2020
Subfm. Epyrinae Kieffer 1914
Subfm. †Lancepyrinae Azevedo and Azar 2012
Subfm. Mesitiinae Kieffer 1914
G. †Nadezhdabythus Zhang et al. 2020
Subfm. Pristocerinae Mocsary 1881
Subfm. †Protopristocerinae Nagy 1974
Subfm. Scleroderminae Kieffer 1914
Invalid names: Epyrinae Kieffer 1914 [invalid subgroup], Fushunochrysidae Hong 2002 [synonym]
Fm. Chrysididae Latreille 1802 [cuckoo wasp]
+
Subfm. Amiseginae Mocsary 1889
G. †Auricleptes Lucena and Melo 2018
G. †Burmasega Lucena and Melo 2018
Subfm. Chrysidinae Latreille 1802
Subfm. Cleptinae Latreille 1802
G. †Dahurochrysis Rasnitsyn 1990
G. †Ectenobythus Engel et al. 2013
G. †Necrobythus Engel et al. 2013
G. †Protochrysidis Carpenter 1986
+
Invalid names: Eochrysis Doweld 2015 [objective synonym], Protochrysis Bischoff 1916 [replaced]
G. †Sphakelobythus Engel et al. 2013
Fm. †Chrysobythidae Melo and Lucena 2019
+
G. †Aureobythus Melo and Lucena 2019
G. †Bethylochrysis Melo and Lucena 2019
G. †Chrysobythus Melo and Lucena 2019
Fm. †Chrysopsenellidae Lepeco and Melo 2023
+
G. †Chrysopsenella Lepeco and Melo 2023
Fm. Dryinidae Haliday 1833
+
Subfm. Anteoninae Kieffer 1907
Subfm. Aphelopinae Perkins 1912
Subfm. Apodryininae Olmi 1984
Subfm. †Archaeodryininae Olmi et al. 2020
Subfm. Bocchinae Richards 1939
Subfm. †Burmadryininae Olmi et al. 2014
Subfm. Dryininae Haliday 1833
Subfm. †Palaeoanteoninae Olmi and Bechly 2001
Subfm. †Protodryininae Olmi and Guglielmino 2012
Subfm. †Raptodryininae Olmi et al. 2020
Subfm. Thaumatodryininae Perkins 1905
Fm. Embolemidae Forster 1856
+
G. Ampulicomorpha Ashmead 1893
G. †Baissobius Rasnitsyn 1975
G. †Cretembolemus Olmi et al. 2014
G. †Embolemopsis Olmi et al. 2010
G. Embolemus Westwood 1833
G. †Ponomarenkoa Olmi 2010
+
Invalid names: Laberites Ponomarenko 1988 [replaced]
Invalid names: Embolemus breviscapus Brues 1933 [nomen dubium], Ponomarenkoinae Olmi 2010 [invalid subgroup]
Fm. †Plumalexiidae Brothers 2011
+
G. †Plumalexius Brothers 2011
Fm. †Radiophronidae Ortega-Blanco et al. 2010
+
G. †Microcostaphron Ortega-Blanco et al. 2010
G. †Radiophron Ortega-Blanco et al. 2010
Fm. Sclerogibbidae Ashmead 1902 [sclerogibbid wasp]
+
G. †Burmasclerogibba Perkovsky et al. 2020
G. †Chosia Fujiyama 1994
G. †Cretosclerogibba Perkovsky et al. 2020
G. †Edrossia Perkovsky et al. 2020
G. †Gallosclerogibba Perrichot et al. 2020
G. Sclerogibba Riggio and De Stefani-Perez 1888
Subfm. Sclerogibbinae Ashmead 1902
Subfm. †Sclerogibbodinae Engel and Grimaldi 2006
Fm. Scolebythidae Evans 1963
+
Subfm. †Cretopsenellinae Lepeco and Melo 2023
Subfm. Pristapenesiinae Engel et al. 2013
Subfm. †Rhynchopsenellinae Lepeco and Melo 2023
Subfm. Scolebythinae Evans 1963
Fm. †Sirenobethylidae Wu et al. 2025
+
G. †Sirenobethylus Wu et al. 2025
Invalid names: Bethyloidea [synonym], Plumariidae [empty]
G. †Cretopone Dlussky 1975
+
†Cretopone magna Dlussky 1975
G. †Dolichomyrma Dlussky 1975
+
†Dolichomyrma latipes Dlussky 1975
†Dolichomyrma longiceps Dlussky 1975
Superfm. Formicoidea Latreille 1802
+
G. †Camelomecia Barden and Grimaldi 2016
+
†Camelomecia janovitzi Barden and Grimaldi 2016
G. †Camelosphecia Boudinot et al. 2020
+
†Camelosphecia fossor Boudinot et al. 2020
†Camelosphecia venator Boudinot et al. 2020
Fm. Formicidae Latreille 1802 [ant]
+
Unr. Antennoclypeata Boudinot et al. 2020
Subfm. †Armaniinae Dlussky 1983
G. †Calyptites Scudder 1877
G. †Cretomyrma Dlussky 1975
G. †Dlusskyidris Bolton 1994
+
Invalid names: Palaeomyrmex Dlussky 1975 [replaced]
Subfm. Ecitoninae Forel 1893
G. †Eoformica Cockerell 1921
G. †Fonsecahymen Martins-Neto and Mendes 2002
Subfm. †Haidomyrmecinae Bolton 2003 [hell ant]
G. †Klondikia Dlussky and Rasnitsyn 2002
G. †Kohlsimyrma Dlussky and Rasnitsyn 2002
G. †Myrmicites Förster 1891
Subfm. †Sphecomyrminae Wilson and Brown 1967
Subfm. †Zigrasimeciinae Borysenko 2017
Invalid names: Camponotus brodiei Donisthorpe 1920 [nomen dubium], Eocynipites Hong 2002 [nomen nudum], Formica gibbosa Presl 1822 [nomen dubium], Formica lucida Giebel 1856 [nomen dubium], Formica luteola Presl 1822 [nomen dubium], Formica macrognatha Presl 1822 [nomen dubium], Formica nigra Presl 1822 [nomen dubium], Formica obtecta Heer 1849 [nomen dubium], Formica obvoluta Heer 1849 [nomen dubium], Formica oculata Heer 1849 [nomen dubium], Formica parvula Presl 1822 [nomen dubium], Formica quadrata Holl 1829 [nomen dubium], Formica trigona Presl 1822 [nomen dubium], Gesomyrmex miegi Theobald 1937 [nomen dubium], Leucotaphus permancus Cockerell 1927 [nomen dubium], Mutilla tenera Förster 1891 [nomen dubium], Myrmica venusta Heer 1867 [nomen dubium], Plagiolepis succini André 1895 [nomen dubium], Ponera croatica Heer 1849 [nomen dubium], Sphecomyrmicinae Wilson et al. 1976 [empty], Sphecomyrmines Boudinot et al. 2020 [invalid subgroup]
Fm. †Holopsenellidae Engel et al. 2016
+
G. †Holopsenella Engel et al. 2016
+
†Holopsenella antiqua Lepeco and Melo 2021
†Holopsenella burmitica Lepeco and Melo 2021
†Holopsenella gothica Lepeco and Melo 2021
†Holopsenella primotica Engel et al. 2016
G. †Khetania Dlussky 1999
+
†Khetania mandibulata Dlussky 1999
Fm. †Melittosphecidae Poinar and Danforth 2006
+
G. †Melittosphex Poinar and Danforth 2006
+
†Melittosphex burmensis Poinar and Danforth 2006
G. †Mesosirex Hong 1984
+
†Mesosirex volantis Hong 1984
G. †Petropone Dlussky 1975
+
†Petropone petiolata Dlussky 1975
Superfm. Pompiloidea Latreille 1804
+
Fm. †Burmusculidae Zhang and Rasnitsyn 2018
+
G. †Burmusculus Zhang and Rasnitsyn 2018
Fm. Mutillidae Latreille 1802 [velvet ant]
+
G. †Cretavus Sharov 1957
G. †Mesomutilla Zhang 1985
Subfm. Mutillinae Latreille 1802
Subfm. Sphaeropthalminae Schuster 1949
Invalid names: Cretavidae Sharov 1957 [synonym]
Fm. Myrmosidae Fox 1894
+
Subfm. Kudakrumiinae Krombein 1979
Tr. Myrmosini Fox 1894
Invalid names: Kudakrumiini Krombein 1979 [empty]
Fm. Pompilidae Latreille 1804 [spider wasp]
+
Subfm. Pepsinae Lepeletier 1845
Subfm. Pompilinae Latreille 1804
G. †Pompilites Rodriguez 2017
Invalid names: Psammocharidae [synonym]
Fm. Sapygidae Latreille 1810
+
Subfm. †Cretosapyginae Bennett and Engel 2005
Subfm. Fedtschenkiinae André 1899
Superfm. Scolioidea Latreille 1802
+
G. †Oryctopterus Rasnitsyn 1975
+
†Oryctopterus dubius Rasnitsyn 1975
Fm. Scoliidae Latreille 1802 [hairy flower wasp]
+
G. †Araripescolia Nel et al. 2013
Subfm. †Archaeoscoliinae Rasnitsyn 1993
Subfm. †Palaeoscoliinae Antropov 2014
Subfm. Proscoliinae Rasnitsyn 1977
Subfm. Scoliinae Latreille 1802
Invalid names: Bradynobaenidae Saussure 1892 [empty]
Fm. †Stigmaphronidae Kozlov 1975
+
G. †Allocotidus Muesebeck 1963
+
†Allocotidus bruesi Muesebeck 1963
†Allocotidus melpomene Kozlov 1975
G. †Aphrostigmon Rasnitsyn 1991
+
†Aphrostigmon bontsaganicum Rasnitsyn 1991
†Aphrostigmon priapulus Rasnitsyn 1991
†Aphrostigmon vitimense Rasnitsyn 1991
G. †Burmaphron Engel and Grimaldi 2009
+
†Burmaphron iratxoak Ortega-Blanco et al. 2011
†Burmaphron jentilak Ortega-Blanco et al. 2011
†Burmaphron prolatum Engel and Grimaldi 2009
†Burmaphron sorginak Ortega-Blanco et al. 2011
†Burmaphron tridentatum Engel and Grimaldi 2009
G. †Elasmomorpha Kozlov 1975
