PBDB Taxon

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Chirotheriidae

Reptilia - Chirotheriidae

Taxonomy

Chirotheriidae was named by Abel (1935).

It was assigned to Ornithosuchoidea by Kuhn (1964); to Thecodontia by Gand (1974); and to Pseudosuchia by Peabody (1948), Bock (1952), Baird (1957), Kuhn (1963), Kuhn (1966), Haubold (1969), Haubold (1971), Haubold (1974), Calvo (2007).

Synonyms

Paratrisauropus was named by Ellenberger (1970). It is considered to be a form taxon.

It was considered a nomen dubium by Cavin et al. (2012); it was synonymized subjectively with Grallator by Klein and Lucas (2021).

It was assigned to Ornithischia by Ellenberger (1972); to Ornithopoda by Ellenberger (1974), Romero Molina et al. (2003); and to Dinosauriformes by Irmis et al. (2007).
Deuterosauropodopus was named by Ellenberger (1970). It is considered to be a form taxon.

It was synonymized subjectively with Brachychirotherium by Olsen and Galton (1984); it was considered a nomen dubium by Cavin et al. (2012).

It was assigned to Sauropoda by Ellenberger (1970), Ellenberger (1972).
Deuterosauropodopus minor was named by Ellenberger (1970). It is considered to be a form taxon. It is a trace fossil. Its type locality is Seobeng tracksite (zone A/4), which is in a Sinemurian terrestrial sandstone in the Clarens Formation of Lesotho.

It was recombined as Brachychirotherium minor by Olsen and Galton (1984); it was considered a nomen dubium by D'Orazi Porchetti and Nicosia (2007).
Swinnertonichnus was named by Sarjeant (1967). It is considered to be a form taxon.

It was considered a nomen dubium by Klein and Lucas (2021).

It was assigned to Anchisauripodidae by Haubold (1971), Demathieu and Haubold (1972), Haubold (1974); and to Coelurosauria by Sarjeant (1975).
Swinnertonichnus mapperleyensis was named by Sarjeant (1967). It is considered to be a form taxon. Its type specimen is P.C.3315, a footprint, and it is a trace fossil. Its type locality is Mapperley Park tracksite, which is in an Anisian terrestrial sandstone in the Keuper Waterstones Formation of the United Kingdom.

It was considered a nomen dubium by Klein and Lucas (2021).
Shimmelia was named by Casamiquela (1964). It is considered to be a form taxon. Its type is Shimmelia chirotheroides.

It was synonymized subjectively with Chirotherium by Leonardi (1981); it was considered a nomen dubium by Melchor and de Valais (2006).

It was assigned to Archosauria by Casamiquela (1964); to Chirotherioidea by Casamiquela (1975); and to Chirotheridae by Calvo (2007).
Shimmelia chirotheroides was named by Casamiquela (1964). It is considered to be a form taxon. Its type specimen is MLP 60-XI-31-1, a footprint, and it is a trace fossil. Its type locality is Cantera Vieja, Tscherig Farm, which is in a Wuchiapingian/Olenekian terrestrial sandstone in the Vera Formation of Argentina. It is the type species of Shimmelia.

It was considered a nomen dubium by Melchor and de Valais (2006).
Chelonichnium was named by Schimper (1853). Its type is Chelonichnium vogesiacum.

It was considered a nomen dubium by Klein and Lucas (2021).

It was assigned to Chelonia by Schimper (1853).
Chirotherium angustum was named by Huene (1941). It is considered to be a form taxon. It is a trace fossil. Its type locality is Agnano tracksite, Monte Pisano, which is in a Carnian delta plain siltstone/claystone in the Quarziti di Monte Serra Formation of Italy.

It was considered a nomen dubium by Haubold (1971); it was synonymized subjectively with Chirotherium barthii by Klein and Lucas (2021).
Brachychirotherium tintanti was named by Demathieu (1971). It is considered to be a form taxon. Its type specimen is 21.1 W1, a footprint, and it is a trace fossil. Its type locality is Chasselay tracksite, level C (BQ), which is in an Anisian/Ladinian terrestrial horizon in the Grès Inférieurs du Lyonnais Formation of France.

