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Buccinidae (true whelk)

Gastropoda - Neogastropoda - Buccinidae

Taxonomy
Buccinidae was named by Rafinesque (1815) [m. kosnik taxon no: 5368]. It is extant. Its type is Buccinum.

It was assigned to Rachiglossa by Martin (1904); to Rachigdossa by Olsson (1931); to Rhachiglossa by Grant and IV and Gale (1931); to Gastropoda by Turner (1938); to Prosobranchiata by Harris and Palmer (1947); to Muricacea by Squires (1984); to Neogastropoda by Addicott (1970), Givens and Kennedy (1976), Vermeij (1997), Squires (1999), Squires and Saul (2000), Wienrich (2001); to Buccinacea by Rivera (1957), Sohl (1964), Hickman (1980), Squires et al. (1989), Stilwell and Zinsmeister (1992), Chiesa et al. (1995), Harzhauser (2002); to Muricoidea by Ponder and Warén (1988), Beesley (1998), Todd (2001), Squires and Saul (2001), Squires and Saul (2003); and to Buccinoidea by Jeffery (2003), Stilwell et al. (2004), Harzhauser (2004), Bouchet et al. (2005), Squires and Saul (2006), Landau and Marques da Silva (2010), DeVries (2016), Amano and Oleinik (2016), Bouchet et al. (2017), Kantor et al. (2022), Schnetler et al. (2024).

Subtaxa
Accidenticulabrum, Agasoma, Aliofusus, Amarophos, Anomalisipho, Aquilofusus, Austrocominella, Babylonia, Bailya, Beringiinae (syn. Metajapelioninae), Bolis, Brachysphingus, Buccininae, Buccinopsis (syn. Ripleyia, Seminola), Buccitriton, Burnupena, Cantharus, Cymatophos, Cyrtochetus, Dipsaccus, Donovaniinae, Editharus, Egotistica, Eobuccinella, Euthrenopsis, Hanetia, Hindsia, Janiopsis, Kaiparanura, Kergipenion, Khetella, Lacinia, Laevibuccinum, Latrunculus, Loochooia, Metaphos, Metula (syn. Antemetula), Misifulgur, Monoptygma, Monostiolum, Nawenia, Neosconsia, Neptuneinae (syn. Chrysodominae), Nicema, Odontobasis, Ornopsis, Pangoa, Paracominia, Parancistrolepidinae (syn. Ancistrolepidinae, Brevisiphoniinae), Parkeristoma, Parvisipho, Pegocomptus, Perunassa, Pisaninae, Plicifusus (syn. Quasisipho), Pomahakia, Pseudofulgur, Pseudolivinae, Pseudoneptunea, Ptychosalpinx, Sagenella, Scalaspira, Searlesia, Siphonaliinae, Stantonella, Suessionia (syn. Fusopsis), Suessonia, Terebrifusus, Tritonidea, Trominina, Truncaria, Varicosipho, Volutopsiinae, Wrigleya, Zaglenum, Zelandiella
Species lacking formal opinion data
B. montensis
Entered
by M. Kosnik on 2002-07-27; modified by P. Novack-Gottshall on 2026-06-26

Synonyms
Synonymy list
YearName and author
1815Buccinidae Rafinesque
1865Sagenella Conrad
1904Buccinidae Martin p. 183
1931Buccinidae Grant, IV and Gale p. 667
1931Buccinidae Olsson p. 99
1938Buccinidae Turner p. 77
1947Buccinidae Harris and Palmer p. 353
1957Buccinidae Rivera p. 181
1964Buccinidae Sohl p. 372
1970Buccinidae Addicott p. 94
1976Buccinidae Givens and Kennedy p. 970
1980Buccinidae Hickman p. 55
1984Buccinidae Squires p. 30
1988Buccinidae Ponder and Warén p. 305
1989Buccinidae Squires et al. p. 212
1992Buccinidae Stilwell and Zinsmeister p. 121
1995Buccinidae Chiesa et al. p. 56
1997Buccinidae Vermeij p. 47
1998Buccinidae Beesley p. 825
1999Buccinidae Squires p. 19
2000Buccinidae Squires and Saul p. 118
2001Buccinidae Squires and Saul p. 57
2001Buccinidae Todd
2001Buccinidae Wienrich p. 483
2002Buccinidae Harzhauser p. 99
2002Sagenella Sepkoski
2003Buccinidae Jeffery
2003Buccinidae Squires and Saul p. 152
2004Buccinidae Harzhauser p. 136
2004Buccinidae Stilwell et al. p. 38
2005Buccinidae Bouchet et al. p. 254
2006Buccinidae Squires and Saul p. 70
2010Buccinidae Landau and Marques da Silva p. 70
2016Buccinidae Amano and Oleinik p. 159
2016Buccinidae DeVries p. 103
2017Buccinidae Bouchet et al. p. 349
2022Buccinidae Kantor et al.
