PBDB Collection

Flat Creek 5 (Cretaceous of the United States)

Also known as UCMP V-73087; UWBM C1614; UCMP V73087

Where: Garfield County, Montana (47.5° N, 106.4° W: paleocoordinates 54.2° N, 79.5° W)

• coordinate based on unpublished field data

• small collection-level geographic resolution

When: Upper Member (Hell Creek Formation), Lancian (70.6 - 66.0 Ma)

• bed-level stratigraphic resolution

Environment/lithology: terrestrial; silty sandstone

Size class: mesofossils

Reposited in the UCMP

Collection methods: bulk, surface (float), sieve,

Primary reference: J. D. Archibald. 1982. A study of Mammalia and geology across the Cretaceous-Tertiary boundary in Garfield County, Montana. University of California Publications in Geological Sciences 122:1-286 [J. Alroy/J. Alroy/M. Carrano]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 14549: authorized by John Alroy, entered by John Alroy on 18.02.1993, edited by Matthew Carrano and Patricia Holroyd

Creative Commons license: CC BY (attribution)

Taxonomic list

• this appears to actually post-date Archibald and Clemens 1984, so their "Flat Creek" list should be ignoredupper level of Flat Creek sectionNISP 217 (minimum; maximum 222)

Show authors, comments, and common names

unclassified
-	Galagadon nordquistae¹⁴ Galagadon nordquistae¹⁴ UCMP 287558–287560
Mammalia
Theriamorpha - Pediomyidae	"Pediomys florencae" = Protolambda florencae, "Pediomys hatcheri" = Protolambda hatcheri, "Pediomys krejcii" = Leptalestes krejcii, "Pediomys cf. cooki" = Leptalestes cooki, Pediomys elegans "Pediomys florencae" = Protolambda florencae Clemens 1966 metatherian "Pediomys hatcheri" = Protolambda hatcheri Osborn 1898 metatherian "Pediomys krejcii" = Leptalestes krejcii Clemens 1966 metatherian "Pediomys cf. cooki" = Leptalestes cooki Clemens 1966 metatherian Pediomys elegans Marsh 1889 metatherian
Theriamorpha - Herpetotheriidae	"Alphadon jasoni" = Nortedelphys jasoni⁸ "Alphadon jasoni" = Nortedelphys jasoni⁸ Storer 1991 marsupial most of the A. marshi/wilsoni sample
Theriamorpha - Stagodontidae	Didelphodon vorax Didelphodon vorax Marsh 1889 metatherian (minimum; maximum 6)
Polydolopimorphia - Glasbiidae	Glasbius twitchelli n. sp. Glasbius twitchelli n. sp. Archibald 1982 metatherian
Theriamorpha - Alphadontidae	Alphadon marshi, Turgidodon rhaister¹ Alphadon marshi Simpson 1927 metatherian includes "A. wilsoni": (32 + 23) Turgidodon rhaister¹ Clemens 1966 metatherian
Theriamorpha -	Protalphadon foxi n. sp.⁸ Protalphadon foxi n. sp.⁸ Johanson 1996 metatherian
Theriamorpha - Gypsonictopidae	Gypsonictops cf. illuminatus Gypsonictops cf. illuminatus Lillegraven 1969 eutherian
Cimolesta - Cimolestidae	Cimolestes incisus, Cimolestes stirtoni, "Cimolestes propalaeoryctes" = Scollardius propalaeoryctes, Batodon tenuis Cimolestes incisus Marsh 1889 eutherian Cimolestes stirtoni Clemens 1973 eutherian "Cimolestes propalaeoryctes" = Scollardius propalaeoryctes Lillegraven 1969 eutherian Batodon tenuis Marsh 1892 eutherian
Multituberculata - Neoplagiaulacidae	? Neoplagiaulax burgessi n. sp., Mesodma hensleighi, Mesodma cf. formosa, Mesodma thompsoni ? Neoplagiaulax burgessi n. sp. Archibald 1982 multituberculate Mesodma hensleighi Lillegraven 1969 multituberculate Mesodma cf. formosa Marsh 1889 multituberculate Mesodma thompsoni Clemens 1964 multituberculate