It was considered a nomen dubium by Klein and Lucas (2021).
Dahutherium agilis was named by Montenat (1968). It is considered to be a form taxon. It is a trace fossil. Its type locality is Ailhon tracksite, Daüs, which is in an Anisian/Ladinian terrestrial sandstone in the Grès Inférieurs du Roubreau Formation of France. It is the type species of Dahutherium.

It was considered a nomen nudum by Klein and Lucas (2021).
Pachysaurichnium was named by Demathieu and Weidmann (1982). It is considered to be a form taxon.

It was considered a nomen dubium by Cavin et al. (2012), Klein and Lucas (2021).
Pseudochirotherium was named by Biron and Dutuit (1981). It is considered to be a form taxon. Its type is Pseudochirotherium oukaimedensis.

It was considered a nomen dubium by Klein and Lucas (2021).

It was assigned to Chirotheriidae by Biron and Dutuit (1981).
Pseudochirotherium oukaimedensis was named by Biron and Dutuit (1981). It is considered to be a form taxon. It is a trace fossil. Its type locality is Ourika Valley tracksites (general), which is in a Carnian terrestrial sandstone in the Oukaïmeden Sandstone Formation of Morocco. It is the type species of Pseudochirotherium.

It was considered a nomen dubium by Klein and Lucas (2021).
Pentichnus was named by Biron and Dutuit (1981). It is considered to be a form taxon.

It was considered a nomen dubium by Klein and Lucas (2021).

It was assigned to Dicynodontia by Biron and Dutuit (1981).
Pentichnus largus was named by Biron and Dutuit (1981). It is considered to be a form taxon. Its type specimen is ARG. 1018, 1012, a footprint, and it is a trace fossil. Its type locality is Locality XI, track level (Argana Basin), which is in a Carnian terrestrial horizon in the Timezgadiouine Formation of Morocco.

It was considered a nomen dubium by Klein and Lucas (2021).
Chirotherium ibericus was named by Navás (1906). It is considered to be a form taxon. It is a trace fossil.

It was considered a nomen dubium by Haubold (1971); it was synonymized subjectively with Chirotherium barthii by Klein and Lucas (2021).
Chirotherium catalaunicum was named by Casanovas-Cladellas et al. (1979). It is considered to be a form taxon. It is a trace fossil.

It was considered a nomen dubium by Klein and Lucas (2021).
Deuterosauropodopus sedunensis was named by Demathieu and Weidmann (1982). It is considered to be a form taxon. Its type specimen is track no. 26, a footprint, and it is a trace fossil. Its type locality is Col de Corbeaux-Vieil Emosson tracksite, which is in a Ladinian/Carnian lagoonal sandstone in the Grès bigarre Formation of Switzerland.

It was considered a nomen dubium by Cavin et al. (2012).
Pachysaurichnium emossonense was named by Demathieu and Weidmann (1982). It is considered to be a form taxon. Its type specimen is track no. 11, a footprint, and it is a trace fossil. Its type locality is Col de Corbeaux-Vieil Emosson tracksite, which is in a Ladinian/Carnian lagoonal sandstone in the Grès bigarre Formation of Switzerland.

It was considered a nomen dubium by Cavin et al. (2012).
Chirotherium lulli was named by Bock (1952). It is considered to be a form taxon. Its type specimen is LC S491, a footprint, and it is a trace fossil. Its type locality is Smith Clark's quarry, Milford (unit O), which is in a Norian terrestrial horizon in the Passaic Formation of New Jersey.

It was considered a nomen dubium by Klein and Lucas (2021).
Chirotherium wondrai was named by Heller (1952). It is considered to be a form taxon. Its type specimen is at Institut Erlangen, a footprint, and it is a trace fossil. Its type locality is Altselingsback tracksite, which is in a Carnian terrestrial sandstone in the Middle Keuper Formation of Germany.