2024Buccinidae Schnetler et al. p. 16

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—Spiralia
superphylumLophotrochozoa
phylumMollusca
classGastropoda
RankNameAuthor
subclassCaenogastropoda(Cox 1959)
—Sorbeoconcha(Ponder and Lindberg 1997)
—Hypsogastropoda(Ponder and Lindberg 1997)
superorderLatrogastropodaRiedel 2000
orderNeogastropodaThiele 1929
superfamilyBuccinoidea(Rafinesque 1815)
familyBuccinidaeRafinesque 1815
familyBuccinidaeRafinesque 1815

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Fm. Buccinidae Rafinesque 1815 [true whelk]
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G. Accidenticulabrum Vermeij 2000
+
G. †Agasoma Gabb 1869
+
Subg. †Agasoma (Trophosycon) Cooper 1894
†Agasoma acuminatum Anderson and Martin 1914
†Agasoma andersoni Wiedey 1928
+
Invalid names: Agasoma oregonense Anderson and Martin 1914 [replaced]
G. †Aliofusus Stephenson 1941
+
†Aliofusus stamineus Sohl 1964
G. †Amarophos Woodring 1964
+
†Amarophos bothrus Woodring 1964
†Amarophos dentalis Olsson 1922
G. Anomalisipho Dautzenberg and Fischer 1912
G. †Aquilofusus Kautsky 1925
+
†Aquilofusus elegantulus Philippi 1843
†Aquilofusus waeli Nyst 1852
G. †Austrocominella Ihering 1907
+
G. Babylonia Schlüter 1838
+
Subg. Babylonia (Peridipsaccus) Rovereto 1900
+
Babylonia japonica Reeve 1842
†Babylonia kozaiensis Nomura 1939
†Babylonia pangkaensis Martin 1895
†Babylonia spirata Linnaeus 1758
Babylonia zeylanica Lamarck 1822
G. Bailya Smith 1944
+
Bailya intricata Dall 1884
Subfm. Beringiinae Golikov and Starobogatov 1975
+
G. †Beringion Habe and Ito 1965
G. Beringius Dall 1886
+
Subg. Beringius (Beringius) Dall 1886
+
G. Metajapelion Goryachev 1987
G. †Neoberingius Habe and Ito 1965
Invalid names: Metajapelioninae Goryachev 1987 [synonym]
G. †Bolis Gardner 1939
G. †Brachysphingus Gabb 1869
+
†Brachysphingus clarki Weaver 1912
†Brachysphingus gibbosus Nelson 1925
†Brachysphingus mammilatus Clark and Woodford 1927
+
Invalid names: Brachysphingus gabbi Stewart 1927 [synonym], Brachysphingus liratus Gabb 1864 [replaced]
Subfm. Buccininae Rafinesque 1815
+
G. †Angistoma Sandberger 1861
G. †Austrobuccinum Stilwell and Zinsmeister 1992
+
†Austrobuccinum tuberosum Stilwell and Zinsmeister 1992
G. Buccinum Linnaeus 1758
+
Buccinum aciculatum Gmelin 1791
†Buccinum amoenum Conrad 1833
†Buccinum bistriatum Lamarck 1803
†Buccinum clavatulatum Lamarck 1803
Buccinum cyaneum Bruguière 1789
†Buccinum declive Wood 1879
†Buccinum decussatum Lamarck 1803
Buccinum digitale Dillwyn 1817
†Buccinum dubium Martin 1879
Buccinum finmarkianum Verkrüzen 1875
Buccinum glaciale Linnaeus 1761
†Buccinum jordani Hertlein 1925
†Buccinum junghuhni Martin 1879
Buccinum laevigatum Gmelin 1791
†Buccinum mississppiensis Conrad 1848
Buccinum modestum Powys 1835
†Buccinum nudum Wood 1879
Buccinum ochotense Middendorf 1848
Buccinum plectrum Stimpson 1865
Buccinum polare Gray 1839
+
Invalid names: Buccinum totteni Stimpson 1861 [synonym]
Buccinum scalariforme Möller 1842