Multituberculata - Cimolodontidae	Cimolodon nitidus Cimolodon nitidus Marsh 1889 multituberculate
Multituberculata -	Paracimexomys priscus Paracimexomys priscus Lillegraven 1969 multituberculate
Multituberculata - Cimolomyidae	Meniscoessus robustus, Essonodon browni, Cimolomys gracilis Meniscoessus robustus Marsh 1889 multituberculate Essonodon browni Simpson 1927 multituberculate Cimolomys gracilis Marsh 1889 multituberculate
Reptilia
Squamata - Globauridae	Contogenys sloani⁶ Contogenys sloani⁶ Estes 1969 squamates UCMP 130732
Squamata - Chamopsiidae	Leptochamops denticulatus⁶, Chamops segnis⁶, Haptosphenus placodon⁶, Peneteius aquilonius¹¹ Leptochamops denticulatus⁶ Gilmore 1928 squamates Chamops segnis⁶ Marsh 1892 squamates Haptosphenus placodon⁶ Estes 1964 squamates Peneteius aquilonius¹¹ Estes 1969 squamates teiid lizard; UCMP 123325 9left maxilla), 124744 (right dentary)
Squamata - Xenosauridae	Exostinus sp.⁶, Exostinus lancensis⁶ Exostinus sp.⁶ Cope 1873 squamates Exostinus lancensis⁶ Gilmore 1928 squamates
Squamata - Anguidae	Odaxosaurus piger⁶ Odaxosaurus piger⁶ Gilmore 1928 squamates
Testudinata -	Compsemys victa⁶ Compsemys victa⁶ Leidy 1856 turtle
Testudines - Trionychidae	Trionychidae indet.² Trionychidae indet.² Gray 1825 softshell turtle
Testudines - Adocidae	Adocus sp.⁶ Adocus sp.⁶ Cope 1868 turtle
Loricata -	"Leidyosuchus sternbergi" = Borealosuchus sternbergii⁶, Crocodylia indet.³ "Leidyosuchus sternbergi" = Borealosuchus sternbergii⁶ Gilmore 1910 crocodilian Crocodylia indet.³ crocodilian
Crocodylia - Alligatoridae	Brachychampsa sp.⁶, Brachychampsa montana⁶ Brachychampsa sp.⁶ Gilmore 1911 crocodilian Brachychampsa montana⁶ Gilmore 1911 crocodilian
Ornithischia -	Thescelosaurus sp.⁶ Thescelosaurus sp.⁶ Gilmore 1913 ornithopod
Ornithischia - Hadrosauridae	"Anatosaurus sp." = Edmontosaurus⁶ "Anatosaurus sp." = Edmontosaurus⁶ Lambe 1917 hadrosaurine
Theropoda -	Theropoda indet.⁵, Paronychodon sp.⁶, Paronychodon lacustris⁵, cf. Paronychodon lacustris¹² Theropoda indet.⁵ theropod Paronychodon sp.⁶ Cope 1876 maniraptoran Paronychodon lacustris⁵ Cope 1876 maniraptoran cf. Paronychodon lacustris¹² Cope 1876 maniraptoran UCMP 187159, 123342, 187156, 187162, 1871464
Theropoda - Troodontidae	cf. Pectinodon sp.⁹ cf. Pectinodon sp.⁹ Carpenter 1982 maniraptoran UCMP 186885, 214060
Theropoda - Dromaeosauridae	Dromaeosauridae indet.⁵, Richardoestesia sp.⁶, Richardoestesia cf. isosceles¹², Richardoestesia cf. gilmorei⁹, Saurornitholestinae indet.⁹ Dromaeosauridae indet.⁵ Colbert and Russell 1969 maniraptoran Richardoestesia sp.⁶ Currie et al. 1990 maniraptoran Richardoestesia cf. isosceles¹² Sankey 2001 maniraptoran UCMP 213922–214006 Richardoestesia cf. gilmorei⁹ Currie et al. 1990 maniraptoran UCMP 187163 Saurornitholestinae indet.⁹ Longrich and Currie 2009 maniraptoran UCMP 124987, 187041
Aves
-	Aves indet.¹² Aves indet.¹² Linnaeus 1758 bird UCMP 186826, 214056-58
Reptilia
Theropoda - Tyrannosauridae	Aublysodon mirandus⁵ Aublysodon mirandus⁵ Leidy 1868 tyrannosaurid
Choristodera -	Champsosaurus sp.⁶, Champsosaurus natator⁶ Champsosaurus sp.⁶ Cope 1876 choristodere Champsosaurus natator⁶ Parks 1933 choristodere
Amphibia
Temnospondyli - Albanerpetontidae	Albanerpetontidae indet.⁷ Albanerpetontidae indet.⁷ Fox and Naylor 1982 tetrapod