It was considered a nomen dubium by Klein and Lucas (2021).
Thecodontichnus fucinii was named by Huene (1941). It is a trace fossil. Its type locality is Agnano tracksite, Monte Pisano, which is in a Carnian delta plain siltstone/claystone in the Quarziti di Monte Serra Formation of Italy.

It was considered a nomen dubium by Haubold (1971).
Chirotherium huberi was named by Bock (1952). Its type specimen is ANSP 15211, a footprint, and it is a trace fossil. Its type locality is Huber's Quarry, Blue Bell, which is in a Norian fluvial-lacustrine claystone in the Lockatong Formation of Pennsylvania.

It was considered a nomen dubium by Haubold (1971); it was considered a nomen dubium by Klein and Lucas (2021).
Dahutherium was named by Montenat (1968). Its type is Dahutherium agilis.

It was considered a nomen nudum by Klein and Lucas (2021).

It was assigned to Pseudosuchia by Lapparent and Montenat (1967); to Batrachopodidae by Haubold (1969); to Batrachopodidae by Haubold (1971), Demathieu and Haubold (1972), Gand (1974); and to Protosuchia by Haubold (1974).

Synonymy list

Year	Name and author
1853	Chelonichnium Schimper p. 10
1906	Chirotherium ibericus Navás
1935	Chirotheriidae Abel p. 67
1941	Chirotherium angustum Huene p. 5
1941	Thecodontichnus fucinii Huene p. 5–6
1948	Chirotheriidae Peabody p. 340
1948	Chirotherium angustum Peabody p. 347
1952	Chirotheriidae Bock p. 410
1952	Chirotherium lulli Bock pp. 415-416
1952	Chirotherium huberi Bock pp. 416-417 figs. Pl. 47:1
1952	Chirotherium wondrai Heller p. 131 fig. 1
1952	Chirotherium angustum Heller p. 134
1954	Chirotherium lulli Baird p. 167
1954	Chirotherium angustum Baird p. 174
1954	Chirotherium wondrai Baird p. 174
1957	Chirotheriidae Baird p. 473
1957	Chirotherium lulli Baird p. 482
1957	Chirotherium wondrai Baird p. 482
1958	Chirotherium angustum Kuhn p. 21
1958	Chirotherium huberi Kuhn p. 21
1958	Chirotherium lulli Kuhn p. 21
1958	Chirotherium wondrai Kuhn p. 21
1958	Thecodontichnus fucinii Kuhn p. 23
1959	Chirotherium lulli Bock p. 132
1959	Chirotherium wondrai Schmidt p. 103
1959	Thecodontichnus fucinii Schmidt p. 105
1959	Chirotherium lulli Schmidt p. 106
1963	Chirotheriidae Kuhn p. 67
1963	Chirotherium lulli Kuhn p. 72
1963	Chirotherium wondrai Kuhn p. 74
1963	Thecodontichnus fucinii Kuhn p. 88
1964	Shimmelia Casamiquela p. 139
1964	Shimmelia chirotheroides Casamiquela pp. 139-140 figs. Pl. XV, XVI
1964	Chirotheriidae Kuhn p. 320
1966	Chirotheriidae Kuhn p. 75
1967	Dahutherium Lapparent and Montenat p. 33
1967	Swinnertonichnus Sarjeant p. 332
1967	Swinnertonichnus mapperleyensis Sarjeant p. 333 figs. 3; Pl. 14
1968	Dahutherium Montenat p. 373
1968	Dahutherium agilis Montenat p. 373 figs. Pl. 1"4; Pl. II:8–9
1969	Chirotheriidae Haubold p. 91
1969	Dahutherium Haubold p. 97
1970	Paratrisauropus Ellenberger p. 345
1970	Deuterosauropodopus Ellenberger p. 346
1970	Deuterosauropodopus minor Ellenberger p. 346
1971	Brachychirotherium tintanti Demathieu p. 814
1971	Chirotheriidae Haubold p. 52