+
Invalid names: Buccinum tenue Gray 1839 [replaced]
†Buccinum striatulum Lamarck 1803
†Buccinum strigillatum Dall 1891
+
Invalid names: Buccinum saundersi Martin 1914 [synonym]
†Buccinum tenebrosum Hancock 1846
+
Invalid names: Buccinum bogachielii Reagan 1909 [synonym]
†Buccinum terebrale Lamarck 1803
Buccinum undatum Linnaeus 1758
†Buccinum vinculum Hall and Meek 1855
Invalid names: Bathybuccinum Golikov and Sirenko 1988 [synonym], Madiella Wenz 1943 [synonym], Volutharpa Fisher 1855 [synonym]
G. †Coptochetus Cossmann 1889
G. †Corneobuccinum Golikov and Gulbin 1977
G. †Eripachya Gabb 1869
+
†Eripachya jalama Squires and Saul 2003
†Eripachya ponderosa Gabb 1864
†Eripachya vaccina Saul and Popenoe 1993
G. †Euryochetus Cossmann 1896
G. †Exilioidea Grant, IV and Gale 1931
+
Exilioidea rectirostris Carpenter 1864
G. Fax Iredale 1925
+
†Fax cominelloides Tate 1888
G. †Kapala Ponder 1982
Tr. Liomesini Fischer 1884
+
G. Liomesus Stimpson 1865
+
†Liomesus africanus Newton 1922
†Liomesus dalei Sowerby 1825
†Liomesus sulculatus Dall 1909
Tritonium (Buccinum) ooides Middendorff 1848
G. †Plicibuccinum Golikov and Gulbin 1977
G. †Pseudofax Finlay and Marwick 1937
+
†Pseudofax costellatus Griffin and Hünicken 1994
†Pseudofax ordinarius Marshall 1917
†Pseudofax paucus Stilwell et al. 2004
†Pseudofax solitaria Chiesa et al. 1995
†Pseudofax suroinflatus Stilwell and Zinsmeister 1992
†Pseudofax weddellensis Stilwell and Zinsmeister 1992
G. †Strombinophos Pilsbry and Olsson 1941
+
†Strombinophos loripanus Pilsbry and Olsson 1941
†Strombinophos maxwelli Olsson and Harbison 1953
†Strombinophos mimicus Woodring 1964
†Strombinophos vadosus Gardner 1926
G. Thysanobuccinum Golikov and Gulbin 1980
+
Thysanobuccinum tunicatum Golikov and Gulbin 1977
Invalid names: Buccinini Rafinesque 1815 [empty], Buccinulini Finlay 1928 [empty], Colini Gray 1857 [empty]
G. †Buccinopsis Conrad 1857
+
†Buccinopsis crassa Wade 1917
†Buccinopsis solida Wade 1917
+
Invalid names: Ripleyia Cossmann 1920 [synonym], Seminola Wade 1918 [synonym]
+
†Buccitriton sagenum Conrad 1833
+
Invalid names: Buccinum (Nassa) cancellatum Lea 1833 [synonym]
†Buccitriton texanum Gabb 1861
+
Invalid names: Buccitriton altum Conrad 1965 [synonym]
G. †Burnupena Iredale 1918
+
†Burnupena aestus Kensley and Pether 1986
†Burnupena rogersi Kensley and Pether 1986
G. Cantharus Röding 1798
+
Subg. †Cantharus (Calicantharus) Clark 1938
+
†Calicantharus branneri Clark and Arnold 1923
†Calicantharus dalli Clark 1918
Calicantharus fortis Carpenter 1866
+
Invalid names: Calicantharus angulatus Arnold 1907 [synonym]
+
Invalid names: Cantharus ashleyi Carson 1925 [synonym], Cantharus breaensis Carson 1925 [synonym], Cantharus elsmerensis Carson 1925 [synonym], Chrysodomus arnoldi Rivers 1904 [synonym]
†Calicantharus kernensis Anderson and Martin 1914
Subg. Cantharus (Cantharus) Röding 1798
Subg. †Cantharus (Endopachychilus) Cossman 1889
+
†Endopachychilus blakianus Anderson and Hanna 1925