G. †Elasmophron Engel and Grimaldi 2009
+
†Elasmophron kurthi Engel and Grimaldi 2009
†Elasmophron mari Ortega-Blanco et al. 2011
G. †Hippocoon Kozlov 1975
+
†Hippocoon basajauni Ortega-Blanco et al. 2011
†Hippocoon evadne Kozlov 1975
G. †Libanophron Engel and Grimaldi 2009
+
†Libanophron astarte Engel and Grimaldi 2009
†Libanophron sugaar Ortega-Blanco et al. 2011
G. †Stigmaphron Kozlov 1975
+
†Stigmaphron orphne Kozlov 1975
G. †Tagsmiphron Engel and Grimaldi 2009
+
†Tagsmiphron ascalaphus Engel and Grimaldi 2009
†Tagsmiphron canadense Engel and Grimaldi 2009
†Tagsmiphron exitorum McKellar and Engel 2011
†Tagsmiphron gigas Engel and Grimaldi 2009
†Tagsmiphron leucki McKellar and Engel 2011
†Tagsmiphron muesebecki Engel and Grimaldi 2009
†Tagsmiphron olentzero Ortega-Blanco et al. 2011
†Tagsmiphron spiculum McKellar and Engel 2011
Superfm. Thynnoidea Shuckard 1841
+
Fm. Thynnidae Shuckard 1841
+
Subfm. Anthoboscinae Rohwer 1911
Subfm. Methochinae Rohwer 1916
Subfm. Myzininae
Superfm. Tiphioidea Leach 1815
+
Fm. Sierolomorphidae Brues and Melander 1932 [sierolomorphid wasp]
+
Subfm. †Loreisomorphinae Álvarez-Parra and Engel 2024
Subfm. Sierolomorphinae Brues and Melander 1932
Superfm. Vespoidea Latreille 1802
+
Fm. †Falsiformicidae Rasnitsyn 1975
+
G. †Burmomyrma Dlussky 1996
G. †Falsiformica Rasnitsyn 1975
G. †Falsiformix Zhang and Rasnitsyn 2024
G. †Siccibythus Cockx and McKellar 2016
G. †Taimyrisphex Evans 1973
+
G. †Cretolixon Lohrmann 2020
G. †Eorhopalosoma Engel 2008
G. Rhopalosoma Cresson 1865
Fm. Tiphiidae Leach 1815
+
Subfm. †Burmatiphiinae Zheng et al. 2022
Subfm. †Dryophiinae Antropov 2014
G. Paratiphia Saussure and Sichel 1864
Subfm. †Thanatotiphiinae Engel et al. 2009
Subfm. Tiphiinae Leach 1815
Fm. Vespidae Latreille 1802
+
G. Agelaia Lepeletier de Saint-Fargeau 1835
G. †Archaeovespa Wu et al. 2020
Subfm. Eumeninae Leach 1815 [potter wasp]
Subfm. Euparagiinae Ashmead 1902
Subfm. Polistinae Lepeletier de Saint-Fargeau 1835
G. Polybia Lepeletier de Saint-Fargeau 1835
Subfm. †Priorvespinae Carpenter and Rasnitsyn 1990
Subfm. †Protovespinae Perrard and Carpenter 2017
Subfm. †Vespatulinae Jouault 2023
Subfm. Vespinae Latreille 1802
Invalid names: Bocchus cenomanianus Olmi et al. 2010 [nomen dubium], Bombus pusillus Menge 1856 [nomen dubium]
G. †Apocritites Rasnitsyn and Jarzembowski 1998
+
†Apocritites distinctus Rasnitsyn and Jarzembowski 1998
Infraor. Brachycleistogastromorpha Kovalev 1994
+
Superfm. †Brachycleistogastroidea Kovalev 1994
+
Fm. †Brachycleistogastridae Kovalev 1994
+
G. †Brachycleistogaster Rasnitsyn 1975
G. †Microcleistogaster Rasnitsyn 1975
G. †Neocleistogaster Rasnitsyn 1986
Invalid names: Cleistogastroidea Rasnitsyn 1975 [empty]
Superfm. Ceraphronoidea Haliday 1833
+
Fm. †Ceospilidae Engel 2025
+
G. †Ceospilus Engel 2025
+
†Ceospilus phoenicius Engel 2025
Fm. Ceraphronidae Haliday 1833
+
G. Aphanogmus Thomson 1858
+
†Aphanogmus serrulatus Meunier 1916
G. Ceraphron Jurine 1807
+
†Ceraphron ceuthonymus Engel 2013
†Ceraphron fasciatus Meunier 1916
Invalid names: Ceraphronini Haliday 1833 [empty]
Fm. †Maimetshidae Rasnitsyn 1975
+
Subfm. †Maimetshinae Rasnitsyn 1975
+
G. †Afrapia Rasnitsyn and Brothers 2009
+
Invalid names: Afromaimetsha Rasnitsyn and Brothers 2009 [synonym]
G. †Crucimaimetsha Li et al. 2025
G. †Guyotemaimetsha Perrichot et al. 2004
+
Invalid names: Burmaimetsha Perrichot 2013 [synonym]
G. †Maimetshasia Perrichot 2013
G. †Turgonalus Rasnitsyn 1990
+
Invalid names: Ahiromaimetsha Perrichot et al. 2011 [synonym], Turgonaliscus Engel 2016 [synonym]
Invalid names: Ahiromaimetshini Engel 2016 [invalid subgroup], Cretogonalinae Rasnitsyn 1977 [synonym], Maimetshini Rasnitsyn 1975 [invalid subgroup]
Subfm. †Zorophratrinae Engel 2016
+
G. †Zorophratra Engel 2016
Fm. Megaspilidae Ashmead 1893
+
G. Lagynodes Forster 1841
+
†Lagynodes primordialis Brues 1940
†Lagynodes pseudocarinatus Szabó and Oehlke 1986
Subfm. Lagynodinae Masner and Dessart 1967
+
G. †Prolagynodes Alekseev and Rasnitsyn 1981
Subfm. Megaspilinae Ashmead 1893
+
G. Conostigmus Dahlbom 1858
G. Dendrocerus Ratzeburg 1852
+
Invalid names: Lygocerus forster 1856 [synonym]
Invalid names: Calliceratidae [synonym]
Superfm. Evanioidea Latreille 1802
+
Unr. Neoevanioides Engel 2006
+
Fm. †Anomopterellidae Rasnitsyn 1975
+
G. †Anomopterella Rasnitsyn 1975
G. †Choristopterella Li et al. 2013
G. †Synaphopterella Li et al. 2013
Unr. Aulaciformes Grimaldi and Engel 2005
+
Fm. †Baissidae Rasnitsyn 1975
+
Invalid names: Manlayidae Rasnitsyn 1986 [synonym]
Unr. Euaulacides Engel 2006
Unr. Evaniiformes Grimaldi and Engel 2005
+
Fm. †Andreneliidae Rasnitsyn and Martínez-Delclòs 2000
Fm. Evaniidae Latreille 1802 [ensign wasp]
+
Invalid names: Cretevaniidae Rasnitsyn 1975 [synonym]
Unr. †Protoevanioides Jouault et al. 2021
+
Fm. †Othniodellithidae Engel and Huang 2016
+
G. †Keratodellitha Jouault et al. 2021
G. †Othniodellitha Engel and Huang 2016
G. †Xenodellitha Engel 2017
Fm. †Praeaulacidae Rasnitsyn 1972
+
G. †Albiogonalys Nel et al. 2003
Subfm. †Cretocleistogastrinae Rasnitsyn 1990
Subfm. †Nevaniinae Zhang and Rasnitsyn 2007
Subfm. †Praeaulacinae Rasnitsyn 1972
Fm. †Vectevaniidae Li et al. 2018
+
G. †Vectevania Cockerell 1922
+
†Vectevania vetula Cockerell 1922
Invalid names: Asiapterites Hong 2002 [nomen nudum], Euryopterites Hong 2002 [nomen nudum], Oblongigastrula Hong 2002 [nomen nudum]
Superfm. Ichneumonoidea Latreille 1802
+
Fm. Braconidae Nees ab Esenbeck 1811
+
Subfm. Agathidinae Haliday 1833
+
G. Agathis Latreille 1804
G. Bassus Fabricius 1804
+
Invalid names: Microdus Nees 1812 [objective synonym]
G. Cremnops Förster 1862
G. Earinus Wesmael 1837
G. Aivalykus Nixon 1938
+
†Aivalykus dominicanus Zuparko and Poinar 1997
G. Alysia Latreille 1804
+
†Alysia exigua Brues 1910
†Alysia latifrons Statz 1936
†Alysia meunieri Theobald 1937
†Alysia petrina Brues 1910
†Alysia phanerognatha Cockerell 1927
†Alysia ruskii Cockerell 1913
Subfm. Alysiinae Leach 1815
+
Tr. Alysiini Förster 1862
G. Tanycarpa Förster 1862
Subfm. Aphidiinae Haliday 1833
+
Tr. Aphidiini Haliday 1833
G. Aphidius Nees 1818
G. †Archephedrus Ortega-Blanco et al. 2009
G. †Archipraon Stary 1973
G. Diaeretus Förster 1862
Tr. Ephedrini Mackauer 1961
G. †Holocnomus Quilis Pérez 1940
G. †Oligoaphidius Quilis Pérez 1940
G. †Palaeoxys Mackauer 1961
G. †Praeaphidius Stary 1973
G. Praon Haliday 1833
G. †Propraon Brues 1933
G. †Protephedrus Quilis Pérez 1940
G. †Protodiaeretiella Stary 1973
G. †Pseudaphidius Stary 1973
G. Trioxys Haliday 1833
Subfm. Apozyginae Mason 1978
+
G. †Aenigmabracon Perrichot et al. 2009
G. Austrohelcon Turner 1918
Subfm. Betylobraconinae Tobias 1979
+
G. Mesocentrus Szépligeti 1900
Invalid names: Betylobraconini Tobias 1979 [invalid subgroup]
Subfm. Brachistinae Förster 1862
+
Tr. Blacini Foerster 1862
Tr. Brachistini Förster 1862
Tr. Diospilini Förster 1862
Invalid names: Triaspidinae Viereck 1918 [synonym]
Subfm. Braconinae Nees ab Esenbeck 1811
+
Tr. Aphrastobraconini Ashmead 1900
G. †Bembracon Belokobylskij 2014
Tr. Braconini Nees ab Esenbeck 1811
Subfm. Cardiochilinae Ashmead 1900