1971	Chirotherium lulli Haubold p. 55
1971	Chirotherium wondrai Haubold p. 55
1971	Brachychirotherium tintanti Haubold p. 57
1971	Dahutherium Haubold p. 63
1971	Swinnertonichnus Haubold p. 69
1971	Swinnertonichnus mapperleyensis Haubold p. 69
1971	Shimmelia chirotheroides Haubold p. 101
1972	Brachychirotherium tintanti Demathieu and Haubold p. 805
1972	Chirotherium wondrai Demathieu and Haubold p. 805
1972	Dahutherium Demathieu and Haubold p. 805
1972	Dahutherium agilis Demathieu and Haubold p. 805
1972	Swinnertonichnius Demathieu and Haubold p. 806
1972	Swinnertonichnius mapperleyensis Demathieu and Haubold p. 806
1972	Deuterosauropodopus minor Ellenberger p. 90
1972	Deuterosauropodopus Ellenberger p. 152
1972	Paratrisauropus Ellenberger p. 152
1974	Paratrisauropus Ellenberger p. 17
1974	Chirotheriidae Gand p. 8
1974	Dahutherium Gand p. 9
1974	Chirotheriidae Haubold p. 39
1974	Dahutherium Haubold p. 40
1974	Swinnertonichnus Haubold p. 43
1974	Swinnertonichnus mapperleyensis Haubold p. 115
1974	Brachychirotherium tintanti Haubold p. 117
1974	Dahutherium agilis Haubold p. 117
1974	Chirotherium lulli Haubold p. 122
1974	Chirotherium wondrai Haubold p. 122
1975	Shimmelia Casamiquela p. 558
1975	Shimmelia chirotheroides Casamiquela p. 558
1975	Swinnertonichnus Sarjeant p. 304
1975	Swinnertonichnus mapperleyensis Sarjeant p. 307
1976	Chirotherium wondrai Courel and Demathieu p. 204
1979	Chirotherium catalaunicum Casanovas-Cladellas et al.
1980	Chirotherium angustum Tongiorgi p. 81
1980	Thecodontichnus fucinii Tongiorgi p. 81
1981	Pseudochirotherium Biron and Dutuit p. 409
1981	Pseudochirotherium oukaimedensis Biron and Dutuit p. 409 figs. fig. 6, Pl. IIM
1981	Pentichnus Biron and Dutuit p. 410
1981	Pentichnus largus Biron and Dutuit p. 410 figs. fig. 7, Pl. IIH, I
1981	Shimmelia chirotheroides Leonardi p. 931
1982	Deuterosauropodopus sedunensis Demathieu and Weidmann p. 746 figs. 13B, 14A
1982	Pachysaurichnium Demathieu and Weidmann p. 749
1982	Pachysaurichnium emossonense Demathieu and Weidmann p. 749 figs. 13C, 14B
1984	Brachychirotherium minor Olsen and Galton p. 96 fig. 3
1986	Chirotherium huberi Conrad and Lockley p. 13
1986	Chirotherium lulli Olsen and Baird p. 21
1987	Chirotherium huberi Conrad et al. p. 131
2003	Paratrisauropus Romero Molina et al. p. 24
2003	Chirotherium lulli Szajna and Hartline p. 268
2007	Chirotheridae Calvo p. 318
2007	Shimmelia Calvo p. 318
2007	Shimmelia chirotheroides Calvo p. 318–319
2007	Brachychirotherium tintanti Gand et al. p. 12
2007	Dahutherium agilis Gand et al. p. 20
2007	Paratrisauropus Irmis et al. p. 16
2012	Brachychirotherium tintanti Gand et al. p. 68
2025	Dahutherium agilis Foster et al. p. 315

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Deuterostomia
phylum	Chordata	Haeckel 1874
subphylum	Vertebrata
superclass	Gnathostomata
—	Osteichthyes	()
subclass	Sarcopterygii	()
subclass	Dipnotetrapodomorpha	(Nelson 2006)
subclass	Tetrapodomorpha	()
—	Tetrapoda
—	Reptiliomorpha
—	Anthracosauria
subclass	Amphibiosauria	Kuhn 1967