Subg. †Cantharus (Zeapollia) Finlay 1926
+
†Zeapollia brevis Tate 1888
†Zeapollia purpuroides Johnston 1879
†Cantharus andoi Matsubara et al. 2010
†Cantharus brongniartianus Orbigny 1850
†Cantharus bucklandi d'Archiac 1850
†Cantharus cowlitzensis Turner 1938
†Cantharus erythrostoma Reeve 1849
Cantharus lugubris Adams 1852 [ribbed cantharus]
†Cantharus luliana Martin 1884
†Cantharus martiniana Noetling 1901
†Cantharus perrini Dickerson 1915
†Cantharus polygona Lamarck 1803
†Cantharus varicosus Tate 1888
G. Cymatophos Pilsbry and Olsson 1941
+
†Cymatophos acolus Woodring 1964
†Cymatophos beteyensis Olsson 1922
Cymatophos crassus Hinds 1843
†Cymatophos galerus Pilsbry and Olsson 1941
†Cymatophos glareosus Jung 1971
†Cymatophos hodsoni Olsson 1932
†Cymatophos paraguanensis Hodson and Hodson 1931
†Cymatophos subsemicostatus Brown and Pilsbry 1911
†Cymatophos tuberaensis Anderson 1929
†Cymatophos turbacoensis Anderson 1929
†Cymatophos veatchi Olsson 1922
+
†Cymatophos veatchi erymnus Woodring 1964
Invalid names: Phos (Phos) buchivacoensis Hodson 1931 [synonym]
G. †Cyrtochetus Cossmann 1899
+
Subg. †Cyrtochetus (Cyrtochetus) Cossmann 1899
+
†Cyrtochetus chiraensis Olsson 1931
G. Dipsaccus Deshayes 1832
G. †Editharus Vermeij 2001
+
†Editharus angulilabris Vermeij 2001
†Editharus berellensis de Laubrière and Carez 1880
†Editharus ditropis Bayan 1870
†Editharus dumasi Cossmann 1897
†Editharus labiatus Sowerby 1823
†Editharus marcellini Cossman 1902
†Editharus polygonus Lamarck 1803
†Editharus roncanus d'Orbigny 1850
+
Invalid names: Fusus rarispinatus Schauroth 1965 [synonym]
†Editharus vasseuri Cossmann 1897
G. †Egotistica Marwick 1934
+
†Egotistica scirrhoma Marwick 1934
G. †Eobuccinella Stilwell and Zinsmeister 1992
+
†Eobuccinella brucei Stilwell and Zinsmeister 1992
G. Euthrenopsis Powell 1929
G. Hanetia Jousseaume 1880
+
†Hanetia boggsi Pilsbry and Olsson 1941
†Hanetia cymioides Pilsbry and Olsson 1941
†Hanetia dalli Brown and Pilsbry 1911
+
†Hanetia dalli protera Woodring 1964
Invalid names: Coralliophila gatunensis Toula 1911 [synonym], Coralliophila incerta Toula 1911 [synonym]
†Hanetia ecuadorensis Pilsbry and Olsson 1941
†Hanetia mengeana Dall 1890
G. Hindsia Adams 1853
+
Subg. Hindsia (Nihonophos) MacNeil 1960
+
†Hindsia granosa Sowerby 1840
†Hindsia mekranica Vredenburg 1925
†Hindsia neastriatula Noetling 1901
†Hindsia pardalis Noetling 1895
Hindsia varicifera Adams 1853
G. †Janiopsis Rovereto 1899
+
†Janiopsis californiensis Palmer 1923
†Janiopsis ewekoroensis Adegoke 1977
†Janiopsis nigeriensis Newton 1922
†Janiopsis sanmarcoensis Givens and Kennedy 1976
G. †Kaiparanura Laws 1944
G. †Kergipenion Fletcher 1938
+
†Kergipenion dubia Fletcher 1938
G. †Khetella Beisel 1977
+
†Khetella trautscholdi Blagovetshenskiy and Shumilkin 2006
G. †Lacinia Conrad 1833
+
†Lacinia alveata Conrad 1833
†Lacinia claibornensis Palmer 1937
†Lacinia dubia Merriam and Turner 1937
†Lacinia minbuensis Vredenburg 1921
G. †Laevibuccinum Conrad 1865