+
G. †Oligoneuroides Brues 1910
Subfm. Charmontinae van Achterberg 1979
+
Tr. †Palaeocharmontini Belokobylskij et al. 2010
Subfm. Cheloninae Nees 1816
+
G. †Anacanthobracon Brues 1939
G. Ascogaster Wesmael 1835
Tr. †Chelonohelconini Tobias 1987
G. Chelonus Panzer 1806
G. †Diodontogaster Brues 1933
G. Phanerotoma Wesmael 1838
Tr. Phanerotomini Baker 1926
G. Coeloreuteus Roman 1910
+
†Coeloreuteus gracilior Brues 1933
†Coeloreuteus resinae Brues 1933
G. †Digastrotheca Brues 1933
+
Subfm. †Diospilitinae Tobias 1987
+
G. †Diospilites Brues 1933
Subfm. Doryctinae Förster 1862
+
G. Doryctes Haliday 1839
G. †Eocenhecabolus Belokobylskij 2023
Tr. Hecabolini Förster 1862
G. Ontsira Cameron 1900
G. †Palaeorhoptrocentrus Belokobylskij 2023
Tr. Spathiini Förster 1862
G. †Electrohelcon Brues 1933
+
†Electrohelcon foveatus Brues 1933
†Electrohelcon grandis Brues 1933
†Electrohelcon minor Brues 1933
G. †Eobracon Cockerell 1921
+
†Eobracon cladurus Cockerell 1921
G. †Eobraconus Rasnitsyn 1985
+
†Eobraconus inopinatus Rasnitsyn 1983
Invalid names: Eobracon Rasnitsyn 1983 [replaced]
Subfm. Euphorinae Förster 1862
+
Tr. Centistini Čapek 1970
Tr. Euphorini Förster 1862
Tr. Meteorini Cresson 1887
Tr. Neoneurini Bengtsson 1918
Tr. †Oncometeorini Tobias 1987
G. †Palaeorionis Belokobylskij and Hovorka 2022
Tr. Perilitini Förster 1862
Tr. †Prosyntretini Tobias 1987
Subfm. Exothecinae Förster 1862
+
G. Chremylus Haliday 1833
G. Colastes Haliday 1833
+
Invalid names: Exothecus Wesmael 1838 [synonym]
Tr. Hormiini Förster 1862
G. †Palaeocolastes Belokobylskij and Zaldívar-Riverón 2021
G. †Palaeorhyssalus Brues 1933
Tr. Rhysipolini Belokobylskij 1984
Subfm. Helconinae Förster 1862
+
Tr. †Acampsohelconini Tobias 1987
G. Aspicolpus Wesmael 1836
G. †Calyptoides Cockerell 1921
G. Eumacrocentrus Ashmead 1900
Tr. Helconini Förster 1862
G. Hellenius Tobias 1982
Subfm. Homolobinae van Achterberg 1979
+
G. Homolobus Förster 1862
G. Hormiopterus Giraud 1869
+
†Hormiopterus petrinus Scudder 1877
Subfm. Ichneutinae Förster 1862
+
Tr. Ichneutini Förster 1862
Tr. Muesebeckiini Mason 1969
Tr. Proteropini van Achterberg 1976
G. Iphiaulax Förster 1862
Subfm. Masoninae van Achterberg 1995
+
Tr. Masonini van Achterberg 1995
Subfm. †Megalyrhyssalinae Belokobylskij and Jouault 2021
+
G. †Megalyrhyssalus Belokobylskij and Jouault 2021
Subfm. Microgastrinae Förster 1862
+
G. Apanteles Förster 1862
G. †Dacnusites Cockerell 1921
G. Microgaster Latreille 1804
G. Microplitis Förster 1862
G. †Palaeomicrogaster Belokobylskij 2014
G. Semionis Nixon 1965
Invalid names: Semionini Tobias 1987 [invalid subgroup]
Subfm. Microtypinae Szepligeti 1908
+
G. Microtypus Ratzeburg 1848
+
Invalid names: Diospiloides Cockerell 1921 [synonym]
Subfm. Miracinae
+
G. †Miracoides Brues 1933
Subfm. Opiinae Blanchard 1845
+
G. Biosteres Förster 1862
G. Diachasma Förster 1862
+
Invalid names: Bathystomus Förster 1862 [synonym]
G. Opius Wesmael 1835
Subfm. Orgilinae Ashmead 1900
+
G. Orgilus Haliday 1833
G. †Paleolebus Žalohar et al. 2025
+
†Paleolebus davois Žalohar et al. 2025
G. Phanomeris Förster 1862
Subfm. †Protobraconinae van Achterberg and Chen 2020
+
G. †Chainochora van Achterberg and Chen 2021
G. †Kleistochora van Achterberg and Chen 2021
G. †Protobracon van Achterberg and Chen 2020
G. †Rhetinorhyssalites Engel et al. 2017
G. †Tibialobracon van Achterberg and Chen 2020
Subfm. †Protorhyssalinae Basibuyuk et al. 1999
+
G. †Archaeorhyssalus Engel 2016
G. †Burmabracon Li et al. 2021
G. †Cretorhyssalus Belokobylskij 2012
G. †Diorhyssalus Engel 2016
G. †Magadanobracon Belokobylskij 2012
G. †Protorhyssalodes Perrichot et al. 2009
G. †Protorhyssalopsis Ortega-Blanco et al. 2011
G. †Protorhyssalus Basibuyuk and Quicke 1999
G. †Rhetinorhyssalus Engel 2016
G. †Seneciobracon Engel and Huang 2018
G. †Utrillabracon Álvarez-Parra and Engel 2022
G. †Pyramidibracon van Achterberg and Chen 2020
+
†Pyramidibracon clypeatus Chen and van Achterberg 2020
G. Rhaconotus Ruthe 1854
Subfm. Rhyssalinae Förster 1862
+
G. Dolopsidea Hincks 1944
G. Doryctomorpha Ashmead 1900
G. Oncophanes Förster 1862
G. Rhyssalus Haliday 1833
Subfm. Rogadinae Förster 1862
+
Tr. Clinocentrini van Achterberg 1991
Tr. Rogadini Förster 1862
G. Semirhytus Szepligeti 1902
+
†Semirhytus caudatus Brues 1933
Subfm. Sigalphinae Haliday 1833
+
G. Urosigalphus Ashmead 1889
G. Sigalphus Latreille 1802
G. †Sinobracon Hong 1974
+
†Sinobracon speciosus Hong 1974
G. Snellenius Westwood 1882
+
†Snellenius succinalis Brues 1933
G. †Stephanorhyssalus Belokobylskij and Jouault 2021
+
†Stephanorhyssalus longiscapus Belokobylskij and Jouault 2021
G. †Vertibracon van Achterberg and Chen 2023
+
†Vertibracon brevistigma Chen and van Achterberg 2023
G. Xenarcha Förster 1862
Invalid names: Eoichneumonidae Jell and Duncan 1986 [synonym], Fushunobracon Hong 2002 [nomen nudum], Microgasterinae Förster 1862 [empty], Seneciobraconinae Engel and Huang 2018 [empty], Xuaulacites Hong 2002 [nomen nudum]
Fm. Ichneumonidae Latreille 1802 [ichneumon wasp]
+
G. Absyrtus Holmgren 1858
+
†Absyrtus decrepitus Brues 1910
G. Acoenitus Latreille 1809
+
†Acoenitus lividus Heer 1849
G. †Carinibus Spasojevic et al. 2018
+
†Carinibus molestus Spasojevic et al. 2018
G. †Eopimpla Cockerell 1921
+
†Eopimpla grandis Cockerell 1921
G. †Epicharopimpla Zhang et al. 1994
+
†Epicharopimpla achaica Zhang et al. 1994
Subfm. Hybrizoninae Blanchard 1845
+
G. †Astigmaton Kasparyan 2001
G. †Ghilarovites Kasparyan 1988
Tr. †Tobiasitini Kasparyan 1988
G. †Vorstadt Manukyan 2023
Invalid names: Ghilarovitinae Kasparyan 1988 [synonym], Paxylommatinae Förster 1862 [synonym]
+
G. †Albertocryptus McKellar et al. 2013
Subfm. Cryptinae Kirby 1837
Subfm. Ichneumoninae Latreille 1802
G. †Ichneumonites Heer 1867
+
†Ichneumonites bellus Heer 1867
+
Subfm. Labeninae Ashmead 1900
Subfm. †Labenopimplinae Kopylov 2010
+
G. †Agapia Kopylov 2012
G. †Agapteron Kopylov 2012
G. †Armanopimpla Kopylov 2010
G. †Labenopimpla Kopylov 2010
G. †Micropimpla Kopylov 2010
G. †Pareubaeus McKellar et al. 2013
G. †Ramulimonstrum Kopylov 2010
G. †Rugopimpla Kopylov 2010
G. Lapton Nees 1815
+
†Lapton daemon Brues 1910
G. †Lithapechtis Cockerell 1921
+
†Lithapechtis fumosus Cockerell 1921
†Lithapechtis salmacidus Khalaim 2014
Subfm. Lycorininae Cushman and Rohwer 1920
+
G. Lycorina Holmgren 1859
G. †Megatryphon Cockerell 1924
+
†Megatryphon mortiferus Cockerell 1924
G. †Mesopimpla Cockerell 1919
+
†Mesopimpla sequoiarum Cockerell 1919
G. †Mesornatus Spasojevic et al. 2018
+
†Mesornatus markovici Spasojevic et al. 2018
Subfm. Microleptinae Townes 1958
+
G. Microleptes Gravenhorst 1829
G. †Miolyta Zhang 1989
+
†Miolyta volcania Zhang 1989
Subfm. †Novichneumoninae Li et al. 2016
+
G. †Caloichneumon Li et al. 2016
G. †Heteroichneumon Kopylov et al. 2021
G. †Heteropimpla Li et al. 2019
G. †Novichneumon Li et al. 2016
G. †Rasnichneumon Kopylov et al. 2021
G. †Rogichneumon Kopylov et al. 2021
Unr. Ophioniformes Gauld 1985
+
Subfm. Anomaloninae Viereck 1918
Subfm. Banchinae Townes 1951
Subfm. Campopleginae Förster 1869
Subfm. Cremastinae Förster 1869
Subfm. Ctenopelmatinae Förster 1869