Rank	Name	Author
—	Cotylosauria	()
—	Amniota
—	Sauropsida
class	Reptilia
subclass	Eureptilia	()
—	Romeriida
—	Diapsida	()
—	Archosauromorpha	(Huene 1946)
—	Crocopoda
—	Archosauriformes	Gauthier 1986
—	Eucrocopoda
—	Archosauria	()
—	Pseudosuchia	(Zittel 1890)
family	Chirotheriidae	Abel 1935
family	Chirotheriidae	Abel 1935

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Fm. †Chirotheriidae Abel 1935

show all | hide all

G. †Brachychirotherium Beurlen 1950

†Brachychirotherium circaparvum Demathieu 1971

†Brachychirotherium eyermani Baird 1957

†Brachychirotherium gallicum Willruth 1917

†Brachychirotherium harrasense Haubold 1967

†Brachychirotherium hauboldi Ptaszynski 1990

†Brachychirotherium hessei Soergel 1925

†Brachychirotherium kalkowensis Niedzwiedzki and Ptaszynski 2007

†Brachychirotherium kuhni Demathieu and Haubold 1982

†Brachychirotherium lorteti Haubold 1970

†Brachychirotherium pachydactylum Demathieu and Gand 1973

†Brachychirotherium paraparvum Demathieu and Oosterink 1988

†Brachychirotherium parvum Hitchcock 1889

Invalid names: Agrestipus hottoni Weems 1987 [synonym], Chirotherium copei Bock 1952 [synonym]

†Brachychirotherium praeparvum Haubold 1967

†Brachychirotherium thuringiacum Rühle von Lilienstern 1938

Invalid names: Brachychirotherium coburgense Aumann 1957 [synonym], Brachychirotherium dulcis Ellenberger 1970 [synonym], Brachychirotherium hassfurtense Beurlen 1950 [synonym]

†Brachychirotherium wiorense Ptaszynski 2000

G. †Chirotherium Kaup 1835

†Chirotherium atlensis Biron and Dutuit 1981

†Chirotherium barthii Kaup 1835

Invalid names: Chirotherium bairdi Reig 1961 [synonym], Chirotherium gallicum Willruth 1917 [synonym], Chirotherium higuerensis Rusconi 1952 [synonym], Chirotherium kaupii Owen 1842 [synonym], Chirotherium majus Sickler 1836 [synonym], Chirotherium pfeifferi Soergel 1925 [synonym], Saurichnites auraensis Kirchner 1927 [synonym], Saurichnites gambaensis Schuster 1936 [synonym]

†Chirotherium culmbachense Soergel 1925

†Chirotherium ferox Demathieu 1966

†Chirotherium herculis Egerton 1839

†Chirotherium heterodactylum King 1844

†Chirotherium ischigualastianum Huene 1931

†Chirotherium ladinicus Avanzini and Wachtler 2012

†Chirotherium mediterraneum Demathieu and Durand 1991

†Chirotherium peabodyi Faber 1958

†Chirotherium postchirotherioides Rehnelt 1950

†Chirotherium praeparvum Haubold 1967

†Chirotherium reiteri Moore 1873

†Chirotherium rex Peabody 1948

Invalid names: Chirotherium moquiensis Peabody 1948 [synonym]

†Chirotherium sickleri Kaup 1835

Invalid names: Chirotherium beasleyi Nopcsa 1923 [synonym], Chirotherium bipes Berthold 1835 [synonym], Chirotherium bornemanni Willruth 1917 [synonym], Chirotherium minus Sickler 1836 [synonym]

†Chirotherium soergeli Haubold 1969

†Chirotherium storetonense Morton 1863

†Chirotherium swinnertoni Sarjeant 1970

†Chirotherium vorbachi Kirchner 1927

Invalid names: Chirosaurus Kaup 1835 [synonym], Krokodilipus Nopcsa 1923 [synonym], Quadridigitus Biron and Dutuit 1981 [synonym], Saurichnites Geinitz 1861 [synonym], Sphingopus Demathieu 1966 [synonym]