G. Latrunculus Gray 1847
G. Loochooia MacNeil 1960
G. Metaphos Olsson 1964
G. Metula Adams and Adams 1853
+
Subg. Metula (Metula) Adams and Adams 1853
Metula amosi Vanatta 1913
†Metula brazosensis Johnson 1899
†Metula cancellata Gabb 1873
†Metula gabbi Brown and Pilsbry 1911
†Metula gentilicia Harris and Palmer 1947
†Metula gracilis Johnson 1899
†Metula harrisi Dickerson 1916
†Metula harrisi Olsson 1922
Metula hindsii Adams and Adams 1853
†Metula johnsoni Vaughan 1896
†Metula limonensis Olsson 1922
†Metula martini Vredenburg 1925
†Metula peruviana Olsson 1931
†Metula roberti Olsson 1967
†Metula subgracilis Johnson 1899
†Metula taylorensis Mansfield 1940
Invalid names: Antemetula Rehder 1943 [synonym]
G. †Misifulgur DeVries 2016
+
†Misifulgur dockeryi DeVries 2016
†Misifulgur macneili DeVries 2016
†Misifulgur montemarensis DeVries 2016
G. †Monoptygma Lea 1833
G. Monostiolum Dall 1904
+
†Monostiolum petiti Olsson 1967
†Monostiolum thomasi Olsson 1967
G. †Nawenia Ladd 1977
+
†Nawenia bartholomewi Ladd 1977
G. †Neosconsia Olsson 1964
+
†Neosconsia ecuadoriana Olsson 1964
Subfm. Neptuneinae Stimpson 1865
+
G. †Callioloncha Lus 1978
G. †Helicofusus Dall 1916
+
†Latisipho riversi Martin 1914
G. Neptunea Röding 1798
+
Subg. Neptunea (Neptunea) Röding 1798
+
+
Invalid names: Cymatium pacificum Dall 1909 [synonym]
Invalid names: Neptunea (Neptunea) soluta cordata Dall 1919 [nomen nudum]
Subg. Neptunea (Sulcosipho) Dall 1916
+
†Neptunea (Sulcosipho) eatoni Grant, IV and Quayle 1934
+
Invalid names: Chrysodomus (Ancistrolepis) unicus Pilsbry 1905 [synonym]
+
Invalid names: Chrysodomus gettysburgensis Reagan 1909 [synonym]
Neptunea (Sulcosipho) tabulata Baird 1863 [tabled whelk]
+
Invalid names: Buccinum stocki Howe 1922 [synonym], Chrysodomus bairdi Dall 1909 [synonym]
†Neptunea alaquaensis Mansfield 1935
†Neptunea antarctocosta Stilwell and Zinsmeister 1992
Neptunea despecta Linnaeus 1758
†Neptunea diminuta Etherington 1931
†Neptunea gracilis Gabb 1864
Neptunea ithia Dall 1891
Neptunea lyrata Gmelin 1791
†Neptunea modestoidea Takeda 1953
†Neptunea stonei Pilsbry 1893
Invalid names: Chrysodomus Swainson 1840 [synonym]
G. †Paracallioloncha Lus 1989
G. †Tacita Lus 1971
Invalid names: Chrysodominae Dall 1870 [synonym]
G. †Nicema Woodring 1964
+
†Nicema amara Woodring 1964
†Nicema guttifera Grzybowski 1899
†Nicema predistorta Marks 1951
G. †Odontobasis Meek 1876
+
†Odontobasis australis Wade 1926
†Odontobasis sulcata Sohl 1964
G. †Ornopsis Wade 1916
+
†Ornopsis dysis Squires and Saul 2006
G. †Pangoa Marwick 1931
+
†Pangoa mira Marwick 1931
G. †Paracominia Powell and Bartrum 1929
Subfm. †Parancistrolepidinae Habe and Sato 1972
+
G. Ancistrolepis Dall 1895
+
†Ancistrolepis carolineae Squires 1984
†Ancistrolepis clarki Tegland 1933
†Ancistrolepis landesi Tegland 1933
†Ancistrolepis packardi Durham 1944
G. †Bathyancistrolepis Habe and Ito 1968
+
†Bathyancistrolepis mikasaensis Amano and Oleinik 2016
G. Clinopegma Grant and Gale 1931