Subfm. Mesochorinae Förster 1869
Subfm. Metopiinae Förster 1869
Subfm. Ophioninae Shuckard 1840
+
Invalid names: Ophionoidae Förster 1869 [objective synonym]
Subfm. Tersilochinae Schmiedeknecht 1910
+
Invalid names: Porizontinae Förster 1869 [replaced]
Subfm. Tryphoninae Shuckard 1840
+
Invalid names: Tryphonoidae Förster 1869 [objective synonym]
Subfm. Orthopelmatinae Schmiedeknecht 1910
+
G. Orthopelma Taschenberg 1865
Subfm. †Palaeoichneumoninae Kopylov 2009
+
G. †Dischysma Kopylov 2009
G. †Palaeoichneumon Kopylov 2009
G. †Rudimentifera Kopylov 2009
G. †Paratilgidopsis Martins-Neto 1998
+
†Paratilgidopsis praecursora Martins-Neto 1998
Subfm. †Pherhombinae Kasparyan 1988
+
G. †Furpherhombus Manukyan 2025
G. †Pherhombus Kasparyan 1988
Subfm. Phygadeuontinae Förster 1869
+
G. †Armadilleon Spasojevic et al. 2022
G. Brussinocryptus Pagliano and Scaramozzino 1990
+
Invalid names: Neoparacryptus Uchida 1959 [replaced]
Subtr. Ethelurgina Townes 1970
Subtr. Gelina Viereck 1918
G. †Grana Viertler et al. 2024
Subtr. Hemitelina Förster 1869
G. †Madma Viertler et al. 2022
G. †Magnocula Viertler et al. 2023
Subtr. Mastrina Townes 1970
G. †Osparvis Viertler et al. 2024
G. Phygadeuon Gravenhorst 1829
Subtr. Phygadeuontina Förster 1869
Subtr. Stilpnina Förster 1869
Invalid names: Hemigastrini Ashmead 1900 [synonym]
Unr. Pimpliformes Wahl 1990
+
Subfm. Acaenitinae Förster 1869
Subfm. Cylloceriinae Wahl 1990
Subfm. Diplazontinae Viereck 1918
+
Invalid names: Bassoidae Förster 1869 [replaced]
Subfm. Orthocentrinae Förster 1869
+
Invalid names: Plectiscoidae Förster 1869 [synonym]
Subfm. Pimplinae Wesmael 1845
+
Invalid names: Pimploidae Förster 1869 [objective synonym]
Subfm. Poemeniinae Narayanan and Lal 1953
Subfm. Rhyssinae Morley 1913
G. †Polychorella Zhang 1985
+
†Polychorella magica Zhang 1985
G. †Promyrmicium Baroni Urbani 1971
+
†Promyrmicium boreale Heer 1870
Invalid names: Myrmicium Heer 1870 [replaced]
Subfm. Stilbopinae Townes and Townes 1949
+
G. †Rovenosa Khalaim 2011
Subfm. †Tanychorinae Rasnitsyn 1980
+
G. †Amplicella Kopylov 2010
G. †Khasurtella Kopylov 2011
G. †Megachora Kopylov 2010
G. †Paratanychora Zhang and Rasnitsyn 2003
G. †Sinochora Kopylov and Zhang 2015
G. †Tanychora Townes 1973
G. †Tanychorella Rasnitsyn 1975
G. †Taubatehymen Martins-Neto 1998
+
†Taubatehymen minuta Martins-Neto 1998
G. †Tilgidopsis Cockerell 1921
+
†Tilgidopsis haesitans Cockerell 1921
Subfm. †Townesitinae Kasparyan 1994
+
G. †Acourtia Cockerell 1921
Tr. †Rasnitsynitini Kasparyan 1994
Tr. †Townesitini Kasparyan 1994
G. †Trymectus Spasojevic et al. 2018
+
†Trymectus amasidis Cockerell and Le Veque 1931
G. †Tryphopimpla Kopylov 2010
+
†Tryphopimpla xoridoptera Kopylov 2010
+
Subfm. Xoridinae Shuckard 1840
Invalid names: Eucerotinae Seyrig 1934 [empty], Ichneumonidae gabbroensis Handlirsch 1907 [nomen dubium], Liaoichneumon Hong 2002 [nomen nudum], Oxytorinae Thomson 1883 [empty], Polysphincta atavina Cockerell 1921 [nomen dubium]
Fm. †Ichneumonomimidae Rasnitsyn 1975
+
G. †Ichneumonomima Rasnitsyn 1975
+
†Ichneumonomima paradoxa Rasnitsyn 1975
G. †Ovigaster Zhang 1997
+
†Ovigaster cephalotus Zhang 1997
Fm. †Praeichneumonidae Rasnitsyn 1983
+
G. †Praeichneumon Rasnitsyn 1983
+
†Praeichneumon dzhidensis Kopylov 2012
†Praeichneumon townesi Rasnitsyn 1983
G. †Rasnitsynia Pagliano and Scaramozzino 1990
+
Invalid names: Oligoneuroides Zhang 1985 [replaced]
Fm. †Kuafuidae Rasnitsyn and Zhang 2010
+
G. †Arthrogaster Rasnitsyn 1975
+
†Arthrogaster seticornis Rasnitsyn 1975
G. †Kuafua Rasnitsyn and Zhang 2010
+
†Kuafua polyneura Rasnitsyn and Zhang 2010
G. †Leptogastrella Rasnitsyn 1975
+
†Leptogastrella leptogastra Rasnitsyn 1975
Fm. †Limnetidae Hong 1983
+
G. †Limnetus Hong 1983
+
G. †Longhuaia Hong 1984
+
†Longhuaia rara Hong 1984
Superfm. Megalyroidea Schletterer 1889
+
Fm. Megalyridae Schletterer 1889
+
Subfm. †Cleistogastrinae Rasnitsyn 1975
+
G. †Cleistogaster Rasnitsyn 1975
G. †Leptocleistogaster Rasnitsyn 1975
G. †Mesaulacinus Martynov 1925
G. †Scolichneumon Ren 1995
G. †Sinoryssus Hong 1983
G. †Stemmogaster Zhang 1985
G. †Yanocleistogaster Ren 1995
Invalid names: Sinoryssinae Hong 1983 [synonym]
Subfm. Megalyrinae Schletterer 1889
+
Tr. †Kamyristiini Brazidec et al. 2024
Tr. †Megallicini Brazidec et al. 2024
Tr. Megalyrini Schletterer 1889
+
Invalid names: Cretodinapsini Rasnitsyn 1977 [synonym]
Tr. †Valaaini Brazidec et al. 2024
Subfm. †Megazarinae Perrichot 2009
+
Tr. †Megalavini Brazidec et al. 2024
Tr. †Megazarini Perrichot 2009
Invalid names: Dinapsini Waterston 1922 [invalid subgroup]
Superfm. †Panguoidea Li et al. 2019
+
Fm. †Panguidae Li et al. 2019
+
G. †Pangu Li et al. 2019
+
†Pangu antiquum Li et al. 2020
†Pangu yuangu Li et al. 2019
G. †Prosphex Grimaldi and Engel 2019
+
†Prosphex anthophilos Grimaldi and Engel 2019
G. †Protopangu Zhuang et al. 2024
+
†Protopangu valdensis Zhuang et al. 2024
+
G. †Koonwarrus Rasnitsyn and Öhm-Kühnle 2019
+
†Koonwarrus katya Rasnitsyn and Öhm-Kühnle 2019
G. †Parviformosus Barling et al. 2013
+
†Parviformosus wohlrabeae Barling et al. 2013
Unr. Yangprocta Engel 2025
+
Fm. †Chalscelionidae Rasnitsyn and Öhm-Kühnle 2020
+
G. †Chalscelio Rasnitsyn and Brothers 2007
Fm. †Jurapriidae Rasnitsyn 1983
+
G. †Jurapria Rasnitsyn 1983
Fm. †Khutelchalcididae Rasnitsyn et al. 2004
+
G. †Khutelchalcis Rasnitsyn et al. 2004
Fm. †Mesoserphidae Kozlov 1970
+
G. †Amboserphus Li et al. 2016
G. †Apiciserphus Li et al. 2016
G. †Auliserphus Rasnitsyn 1986
G. †Basiserphus Li et al. 2016
G. †Beipiaoserphus Zhang and Zhang 2000
G. †Campturoserphus Rasnitsyn 1986
G. †Choriserphus Li et al. 2016
G. †Cretoserphus Rasnitsyn and Martínez-Delclòs 2000
G. †Juraserphus Zheng and Chen 2017
G. †Karataoserphus Rasnitsyn 1994
G. †Lordoserphus Rasnitsyn 1994
G. †Mesoserphus Kozlov 1968
G. †Novserphus Li et al. 2016
G. †Otlia Zhang 1992
G. †Ozososerphus Li et al. 2016
G. †Scoliuroserphus Rasnitsyn 1986
G. †Sinoserphus Shih et al. 2011
G. †Turgoserphus Rasnitsyn 1990
G. †Udaserphus Rasnitsyn 1983
G. †Yanliaoserphus Shih et al. 2011
Invalid names: Karataoserphinae Rasnitsyn 1994 [invalid subgroup], Mesoserphinae Kozlov 1970 [invalid subgroup]
Fm. †Peleserphidae Zhang et al. 2017
+
G. †Arkadiserphus Rasnitsyn 2021
G. †Peleproctus Zhang et al. 2017
G. †Peleserphus Zhang et al. 2017
Superfm. Proctotrupoidea Latreille 1802
+
G. †Amitchellia Rasnitsyn and Jarzembowski 1998
Fm. Heloridae Forster 1856
G. †Paraulacus Ping 1928
Fm. Pelecinidae Haliday 1840
+
Invalid names: Pelecinopteridae Brues 1933 [synonym]
Fm. Peradeniidae Naumann and Masner 1985
Fm. Proctotrupidae Latreille 1802
+
Invalid names: Serphidae Kieffer 1909 [synonym]
Invalid names: Austroniidae Kozlov 1975 [empty], Fushunoxyela Hong 2002 [nomen nudum], Hemerocynipites Hong 2002 [nomen nudum], Sinocynipites Hong 2002 [nomen nudum]
Unr. Trochantelliferida Engel 2025
+
Unr. Euprocta Engel 2025
Superfm. Ropronioidea Bradley 1905
Fm. †Uroprionulidae Engel 2025
+
G. †Uroprionulus Engel 2025
Unr. Yinprocta Engel 2025
+
Superfm. Cynipoidea Latreille 1802
+
Fm. †Archaeocynipidae Rasnitsyn and Kovalev 1988