G. †Isochirotherium Haubold 1971

†Isochirotherium bipedale Abel 1935

†Isochirotherium circademathieui Gand 1979

†Isochirotherium coltoni Peabody 1948

†Isochirotherium comblei Gand 1979

†Isochirotherium coureli Demathieu 1970

†Isochirotherium delicatum Courel and Demathieu 1976

†Isochirotherium demathieui Haubold 1970

†Isochirotherium felenci Courel and Demathieu 1976

†Isochirotherium gardettensis Petti et al. 2020

†Isochirotherium gierlinskii Ptaszynski 2000

†Isochirotherium herculis Egerton 1839

†Isochirotherium hessbergense Haubold 1970

†Isochirotherium jenense Haubold 1970

†Isochirotherium lomasi Baird 1954

†Isochirotherium marshalli Peabody 1948

†Isochirotherium sanctacrucense Ptaszynski 1990

†Isochirotherium soergeli Haubold 1969

G. †Parasynaptichnium Mietto 1987

G. †Protochirotherium Fichter and Kunz 2004

†Protochirotherium wolfhagense Fichter and Kunz 2004

Invalid names: Palaeochirotherium Fichter and Kunz 2011 [synonym]

G. †Sauropodopus Ellenberger 1970

†Sauropodopus antiquus Ellenberger 1970

Invalid names: Deuterosauropodopus major Ellenberger 1970 [synonym]

G. †Synaptichnium Nopcsa 1923

†Synaptichnium argantobrivense Demathieu and Gand 1981

†Synaptichnium cameronensis Peabody 1948

†Synaptichnium chirotherioides Ptaszynski 1990

†Synaptichnium diabloensis Peabody 1948

†Synaptichnium hildburghausense Lilienstern 1939

†Synaptichnium kotanskii Ptaszynski 2000

†Synaptichnium motutongense Ellenberger 1974

†Synaptichnium priscum Demathieu 1970

†Synaptichnium pseudosuchoides Nopcsa 1923

†Synaptichnium senkowiczowae Niedzwiedzki and Ptaszynski 2007

Invalid names: Brachychirotherium tintanti Demathieu 1971 [nomen dubium], Chelonichnium Schimper 1853 [nomen dubium], Chirotherium angustum Huene 1941 [nomen dubium], Chirotherium catalaunicum Casanovas-Cladellas et al. 1979 [nomen dubium], Chirotherium huberi Bock 1952 [nomen dubium], Chirotherium ibericum Navás 1906 [nomen dubium], Chirotherium lulli Bock 1952 [nomen dubium], Chirotherium wondrai Heller 1952 [nomen dubium], Dahutherium Montenat 1968 [nomen nudum], Deuterosauropodopus Ellenberger 1970 [nomen dubium], Deuterosauropodopus minor Ellenberger 1970 [nomen dubium], Pachysaurichnium Demathieu and Weidmann 1982 [nomen dubium], Paratrisauropus Ellenberger 1970 [nomen dubium], Pentichnus Biron and Dutuit 1981 [nomen dubium], Pseudochirotherium Biron and Dutuit 1981 [nomen dubium], Shimmelia Casamiquela 1964 [nomen dubium], Swinnertonichnus Sarjeant 1967 [nomen dubium], Thecodontichnus fucinii Huene 1941 [nomen dubium]

Diagnosis

Reference	Diagnosis
F. E. Peabody 1948	Relatively narrow, quadrupedal trackways indicating the normal tetrapod walking gait; in the walking gait a small pentadactyl manus impression regularly occurs immediately in front of, but never overlapped by a much larger, pentadactyl pes which generally resembles a reversed human hand. Manus and pes are digitigrade, and in large forms the pes tends to be plantigrade; digits I-IV point more or less forward, manus digit IV is always shorter than III, digit V is divergent and well developed as a crude prop for the foot; strong elaws are indicated on digits I-IV, infrequently on V of manus and pes, claw on III being largest; the footprints mayor may not show specialized metatarsal and phalangeal pads. Clear impressions often show a granular or beaded skin surface. Reptiles represented by trackways probably dinosaur-like in form, and in body propor tions showed arrested tendency toward bipedalism. In smallest species pes only 3 cm.