G. Japelion Dall 1916
G. †Neancistrolepis Habe and Sato 1972
G. †Parancistrolepis Azum 1965
G. Pseudoliomesus Habe and Sato 1973
G. Thalassoplanes Dall 1908
+
Invalid names: Brevisiphonia Lus 1973 [synonym]
Invalid names: Ancistrolepidinae Habe and Sato 1973 [synonym], Brevisiphoniinae Lus 1973 [synonym]
G. †Parkeristoma Adegoke 1977
+
†Parkeristoma guineensis Adegoke 1977
†Parkeristoma reymenti Adegoke 1977
G. †Parvisipho Cossmann 1889
+
†Parvisipho meganosensis Clark and Woodford 1927
G. †Pegocomptus Zinsmeister 1983
+
†Pegocomptus howardi Dickerson 1914
G. †Perunassa Olsson 1932
+
†Nassa (Perunassa) ecuadoriana Pilsbry and Olsson 1941
G. Plicifusus Dall 1902
+
Invalid names: Quasisipho Petrov 1982 [synonym]
G. †Pomahakia Finlay 1927
+
†Pomahakia aberrans Finlay 1927
G. †Pseudofulgur MacNeil and Dockery 1984
Subfm. Pseudolivinae Cossmann 1901
+
G. †Austrosphaera Camacho 1949
+
Invalid names: Seymourosphaera Oleinik and Zinsmeister 1996 [synonym]
G. †Benthobia Dall 1889
+
Invalid names: Nux Barnard 1960 [synonym]
G. †Fulmentum Fischer 1884
+
Invalid names: Sylvanocochlis Melvill 1903 [synonym]
G. †Hubachia Etayo-Serna 1979
+
†Hubachia ramirolobae Etayo-Serna 1979
G. †Hydrotribulus Wade 1916
+
†Hydrotribulus elegans Sohl 1964
†Hydrotribulus nodosus Wade 1916
G. †Luizia Douvillé 1933
G. †Naudoliva Kilburn 1989
G. †Popenoeum Squires et al. 1989
+
†Popenoeum brevis Doncieux 1908
†Popenoeum briarti Vincent 1928
†Popenoeum chavani Glibert 1973
†Popenoeum lacunosum Pacaud and Schnetler 1999
+
Invalid names: Pseudoliva robusta Briart et al. 1970 [synonym]
†Popenoeum maritimus Squires et al. 1989
†Popenoeum maritimus bajaensis Squires et al. 1989
†Popenoeum mutabilis Woods 1922
+
†Popenoeum prima Defrance 1827
†Popenoeum scalina Heilprin 1881
†Popenoeum subcostata Stoliczka 1867
G. Pseudoliva Swainson 1840
+
Subg. †Pseudoliva (Fusopsis) Ravn 1939
†Pseudoliva adelkei Adegoke 1977
†Pseudoliva binodosa Traub 1938
†Pseudoliva coronaria Olsson 1930
†Pseudoliva elegans Hislop 1859
†Pseudoliva funkeana Adegoke 1977
+
†Pseudoliva guppyi Mansfield 1925
†Pseudoliva inornata Dickerson 1915
†Pseudoliva laudunensis Defrance 1826
†Pseudoliva leutweini Bohm 1926
†Pseudoliva lutulenta Kensley and Pether 1986
†Pseudoliva modesta Olsson 1928
†Pseudoliva monilis Olsson 1928
†Pseudoliva packardi Van Winkle 1918
†Pseudoliva perspectiva Conrad 1860
†Pseudoliva praetermissa Pacaud and Schnetler 1999
†Pseudoliva reticulata Waring 1914
†Pseudoliva rogersi Adegoke 1977
†Pseudoliva serrata Traub 1938
†Pseudoliva thielei Bohm 1926
†Pseudoliva tuberculifera Conrad 1860
†Pseudoliva umpquaensis Turner 1938
†Pseudoliva unicarinata Aldrich 1886
†Pseudoliva vetusta Conrad 1833
+
†Pseudoliva vetusta forma Conrad 1833
G. †Ptychosyca Gabb 1876
+
†Ptychosyca inornata Gabb 1876
G. †Strepsidura Swainson 1840
+
Subg. †Strepsidura (Eamesidura) Adegoke 1977
+
†Strepsidura cossmanni Vredenburg 1923
†Strepsidura diabloensis Clark and Woodford 1927
†Strepsidura ficus Gabb 1864