Fm. Cynipidae Latreille 1802
Fm. Figitidae Forster 1869
Fm. †Gerocynipidae Liu and Engel 2007
Fm. Ibaliidae Forster 1869
Fm. Liopteridae Ashmead 1895
Fm. †Protimaspidae Liu and Engel 2007
Fm. †Stolamissidae Liu and Engel 2007
Invalid names: Archaeocynipoidea Rasnitsyn and Kovalev 1988 [synonym], Diplolepididae Latreille 1802 [empty]
Superfm. Platygastroidea Haliday 1833
+
Fm. †Caradiophyodidae Poinar and Vega 2023
Tr. Geoscelionidae Engel and Huang 2016
Fm. Neuroscelionidae Johnson and Austin 2021
Fm. Platygastridae Haliday 1833
Fm. †Proterosceliopsidae Talamas et al. 2019
Subfm. Scelioninae Thomson 1858
+
Invalid names: Platychalidae Kaddumi 2005 [synonym]
Fm. Sparasionidae Dahlbom 1858
Invalid names: Laelius nudipennis Brues 1933 [nomen dubium]
Invalid names: Cynipomorpha Kovalev 1994 [synonym], Microprocta Rasnitsyn and Öhm-Kühnle 2020 [invalid subgroup]
Superfm. Stephanoidea Leach 1815
+
Fm. †Aptenoperissidae Rasnitsyn and Poinar 2016
+
G. †Aptenoperissus Rasnitsyn and Poinar 2016
+
†Aptenoperissus amabilis Zhang and Rasnitsyn 2017
†Aptenoperissus burmanicus Rasnitsyn and Poinar 2016
†Aptenoperissus delicatus Zhang and Rasnitsyn 2017
†Aptenoperissus etius Rasnitsyn and Öhm-Kühnle 2018
†Aptenoperissus formosus Zhang and Rasnitsyn 2017
†Aptenoperissus magnifemoris Rasnitsyn and Öhm-Kühnle 2018
†Aptenoperissus pusillus Rasnitsyn and Öhm-Kühnle 2018
†Aptenoperissus zonalis Zhang and Rasnitsyn 2018
Fm. †Ephialtitidae Handlirsch 1906
+
Subfm. †Ephialtitinae Handlirsch 1906
+
G. †Acephialtitia Li et al. 2015
G. †Altephialtites Rasnitsyn 2008
G. †Asiephialtites Rasnitsyn 1975
G. †Chorephialtites Jia et al. 2025
G. †Cratephialtites Rasnitsyn 1999
G. †Crephanogaster Rasnitsyn 1990
G. †Ephialtites Meunier 1903
G. †Leptephialtites Rasnitsyn 1975
G. †Liadobracona Zessin 1981
G. †Mesephialtites Rasnitsyn 1975
G. †Montsecephialtites Rasnitsyn and Martínez-Delclòs 2000
G. †Parephialtites Rasnitsyn 1975
G. †Praeproapocritus Rasnitsyn and Zhang 2010
G. †Proapocritus Rasnitsyn 1975
G. †Proephialtitia Li et al. 2015
G. †Sessiliventer Rasnitsyn 1975
G. †Sinephialtites Zhang 1986
G. †Sinocephus Hong 1983
+
Invalid names: Hongicephus Wei 1995 [objective synonym]
G. †Stephanogaster Rasnitsyn 1975
G. †Thilopterus Rasnitsyn et al. 2003
G. †Tuphephialtites Zhang et al. 2002
G. †Kotaphialtites Rasnitsyn 2008
+
†Kotaphialtites frankmortoni Rasnitsyn 2008
Subfm. †Symphytopterinae Rasnitsyn 1980
+
G. †Brigittepterus Rasnitsyn et al. 2003
G. †Cephenopsis Hong 1984
G. †Karataus Rasnitsyn 1977
G. †Karataviola Rasnitsyn 1975
G. †Micrephialtites Rasnitsyn 1975
G. †Symphyogaster Rasnitsyn 1975
G. †Symphytopterus Rasnitsyn 1975
+
Invalid names: Sippelipterus Zessin 1985 [synonym]
G. †Trigonalopterus Rasnitsyn 1975
Invalid names: Karataidae Rasnitsyn 1977 [synonym]
Fm. †Myanmarinidae Zhang and Rasnitsyn 2017
+
G. †Myanmarina Zhang and Rasnitsyn 2017
+
†Myanmarina diversa Zheng et al. 2022
†Myanmarina jeannineae Li et al. 2018
†Myanmarina kachin Zhang and Rasnitsyn 2017
†Myanmarina lahu Zhang and Rasnitsyn 2017
†Myanmarina lisu Zhang and Rasnitsyn 2017
†Myanmarina sidorchukae Jouault et al. 2020
Fm. †Ohlhoffiidae Jouault et al. 2020
+
G. †Cretephialtites Rasnitsyn and Ansorge 2000
+
†Cretephialtites pedrerae Rasnitsyn and Ansorge 2000
G. †Myanmephialtites Jouault et al. 2020
+
†Myanmephialtites bashkuevi Jouault et al. 2020
G. †Ohlhoffia Jouault et al. 2020
+
†Ohlhoffia minuta Jouault 2025
†Ohlhoffia robusta Jouault et al. 2020
Fm. Stephanidae Leach 1815 [crown wasp]
+
G. †Aphanostephanus Ge and Tan 2023
+
Subfm. †Lagenostephaninae Ge and Tan 2023
+
G. †Lagenostephanus Li et al. 2017
G. †Neurastephanus Ge and Tan 2023
G. †Squamastephanus Ge and Tan 2025
G. †Tumidistephanus Ge and Tan 2023
G. †Phoriostephanus Engel and Huang 2016
+
†Phoriostephanus exilis Engel and Huang 2016
Subfm. Schlettereriinae Orfila 1949
+
G. †Acanthostephanus Ge and Tan 2025
G. †Archaeostephanus Engel and Grimaldi 2004
G. †Kronostephanus Engel and Grimaldi 2013
Subfm. Stephaninae Leach 1815
+
G. †Burmastephanus Ge and Tan 2025
G. †Denaeostephanus Engel and Grimaldi 2004
G. †Electrostephanus Brues 1933
G. †Protostephanus Cockerell 1906
Invalid names: Electrostephaninae Engel 2005 [synonym]
G. Stephanus Panzer 1805
Invalid names: Eostephanitidae Hong 2002 [invalid subgroup], Ephialtitoidea Handlirsch 1906 [synonym]
Superfm. Trigonalyoidea Cresson 1887
+
Fm. Trigonalidae Cresson 1887
+
G. †Darbigonalus Rasnitsyn 1986
+
†Darbigonalus capitatus Rasnitsyn 1986
G. †Eotrigonalis Poinar 2005
+
†Eotrigonalis balticus Poinar 2005
G. Trigonalys Westwood 1835
+
†Trigonalys bischoffi Statz 1938
Invalid names: Apiaria lapidea Germar 1842 [nomen dubium], Asiacynipites Hong 2002 [nomen nudum], Dendrocerus dubitatus Brues 1937 [nomen dubium], Eucynipites Hong 2002 [nomen nudum], Evaniomorpha Rasnitsyn 1988 [invalid subgroup], Stephanomorpha Leach 1815 [invalid subgroup]
Superfm. Cephoidea Newman 1834 [stem sawfly]
+
Fm. Cephidae Newman 1834 [stem sawfly]
+
Subfm. Cephinae Newman 1834
+
G. Cephus Latreille 1802
G. †Electrocephus Konow 1897
+
G. Janus Stephens 1835
+
†Janus disperditus Cockerell 1913
G. †Mesocephus Rasnitsyn 1968
+
†Mesocephus brachycerus Kopylov and Rasnitsyn 2016
†Mesocephus leleji Kopylov and Rasnitsyn 2016
†Mesocephus sibiricus Rasnitsyn 1968
Subfm. †Cuspilonginae Kopylov and Rasnitsyn 2016
+
G. †Cuspilongus Archibald and Rasnitsyn 2015
+
†Cuspilongus cachecreekensis Archibald and Rasnitsyn 2015
†Cuspilongus ghilarovi Rasnitsyn 1988
Fm. †Sepulcidae Rasnitsyn 1968
+
Subfm. †Ghilarellinae Rasnitsyn 1988
+
G. †Ghilarella Rasnitsyn 1988
+
†Ghilarella alexialis Kopylov and Rasnitsyn 2017
†Ghilarella elegantula Li et al. 2024
†Ghilarella kopylovi Li et al. 2024
†Ghilarella masculina Kopylov and Rasnitsyn 2017
†Ghilarella mercurialis Rasnitsyn 1988
†Ghilarella nivalis Rasnitsyn 1993
G. †Meiaghilarella Rasnitsyn and Martínez-Delclòs 2000
+
†Meiaghilarella cretacica Rasnitsyn and Martínez-Delclòs 2000
Subfm. †Parapamphiliinae Rasnitsyn 1968
+
G. †Micramphilius Rasnitsyn 1993
+
†Micramphilius minutus Rasnitsyn 1993
†Micramphilius mirabilipennis Kopylov and Rasnitsyn 2017
†Micramphilius secundus Rasnitsyn 1993
G. †Pamparaphilius Rasnitsyn 1993
+
†Pamparaphilius khasurtensis Kopylov and Rasnitsyn 2017
†Pamparaphilius mongolensis Rasnitsyn 1993
†Pamparaphilius semen Rasnitsyn 1993
†Pamparaphilius vitimicus Rasnitsyn 1993
G. †Parabakharius Rasnitsyn 1993
+
†Parabakharius parvus Rasnitsyn 1993
G. †Parapamphilius Rasnitsyn 1968
+
†Parapamphilius confusus Rasnitsyn 1968
G. †Shurabisca Rasnitsyn 1968
+
†Shurabisca deteria Rasnitsyn 1993
†Shurabisca liassica Rasnitsyn 1968
G. †Yananphilius Jouault 2024
+
Subfm. †Sepulcinae Rasnitsyn 1968
+
G. †Sepulca Rasnitsyn 1968
+
†Sepulca mirabilis Rasnitsyn 1968
†Sepulca mongolica Rasnitsyn 1993
G. †Sepulenia Rasnitsyn 1968
+
†Sepulenia syricta Rasnitsyn 1968
Subfm. †Trematothoracinae Rasnitsyn 1988
+
G. †Prosyntexis Sharkey 1990
+
†Prosyntexis antennata Li et al. 2023
†Prosyntexis aristovi Li et al. 2023
†Prosyntexis gobiensis Rasnitsyn 1993
†Prosyntexis gouleti Sharkey 1990
+