Measurements

No measurements are available

Composition:

phosphaticsubp

Environment:

terrestrialsubc

Locomotion:

actively mobilec

Diet:

carnivoresubo

Created:

2009-01-03 20:38:45

Modified:

2009-01-03 22:38:45

Source: subo = suborder, subc = subclass, c = class, subp = subphylum

References: Benton 1983, Hendy et al. 2009, Carroll 1988

Age range: base of the Artinskian to the top of the Turonian or 290.10000 to 89.80000 Ma

Collections (216 total)

Time interval	Ma	Country or state	Original ID and collection number
Oldest occurrences
Artinskian - Capitanian	290.1 - 259.51	Poland (Lower Silesia)	Saurichnites divaricatus, Saurichnites gracilis (223853) Saurichnites leisnerianus (187930)
Buntsandstein	259.51 - 251.902	Germany (Bavaria)	Chirotherium vorbachi (205801) Saurichnites auraensis (205802) Saurichnites auraensis, Chirotherium vorbachi (205800)
Wuchiapingian - Olenekian	259.51 - 246.7	Argentina (Río Negro)	Shimmelia chirotheroides (53684)
Induan	251.902 - 249.9	China (Guizhou)	Chirotherium sp. (60516)
Early/Lower Triassic	251.902 - 246.7	Germany (Rhineland-Palatinate)	Chirotherium barthii (222574)
Early/Lower Triassic	251.902 - 246.7	United Kingdom (England)	Chirotherium herculis (164932) Synaptichnium sp., Chirotherium storetonense, Chirotherium minus, Chirotherium vorbachi, Isochirotherium lomasi, Isochirotherium bipedale, Chirotherium sp., Chirotherium minus (164930)
Early/Lower Triassic	251.902 - 246.7	Morocco (Marrakech-Safi)	Synaptichnium sp., Protochirotherium sp. (199897 199898 199899 199900)
Early/Lower Triassic	251.902 - 246.7	Italy (Cuneo)	Chirotherium sp., Isochirotherium sp., Isochirotherium gardettensis (224578)
Early/Lower Triassic	251.902 - 246.7	Niger (Agadez)	Chirotherium sp. (200662)
Early/Lower Triassic	251.902 - 246.7	Spain (Islas Baleares)	Chirotherium sp. (95349)
Youngest occurrences
Time interval	Ma	Country or state	PBDB collection number
Norian - Rhaetian	227.3 - 201.4	Lesotho (Mohale's Hoek)	Deuterosauropodopus major (56020)
Norian - Rhaetian	227.3 - 201.4	USA (Utah)	Brachychirotherium sp. (57012 83555 193017 193018 193020 193244)
Rhaetian	205.7 - 201.4	USA (Utah)	Brachychirotherium sp. (192896 192897 192915 192916 192941)
Rhaetian	205.7 - 201.4	France (Pays de la Loire)	Chirotherium sp. (70801)
Rhaetian	205.7 - 201.4	USA (Colorado)	Brachychirotherium sp. (192942 193012)
Rhaetian	205.7 - 201.4	USA (Pennsylvania)	Brachychirotherium sp. (62767 132244) Brachychirotherium sp., Chirotherium lulli (62766)
Hettangian - Sinemurian	201.4 - 192.9	Lesotho (Quthing)	Pseudotetrasauropus dulcis (56003) Synaptichnium motutongense (84072)
Sinemurian	199.5 - 192.9	Lesotho (Leribe)	Chirotherium sp., Dinosauria indet. (56014)
Late/Upper Kimmeridgian	152.21 - 149.2	France (Ain)	Chelonichnium cerinense (59351)
Turonian	93.9 - 89.8	Egypt (Aswan)	Chirotherium sp. (138680)