†Strepsidura gracilis Riedel 1933
†Strepsidura howardi Dickerson 1916
†Strepsidura kerstingi Oppenheim 1915
†Strepsidura pachecoensis Stanton 1896
†Strepsidura pacifica Woods 1922
†Strepsidura polaris Stilwell et al. 2004
†Strepsidura reticulata Banerjee and Halder 2024
†Strepsidura spirata Newton 1922
Fm. †Strepturidae Cossmann 1901
+
G. Melapium Adams and A. 1853
+
†Melapium hawthornei Kensley and Pether 1986
G. †Sulcoliva Vermeij 1998
G. Zemira Adams and A. 1853
+
†Zemira praecursoria Tate 1888
Invalid names: Eburnopsis Tate 1889 [synonym]
G. Pseudoneptunea Kobelt 1882
+
†Pseudoneptunea burgasensis Karagyuleva 1964
†Pseudoneptunea costellata Grateloup 1847
†Pseudoneptunea curta Sowerby 1818
G. †Ptychosalpinx Gill 1867
+
†Ptychosalpinx altilis Conrad 1832
†Ptychosalpinx lienosa Conrad 1843
†Ptychosalpinx pustulosus Petuch 1988
G. †Scalaspira Conrad 1862
+
Subg. †Scalaspira (Scalaspira) Conrad 1862
+
†Scalaspira strumosa Conrad 1832
†Scalaspira vokesae Petuch 1988
G. Searlesia Harmer 1914
+
Kelletia (Searlesia) dira Reeve 1846 [dire whelk]
+
Invalid names: Fusus incisus Gould 1849 [synonym]
†Searlesia carlsoni Anderson and Martin 1914
†Searlesia dentifera Vermeij 1991
+
Invalid names: Searlesia ravni Schnetler and Beyer 1990 [synonym]
Searlesia dira Reeve 1846 [dire whelk]
†Searlesia dira miocenica Etherington 1931
†Searlesia mitgaui von Koenen 1867
Searlesia modesta Gould 1860
+
Invalid names: Searlesia fuscolabiata Smith 1874 [synonym]
†Searlesia olympicensis Durham 1944
Invalid names: Chrysodomus dirus meridiei Moody 1916 [invalid subgroup]
Subfm. Siphonaliinae Finlay 1928
+
G. Bayerius Olsson 1971
G. †Berylsma Iredale 1924
G. Japeuthria Iredale 1918
G. †Mohnia Friele 1878
G. Pararetifusus Kosuge 1967
G. Phaenomenella Fraussen 2006
G. Siphonalia Adams 1863
+
Subg. †Siphonalia (Siphonalia) Adams 1863
†Siphonalia acuta Rivera 1957
†Siphonalia ariejansseni Schnetler 2001
†Siphonalia belenensis Olsson 1931
†Siphonalia bryani Adegoke 1977
†Siphonalia calvertana Martin 1904
†Siphonalia cotteri Vredenburg 1923
†Siphonalia devexus Conrad 1843
†Siphonalia dilatata Ihering 1899
†Siphonalia guineensis Adegoke 1977
†Siphonalia hadlockensis Durham 1944
†Siphonalia ishikariana Oyama and Mizuno 1958
†Siphonalia jacksonia Harris 1896
†Siphonalia johnsoni Richards 1947
†Siphonalia kalpanae Banerjee and Halder 2024
†Siphonalia lineata Stanton 1896
†Siphonalia nigeriensis Adegoke 1977
†Siphonalia nipponica Oyama and Mizuno 1958
†Siphonalia nuntia Olsson 1930
†Siphonalia packi Dickerson 1917
†Siphonalia paradoxica Jenkins 1864
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†Siphonalia phosoidea Hanna and Israelsky 1925
†Siphonalia pritilatae Banerjee and Halder 2024
†Siphonalia roblini Tenison-Woods 1876
†Siphonalia rodeoensis Trask 1922
†Siphonalia sakakurai Mizuno 1954
†Siphonalia shojii Hirayama 1956
Siphonalia signa Reeve 1846
†Siphonalia spatiosa Tate 1888
†Siphonalia sullivani Harris and Palmer 1947