Invalid names: Prosyntexis legitima Martins-Neto 2007 [synonym]
†Prosyntexis lata Li et al. 2023
†Prosyntexis montsecensis Rasnitsyn and Ansorge 2000
†Prosyntexis okhotensis Rasnitsyn 1993
†Prosyntexis sennlaubi Jouault and Nel 2021
G. †Thoracotrema Rasnitsyn 1988
+
†Thoracotrema caudatum Rasnitsyn 1988
†Thoracotrema mongolicum Rasnitsyn 1993
†Thoracotrema nebulosum Rasnitsyn 1993
†Thoracotrema obscuripenne Rasnitsyn 1993
†Thoracotrema oculatum Rasnitsyn 1988
†Thoracotrema vitimicum Rasnitsyn 1993
†Thoracotrema zonale Rasnitsyn 1993
G. †Trematothorax Rasnitsyn 1988
+
†Trematothorax baissensis Rasnitsyn 1988
†Trematothorax brachyurus Kopylov and Rasnitsyn 2017
†Trematothorax clementsi Rasnitsyn and Jarzembowski 1998
†Trematothorax demeter Rasnitsyn and Kopylov 2014
†Trematothorax extravenosus Kopylov and Rasnitsyn 2017
†Trematothorax ingodensis Rasnitsyn 1993
†Trematothorax kasparyani Rasnitsyn and Kopylov 2014
†Trematothorax liaoningensis Zhang et al. 2001
†Trematothorax valdensis Rasnitsyn and Jarzembowski 1998
†Trematothorax zhangi Kopylov and Rasnitsyn 2017
Invalid names: Trematothoracoides Zhang et al. 2001 [synonym]
Subfm. †Xyelulinae Rasnitsyn 1993
+
G. †Neoxyelula Rasnitsyn 1993
+
†Neoxyelula micron Rasnitsyn 1993
†Neoxyelula obscura Rasnitsyn 1993
†Neoxyelula oligophlebia Rasnitsyn 1993
†Neoxyelula reducta Rasnitsyn 1993
G. †Onokhoius Rasnitsyn 1990
+
†Onokhoius aculeatus Rasnitsyn 1993
†Onokhoius dubius Rasnitsyn 1990
†Onokhoius sogdianus Rasnitsyn 1993
†Onokhoius venustus Kopylov and Rasnitsyn 2017
G. †Xyelula Rasnitsyn 1969
+
†Xyelula alexandri Rasnitsyn 2008
†Xyelula bakharensis Rasnitsyn 1993
†Xyelula benderi Rasnitsyn et al. 2003
†Xyelula hybrida Rasnitsyn 1969
G. †Cratoenigma Krogmann and Nel 2012
+
†Cratoenigma articulata Krogmann and Nel 2012
Superfm. Orussoidea Newman 1834 [wood wasp]
+
Fm. †Burmorussidae Zhang et al. 2020
+
G. †Burmorussus Zhang et al. 2020
+
†Burmorussus mirabilis Zhang et al. 2020
G. †Cretorussus Jouault et al. 2021
+
†Cretorussus vilhelmseni Jouault et al. 2021
G. †Cretovelona Vilhelmsen et al. 2025
+
†Cretovelona orussopteryx Vilhelmsen et al. 2025
Subfm. †Mesorussinae Rasnitsyn 1977
+
G. †Mesorussus Rasnitsyn 1977
+
†Mesorussus taimyrensis Rasnitsyn 1977
Fm. Orussidae Newman 1834 [parasitic wood wasp]
+
G. †Baltorussus Schedl 2011
+
†Baltorussus velteni Schedl 2011
G. †Kryptovelona Vilhelmsen and Perrichot 2024
+
†Kryptovelona carstengroehni Vilhelmsen and Perrichot 2024
G. †Minyorussus Basibuyuk et al. 2000
+
†Minyorussus luzzii Basibuyuk et al. 2000
Subfm. Ophrynopinae Benson 1935
+
G. Ophrynopus Konow 1897
+
†Ophrynopus peritus Engel 2008
Subfm. Orussinae Newman 1834
+
G. Orussus Latreille 1796
+
†Orussus juttagroehnae Vilhelmsen and Perrichot 2024
Fm. †Paroryssidae Martynov 1925
+
G. †Microryssus Rasnitsyn 1968
+
†Microryssus antennatus Rasnitsyn 1968
†Microryssus brachyurus Rasnitsyn 1968
†Microryssus crassipes Rasnitsyn 1968
†Microryssus minus Rasnitsyn 1968
†Microryssus robustus Rasnitsyn 1968
†Microryssus subtilis Rasnitsyn 1968
G. †Paroryssus Martynov 1925
+
†Paroryssus extensus Martynov 1925
G. †Praeoryssus Rasnitsyn 1968
+
†Praeoryssus gracilis Rasnitsyn 1968
†Praeoryssus venosus Rasnitsyn 1968
G. †Renaliculus Lin 1980
+
Invalid names: Karatavitoidea Rasnitsyn 1963 [synonym]
Superfm. Siricoidea Billberg 1820
+
Fm. Anaxyelidae Martynov 1925
+
Subfm. †Anaxyelinae Martynov 1925
+
G. †Anasyntexis Rasnitsyn 1968
+
†Anasyntexis strophandra Rasnitsyn 1968
G. †Anaxyela Martynov 1925
+
†Anaxyela destructa Rasnitsyn 1969
†Anaxyela gracilis Martynov 1925
+
Invalid names: Anaxyela martynovi Cockerell 1928 [synonym]
†Anaxyela nana Rasnitsyn 1968
†Anaxyela parvula Rasnitsyn 1963
G. †Brachysyntexis Rasnitsyn 1969
+
†Brachysyntexis acuta Kopylov et al. 2020
†Brachysyntexis brachyura Rasnitsyn 1968
†Brachysyntexis brevicornis Kopylov et al. 2020
†Brachysyntexis laticella Kopylov et al. 2020
†Brachysyntexis micrura Rasnitsyn 1969
†Brachysyntexis minuta Kopylov et al. 2020
†Brachysyntexis nova Rasnitsyn 1969
†Brachysyntexis robusta Zhang and Rasnitsyn 2006
†Brachysyntexis rohweri Kopylov et al. 2020
†Brachysyntexis sinensis Kopylov et al. 2020
†Brachysyntexis tenebrosa Kopylov 2018
†Brachysyntexis tigris Kopylov 2018
G. †Dolichostigma Rasnitsyn 1968
+
†Dolichostigma tenuipes Rasnitsyn 1968
G. †Kempendaja Rasnitsyn 1968
+
†Kempendaja jacutensis Rasnitsyn 1968
G. †Kulbastavia Rasnitsyn 1968
+
†Kulbastavia grandis Kopylov 2018
†Kulbastavia macrura Rasnitsyn 1963
G. †Mangus Kopylov 2019
+
†Mangus magnus Kopylov 2019
G. †Sphenosyntexis Rasnitsyn 1969
+
†Sphenosyntexis antonovi Rasnitsyn 1963
†Sphenosyntexis pallicornis Rasnitsyn 1969
G. †Syntexyela Rasnitsyn 1968
+
†Syntexyela asiatica Rasnitsyn 1968
†Syntexyela continentalis Zhang and Rasnitsyn 2006
†Syntexyela gracilicornis Rasnitsyn 1968
†Syntexyela inversa Rasnitsyn 1968
†Syntexyela media Rasnitsyn 1963
G. †Urosyntexis Rasnitsyn 1969
+
†Urosyntexis depressa Rasnitsyn 1969
†Urosyntexis drepanura Rasnitsyn 1968
†Urosyntexis magna Rasnitsyn 1968
†Urosyntexis undosa Kopylov 2019
Invalid names: Dolichostigmatinae Rasnitsyn 1968 [synonym], Kempendajinae Rasnitsyn 1980 [synonym]
Subfm. Syntexinae Benson 1935
+
G. †Cretosyntexis Rasnitsyn and Martínez-Delclòs 2000
+
†Cretosyntexis montsecensis Rasnitsyn and Martínez-Delclòs 2000
G. †Curiosyntexis Kopylov 2019
+
†Curiosyntexis magadanicus Kopylov 2019
G. †Curvitexis Jouault et al. 2022
+
†Curvitexis kopylovi Jouault et al. 2022
G. †Daosyntexis Kopylov et al. 2020
+
†Daosyntexis primus Kopylov et al. 2020
G. †Dolichosyntexis Kopylov 2019
+
G. †Eosyntexis Rasnitsyn 1990
+
†Eosyntexis catalonicus Rasnitsyn and Martínez-Delclòs 2000
†Eosyntexis conflata Li et al. 2024
†Eosyntexis parva Ortega-Blanco et al. 2008
†Eosyntexis senilis Rasnitsyn 1990
†Eosyntexis tuffinae Rasnitsyn and Jarzembowski 1998
G. †Hemisyntexis Li et al. 2024
+
†Hemisyntexis lepida Li et al. 2024
G. †Karasyntexis Kopylov and Rasnitsyn 2025
+
†Karasyntexis martynovi Kopylov and Rasnitsyn 2025
G. †Orthosyntexis Gao et al. 2021
+
†Orthosyntexis elegans Gao et al. 2021
†Orthosyntexis mascula Li and Wei 2025
†Orthosyntexis thanti Gao et al. 2021
G. †Parasyntexis Kopylov 2019
+
†Parasyntexis khasurtensis Kopylov 2019
G. †Sclerosyntexis Wang et al. 2020
+
†Sclerosyntexis hirsuta Wang et al. 2020
Invalid names: Paraxiphydriinae Gao et al. 2022 [synonym]
Fm. †Protosiricidae Rasnitsyn and Zhang 2004
+
G. †Protosirex Rasnitsyn 1969
+
†Protosirex xyelopterus Rasnitsyn 1969
Fm. †Pseudosiricidae Handlirsch 1906
+
G. †Myrmicium Westwood 1854
+
†Myrmicium araripterum Freitas et al. 2020
†Myrmicium elongatum Germar 1839
†Myrmicium heerii Westwood 1854
†Myrmicium nanus Handlirsch 1906
†Myrmicium schroeteri Germar 1839
+
Invalid names: Pseudosirex antiquus Germar 1839 [synonym], Pseudosirex compressus Oppenheim 1885 [synonym], Pseudosirex darwini Weyenbergh 1873 [synonym], Pseudosirex deichmuelleri Handlirsch 1906 [synonym], Pseudosirex elegans Oppenheim 1885 [synonym], Pseudosirex gracilis Oppenheim 1885 [synonym], Pseudosirex karschi Oppenheim 1885 [synonym], Pseudosirex minimus Oppenheim 1885 [synonym], Pseudosirex separatus Handlirsch 1906 [synonym], Rhipidorhabdus schroeteri Oppenheim 1885 [synonym], Sphinx snelleni Weyenbergh 1869 [synonym]
Invalid names: Apiaria Germar 1839 [replaced], Fabellovena Oppenheim 1885 [synonym], Hagenia Weyenbergh 1869 [synonym], Pseudosirex Weyenbergh 1873 [synonym], Rhipidorhabdus Oppenheim 1885 [synonym]
Invalid names: Myrmiciidae Maa 1949 [synonym]
G. †Sinanxyela Hong 1982
+
†Sinanxyela dorunensis Hong 1982
Fm. †Sinosiricidae Hong 1975
+
G. †Sinosirex Hong 1975
+
†Sinosirex gigantea Hong 1975
Fm. Siricidae Billberg 1820 [horntail]
+
Subfm. †Auliscinae Rasnitsyn 1968
+
G. †Aulisca Rasnitsyn 1968
+
†Aulisca odontura Rasnitsyn 1968
†Aulisca variicornis Rasnitsyn 1968
G. †Megaulisca Rasnitsyn 1968
+
†Megaulisca grossa Rasnitsyn 1968
G. †Megura Rasnitsyn 1968
+
†Megura magnifica Rasnitsyn 1968
Subfm. †Cratosiricinae Jouault and Nel 2020
+
G. †Cratosirex Jouault and Nel 2020
+
†Cratosirex sennlaubi Jouault and Nel 2020
G. †Cretosirex Wang et al. 2018
+
†Cretosirex xiaoi Wang et al. 2018
G. †Emanrisca Rasnitsyn 1993
+
†Emanrisca empusa Rasnitsyn 1993
G. †Eoteredon Archibald et al. 2021
+
†Eoteredon lacoi Archibald et al. 2021
Subfm. †Gigasiricinae Rasnitsyn 1968
+
G. †Gigasirex Rasnitsyn 1968
+
†Gigasirex longipes Rasnitsyn 1968
G. †Liasirex Rasnitsyn 1968
+
†Liasirex sogdianus Rasnitsyn 1968
G. †Pararchexyela Rasnitsyn 1968
+
†Pararchexyela macroptera Rasnitsyn 1968
Invalid names: Pararchexyelinae Rasnitsyn 1968 [synonym]
Subfm. †Megapteritinae Maa 1949
+
G. †Megapterites Cockerell 1920
+
†Megapterites mirabilis Cockerell 1920
G. †Shurabica Rasnitsyn 1968
Subfm. Siricinae Billberg 1820
+
G. †Eourocerus Archibald and Rasnitsyn 2022
+
†Eourocerus anguliterreus Archibald and Rasnitsyn 2022
G. †Eoxeris Maa 1949
+
†Eoxeris klebsi Brues 1926
G. †Proximoxeris Nel 1991
+
+
Invalid names: Proximoxeris tripolicola Riou 1999 [objective synonym]
Invalid names: Proximoxeris Riou 1999 [objective synonym]
G. Sirex Linnaeus 1761
+
Invalid names: Paururus Konow 1896 [objective synonym]
G. †Urocerites Heer 1867
+
G. Urocerus Geoffroy 1762
+
†Urocerus dorbnikensis Manukyan 2021
Urocerus gigas Linnaeus 1758
†Urocerus ligniticus Piton 1940
†Urocerus patagonicus Fidalgo and Smith 1987
Invalid names: Eosirex Piton 1940 [synonym]
G. Xeris Costa 1894
+
†Xeris muratensis Boderau et al. 2024
G. Xoanon Semenov 1921
+
†Xoanon eocenicus Wedmann et al. 2014
G. †Ypresiosirex Archibald and Rasnitsyn 2015
+
†Ypresiosirex orthosemos Archibald and Rasnitsyn 2015
Subfm. Tremecinae Ashmead 1898
+
G. Eriotremex Benson 1943
G. Tremex Jurine 1807
Subor. Vespina Laicharting 1781
+
Invalid names: Euhymenoptera [synonym]
Superfm. Xiphydrioidea Leach 1819
+
Invalid names: Xiphydriidae Leach 1819 [empty]
Invalid names: Orussomorpha Rasnitsyn 1980 [invalid subgroup]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Kiessling 2004, Bush and Bambach 2015

Age range

Maximum range based only on fossils: base of the Volkhov to the top of the Holocene or 471.30000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 469.1 Ma

Collections (991 total)

Oldest occurrences

Time interval Ma Country or state Original ID and collection number
Volkhov471.3 - 469.1Russian Federation (Leningrag oblast) Aulocopsis simplex (235818)
Sevatian211.18 - 205.7Germany Sepulenia sp. (123417)
Early/Lower Toarcian184.2 - 182.9Germany (Mecklenburg-Vorpommern) Grimmaratavites mirabilis (114993) Xyelula benderi (154298)
Early/Lower Toarcian184.2 - 182.9Germany Ephialtitidae indet., Xyelula benderi (114991)
Early/Lower Toarcian184.2 - 182.9Germany (Niedersachsen) Liadobracona raduhna (139222) Sippelipterus liasinus (114995) Thilopterus lampei (154300) Xyelula benderi (154299)
Early/Lower Toarcian184.2 - 182.9Germany (Bavaria) Xyelula benderi (117794)
Toarcian184.2 - 174.7Germany Brigittepterus brauckmanni, Liadobracona raduhna, Xyelula benderi (104056)
Toarcian184.2 - 174.7Kyrgyzstan (Batken) Proapocritus praecursor, Onokhoius sogdianus (105003) Shurabisca liassica, Sepulenia syricta, Liasirex sogdianus (105124)
Toarcian184.2 - 174.7India (Andhra Pradesh) Xyelula alexandri (79229)
Toarcian184.2 - 174.7India (Adilabad) Kotaphialtites frankmortoni (117538)
Aalenian174.7 - 170.9China (Shaanxi) Yananphilius peizhuangensis (222545)
Bajocian - Bathonian170.9 - 165.3Mongolia (Bayan-Khongor) Parabakharius parvus (124340) Xyelula bakharensis (114615)
Callovian - Oxfordian165.3 - 154.8China (Liaoning) Beipiaosirex parva (154732) Mesohelorus haifanggouensis (157657) Paraulacus sinicus (156597) Praeaulacus scabratus, Praeaulacon grossus, Nevania deviata, Proapocritus habitus (228936) Sinocephus haifanggouensis (154733) Sinoryssus suni (154730)
Callovian - Oxfordian165.3 - 154.8China (Nei Mongol) Cathaypelecinus daohugouensis, Archaeopelecinus tebbei, Archaeopelecinus jinzhouensis (113360) Juraserphus modicus (157555) Karataus exilis, Stephanogaster rasnitsyni, Stephanogaster ningchengensis (179749) Novserphus ningchengensis, Choriserphus gigantus, Choriserphus bellus, Ozososerphus ovatus, Ozososerphus cuboidus, Ozososerphus lepidus, Karataoserphus sinicus, Karataoserphus adaequatus, Karataoserphus gracilentus, Apiciserphus augustus, Basiserphus longus, Sinoserphus wui, Sinoserphus flexilis, Sinoserphus petilus, Sinoserphus grossus, Mesoserphus venustus, Proephialtitia acantha, Proephialtitia tenuata (123761) Praeaulacus rectus, Stephanogaster integra (172659) Praeaulacus subrhombeus, Praeaulacus tenellus, Praeaulacus obtutus, Praeaulacus byssinus, Archaulacus probus, Nevania retenta, Nevania perbella, Nevania robusta, Nevania aspectabilis, Synaphopterella patula, Anomopterella sp., Anomopterella sp. 1, Anomopterella sp. 2, Anomopterella sp. 3, Anomopterella sp. 4, Anomopterella ovalis, Anomopterella coalita, Anomopterella brevis, Anomopterella pygmea, Anomopterella divergens, Anomopterella brachystelis, Anomopterella ampla, Archaeohelorus tensus, Archaeohelorus polyneurus, Symphytopterus graciler, Praeproapocritus flexus, Proapocritus parallelus, Proapocritus bialatus (138748) Praeparyssites orientalis, Praeratavites wuhuaensis, Praeratavites daohugou, Praeratavites perspicuus, Karatavites junfengi, Praeratavitoides amabilis, Postxiphydria ningchengensis, Postxiphydria daohugouensis, Postxiphydroides strenuus, Aulacogastrinus sp., Aulacogastrinus insculptus, Aulacogastrinus hebeiensis, Aulacogastrinus longaciculatus, Sinevania speciosa, Praeaulacus daohugouensis, Praeaulacus robustus, Praeaulacus afflatus, Praeaulacus sculptus, Praeaulacus exquisitus, Praeaulacus orientalis, Praeaulacus scabratus, Sinaulacogastrinus solidus, Sinaulacogastrinus eucallus, Eosaulacus granulatus, Eosaulacus giganteus, Praeaulacon elegantulus, Praeaulacon ningchengensis, Eonevania robusta, Nevania retenta, Nevania exquisita, Nevania malleata, Nevania ferocula, Nevania robusta, Nevania delicata, Anomopterella huangi, Kuafua polyneura, Karataus vigoratus, Karataus exilis, Karataus orientalis, Karataus strenuus, Karataus daohugouensis, Asiephialtites lini, Praeproapocritus vulgatus, Stephanogaster pristinus, Proapocritus sculptus, Proapocritus densipediculus, Proapocritus lini, Proapocritus longantennatus, Proapocritus atropus, Proapocritus elegans, Proapocritus formosus, Brachysyntexis sp., Brachysyntexis brevicornis, Brachysyntexis minuta, Brachysyntexis rohweri, Brachysyntexis laticella, Brachysyntexis acuta, Brachysyntexis brachyura, Brachysyntexis sinensis, Daosyntexis primus (113371) Praeratavites rasnitsyni, Karatavites medius, Karatavites ningchengensis (183155) Sinoserphus wui, Sinoserphus shihae, Sinoserphus lillianae, Yanliaoserphus jurassicus, Archaeohelorus hoi, Archaeohelorus polyneurus (113362)