†Siphonalia sullivani ouachitae Harris and Palmer 1947
†Siphonalia teglandae Durham 1944
†Siphonalia tessaria Olsson 1931
†Siphonalia trilica Rivera 1957
†Siphonalia vistaensis Givens and Kennedy 1976
G. Verconella Iredale 1914
+
†Verconella penrosei Harris and Palmer 1947
G. †Stantonella Wade 1926
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†Stantonella interrupta Conrad 1858
†Stantonella monmouthensis Gardner 1916
†Stantonella ripleyana Conrad 1860
†Stantonella rugosa Stephenson 1923
†Stantonella subnodosa Wade 1926
G. †Suessionia Cossmann 1889
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†Suessionia canalifera Ravn 1939
Invalid names: Fusopsis Ravn 1939 [synonym]
G. †Suessonia Cossmann 1889
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†Suessonia invavica Traub 1979
G. †Terebrifusus Conrad 1865
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†Terebrifusus amoenus Conrad 1833
†Terebrifusus howei Hanna 1927
†Terebrifusus lepus Olsson 1930
†Terebrifusus placitus Olsson 1930
G. †Tritonidea Sowerby 1842
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†Tritonidea africana Adegoke 1977
†Tritonidea borcherti Ihering 1907
†Tritonidea conoidea Traub 1938
†Tritonidea heptagona Vredenburg 1923
†Tritonidea ickei Martin 1914
†Tritonidea kreyenhageni Arnold 1909
†Tritonidea praeundosa Vredenburg 1923
†Tritonidea promensis Vredenburg 1923
G. †Trominina Oyama and Mizuno 1958
+
†Trominina angasiana Yokoyama 1932
†Trominina bicordata Hatai and Matsui 1951
†Trominina ezoana Takeda 1953
†Trominina hokkaidoensis Hayasaka and Uozumi 1951
†Trominina ishikariensis Hayasaka and Matsui 1951
†Trominina japonica Takeda 1953
†Trominina onnaica Yokoyama 1932
†Trominina umbelliformis Hayasaka and Uozumi 1954
G. Truncaria Adams and Reeve 1848
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†Varicosipho asperulus Tate 1888
†Varicosipho labrosus Tate 1888
†Varicosipho styliformis Tenison-Woods 1879
Subfm. Volutopsiinae Habe and Sato 1972
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G. †Habevolutopsius Kantor 1983
G. †Lussivolutopsius Kantor 1983
G. Volutopsius Mörch 1857
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†Volutopsius eurekaensis Martin 1914
G. †Wrigleya Glibert 1963
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†Wrigleya crebrilinea Wright 1927
†Wrigleya transversaria Wrigley 1925
G. †Zaglenum Squires and Saul 2001
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†Zaglenum lomaensis Squires and Saul 2001
†Zaglenum pentzensis Squires and Saul 2001
G. †Zelandiella Finlay 1926
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†Zelandiella nodosa Hutton 1877
†Zelandiella propenodosa Bartrum 1919
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Invalid names: Siphonalia kaawaensis Bartrum 1919 [synonym]
†Zelandiella verrucosa Stilwell and Zinsmeister 1992
Invalid names: Donovaniinae Casey 1904 [empty], Pisaninae [empty], Sagenella Conrad 1865 [invalid subgroup]
Diagnosis
No diagnoses are available