Callovian - Oxfordian165.3 - 154.8China (Hebei) Limnetus wangyingziensis (113476) Sinephialtites glyptus (156314)
Oxfordian161.5 - 154.8Russian Federation (Buryat) Cleistogaster buriatica (105017)
Oxfordian - Kimmeridgian161.5 - 149.2Russian Federation (Buryat) Jurapria sibirica (117675) Udaserphus transbaicalicus, Bethylonymus sibiricus (104950)
Kimmeridgian154.8 - 149.2Kazakhstan Anaxyela dubia (114238) Anaxyela martynovi (122585) Auliserphus parvulus, Mesohelorus muchini, Stephanogaster longipalpa, Anaxyela macrura (31311) Evanigaster petiolatus, Nevania karatau, Brachycleistogaster nigritibialis, Mesaulacinus areolatus, Mesaulacinus tenuis, Cleistogaster caudata, Cleistogaster seticornis, Cleistogaster curtis, Leptocleistogaster oligomera, Oxyuroserphus leucurus, Auliserphus pallidus, Auliserphus caudatus, Auliserphus brachyurus, Scoliuroserphus pallidulus, Campturoserphus pumilus, Campturoserphus gibbus, Bethylonymus curtipes, Bethylonymus magnus, Arthrogaster seticornis, Symphytopterus geniculatus, Symphytopterus obliquus, Symphytopterus pallicornis, Symphytopterus nigripes, Symphytopterus nigricornis, Karataviola micrura, Karataviola leptogastra, Karataus pedalis, Asiephialtites seticornis, Stephanogaster brachyura, Leptephialtites minor, Anaxyelinae indet., Brachysyntexis tigris, Kulbastavia grandis (105426) Evaniops rostratus (115792) Karasyntexis martynovi (242274) Karatavites medius, Praeaulacus obscurus, Praeaulacus leptogaster, Praeaulacites pachygaster, Praeaulacon tenuis, Brachycleistogaster jurassica, Brachycleistogaster nigripes, Mesaulacinus sculpturatus, Leptocleistogaster pallida, Leptocleistogaster alboscuta, Auliserphus niger, Campturoserphus obscurus, Arkadiserphus leleji, Bethylonymellus cervicalis, Bethylonymellus pallens, Bethylonymellus bethyloides, Leptogastrella leptogastra, Symphyogaster cylindrica, Symphytopterus macrophthalmus, Symphytopterus robustus, Symphytopterus minutus, Parephialtites reductus, Stephanogaster longipalpa, Stephanogaster magna, Leptephialtites linearis, Leptephialtites angustus, Leptephialtites stephanocephalus, Leptephialtites caudatus, Leptephialtites picturatus, Sessiliventer oppositus, Xyelula hybrida, Protosirex xyelopterus, Brachysyntexis nova, Sphenosyntexis pallicornis, Urosyntexis depressa (113439) Microryssus antennatus, Microryssus crassipes, Aulacogaster ater, Praeaulacus ventricosus, Praeaulacus cubocephalus, Microcleistogaster dolichura, Brachycleistogaster tredecatoma, Brachycleistogaster nana, Cleistogaster oculata, Karataoserphinus minor, Mesoserphus karatavicus, Bethylonymellus oligocerus, Symphytopterus nigridorsatus, Symphytopterus nigricornis, Trigonalopterus brachycerus, Parapamphilius confusus, Pararchexyela macroptera, Megura magnifica, Syntexyela nana, Syntexyela magna (105970) Microryssus minus, Microryssus brachyurus, Microryssus subtilis, Microryssus robustus, Praeoryssus gracilis, Praeaulacus ramosus, Praeaulacinus minus, Praeaulacinus crassipes, Anomopterella mirabilis, Microcleistogaster parvula, Brachycleistogaster pentedecamera, Brachycleistogaster karatavica, Brachycleistogaster brachyura, Brachycleistogaster tetradecatoma, Leptocleistogaster propodeata, Oxyuroserphus sculpturatus, Karataoserphus dorsoniger, Auliserphus antennatus, Auliserphus jurassicus, Auliserphus interstitialis, Scoliuroserphus propodealis, Protohelorus mesozoicus, Bethylonymellus microgaster, Bethylonymellus euryurus, Bethylonymellus minutus, Bethylonymus microphthalmus, Symphytopterus ater, Symphytopterus nigricornis, Mesephialtites paurocerus, Stephanogaster longipalpa, Leptephialtites pallidus, Sessiliventer rostratus, Megaulisca grossa, Anaxyelidae indet., Anaxyelinae indet., Brachysyntexis micrura, Syntexyela asiatica, Syntexyela drepanura (106101) Microryssus robustus, Praeoryssus venosus, Praeaulacops lucidus, Praeaulacus elegans, Praeaulacus cephalotus, Praeaulacus ramosus, Praeaulacites medius, Praeaulacites nigripes, Praeaulacites minimus, Praeaulacinus minor, Praeaulacon coniventer, Praeaulacon caudatus, Praeaulacon elongatus, Microcleistogaster nigriventris, Microcleistogaster leptocera, Mesaulacinus niger, Leptocleistogaster paurura, Auliserphus imperfectus, Bethylonymellus imperfectus, Bethylonymellus buccatus, Bethylonymus micrurus, Symphytopterus oculatus, Symphytopterus pallipes, Symphytopterus robustus, Trigonalopterus brachycerus, Stephanogaster similis, Stephanogaster longipalpa, Leptephialtites euryarthrus, Leptephialtites tenuicornis, Leptephialtites gigas, Sessiliventer macrurus, Sepulca mirabilis, Gigasirex longipes, Aulisca variicornis, Aulisca odontura, Anaxyelinae indet., Brachysyntexis tenebrosa, Syntexyela brachyura, Syntexyela gracilicornis, Syntexyela inversa, Anasyntexis strophandra (105971) Paroryssus extensus, Karatavites angustus, Praeaulacites medius, Praeaulacites pachygaster, Brachycleistogaster nigripes, Brachycleistogaster scoliura, Brachycleistogaster galkinensis, Mesaulacinus oviformis, Symphytopterus clavicornis, Asiephialtites niger, Anaxyela antonovi, Anaxyela media, Anaxyela parvula, Anaxyela gracilis (115717) Praeaulacites subniger, Praeaulacites medius, Anomopterella stenocera, Brachycleistogaster nigritibialis, Mesaulacinus stenocerus, Leptocleistogaster alboscuta, Scoliuroserphus pallidulus, Bethylonymus pedalis, Bethylonymus nigricornis, Symphytopterus nigricornis, Asiephialtites ventricosus, Stephanogaster pyriformis, Sessiliventer symphytus, Anaxyelidae indet., Anaxyela destructa (106102) Praeaulacus magnus, Brachycleistogaster pallipes, Cleistogaster obscura, Leptocleistogaster picta, Karataoserphus meridionalis, Lordoserphus cyrturus, Bethylonymus robustus, Leptogastrella leptogastra, Micrephialtites minor, Asiephialtites oviventer, Sessiliventer temporalis (106279) Protocyrtus jurassicus (122330)
Early/Lower Tithonian149.2 - 145.06Germany (Bayern) Apiaria lapidea, Belostomum elongatum, Rhipidorhabdus schroeteri, Rhipidorhabdus gracilis, Fabellovena karschi, Fabellovena compressa, Fabellovena elegans, Rhipidorhabdus minimus, Apiaria antiqua, Cryptocephalus mesozoicus (123985) Cryptocephalus mesozoicus, Rhipidorhabdus schroeteri, Sphinx snelleni, Rhipidorhabdus minimus, Pseudosirex darwini (123906) Pseudosirex nanus, Myrmicium nanus, Myrmicium elongatus, Myrmicium schroeteri, Pseudosirex snelleni (123144) Pseudosirex schroeteri, Pseudosirex deichmuelleri, Pseudosirex minimus, Pseudosirex elongatus (133578)
Early/Lower Tithonian149.2 - 145.06Germany Sphinx sp. (115325)