Salientia -	Scotiophryne sp.⁶ Scotiophryne sp.⁶ Estes 1969 frog
Proteida - Batrachosauroididae	Opisthotriton kayi⁶ Opisthotriton kayi⁶ Auffenberg 1961 salamander
Caudata - Urodela	Scapherpeton tectum⁶, Lisserpeton bairdi⁶, Habrosaurus sp.⁶, Habrosaurus dilatus⁶ Scapherpeton tectum⁶ Cope 1876 salamander Lisserpeton bairdi⁶ Estes 1965 salamander Habrosaurus sp.⁶ Gilmore 1928 siren Habrosaurus dilatus⁶ Gilmore 1928 siren
Actinopteri
Amiiformes - Amiidae	Melvius thomasi⁶, Amia uintaensis⁶, Amia fragosa⁶ Melvius thomasi⁶ Bryant 1987 bowfin Amia uintaensis⁶ Leidy 1873 bowfin Amia fragosa⁶ Jordan 1927 bowfin
Lepisosteiformes - Lepisosteidae	Lepisosteus sp.⁶, Lepisosteus occidentalis⁶ Lepisosteus sp.⁶ Agassiz 1843 gar Lepisosteus occidentalis⁶ Leidy 1856 gar
Teleostei -	Teleostei "indet. U-4"⁴, Osteoglossomorpha indet.⁴ Teleostei "indet. U-4"⁴ Müller 1846 UCMP 230626, one abdominal centrum; UCMP 230627, one abdominal centrum; UCMP 230628, two abdominal centra Osteoglossomorpha indet.⁴ Greenwood et al. 1966 = Coriops/Lopadichthys; UCMP 230612, posterior abdominal centrum
Osteoglossiformes -	Wilsonichthys sp.¹⁰, ? Wilsonichthys sp.⁴, Wilsonichthys aridinsulensis⁴ Wilsonichthys sp.¹⁰ Murray et al. 2016 ? Wilsonichthys sp.⁴ Murray et al. 2016 UCMP 230619, centra type BvA Wilsonichthys aridinsulensis⁴ Murray et al. 2016
Hiodontiformes - Hiodontidae	Hiodontidae "indet. small morph"⁴, Hiodontidae "indet. large morph"⁴ Hiodontidae "indet. small morph"⁴ UCMP 230617, four centra of small size, including a first abdominal centrum Hiodontidae "indet. large morph"⁴ UCMP 230616, two centra of large size, including a first abdominal centrum
Gonorhynchiformes - Gonorhynchidae	Notogoneus sp.⁴ Notogoneus sp.⁴ Cope 1885
Ostariophysi -	"Otophysi "indet. U-3/BvD"" = Otophysa⁴ "Otophysi "indet. U-3/BvD"" = Otophysa⁴ UCMP 230603, one anterior Weberian centrum (centrum type U3); UCMP 230604, one anterior Weberian centrum (centrum type U3); UCMP 230620, a third Weberian centrum (centrum type BvC); UCMP 230621, a third Weberian centrum (centrum type BvC)
Acanthomorphata -	"Acanthomorpha "indet. HC-1"" = Acanthomorphata⁴, "Acanthomorpha "indet. HC-2"" = Acanthomorphata⁴, "Acanthomorpha "indet. HC-4"" = Acanthomorphata⁴, "Acanthomorpha "indet. HC-5"" = Acanthomorphata⁴, "Acanthomorpha "indet. HC-6"" = Acanthomorphata⁴ "Acanthomorpha "indet. HC-1"" = Acanthomorphata⁴ Rosen 1973 UCMP 230601, five first abdominal centra; UCMP 230591, four abdominal centra; UCMP 230599, two abdominal centra "Acanthomorpha "indet. HC-2"" = Acanthomorphata⁴ Rosen 1973 UCMP 230593, abdominal centra "Acanthomorpha "indet. HC-4"" = Acanthomorphata⁴ Rosen 1973 "Acanthomorpha "indet. HC-5"" = Acanthomorphata⁴ Rosen 1973 UCMP 230597, first abdominal centrum "Acanthomorpha "indet. HC-6"" = Acanthomorphata⁴ Rosen 1973 UCMP 198876, one abdominal centrum
Teleostei -	Percopsiformes indet.⁴ Percopsiformes indet.⁴ Berg 1940 UCMP 198875, dentary
Acanthomorphata - Moronidae	? Moronidae indet.⁴ ? Moronidae indet.⁴ temperate bass UCMP 198873, dentary
Acanthomorphata - Sciaenidae	Platacodon nanus⁶ Platacodon nanus⁶ Marsh 1889 UCMP 198873, tooth plate; UCMP 198872, teeth
Esociformes - Esocidae	Estesesox foxi¹³ Estesesox foxi¹³ Wilson et al. 1992 UCMP 230608, centrum type NvC; UCMP 230606, centrum type NvD
Salmoniformes -	Esocoidei indet.¹³ Esocoidei indet.¹³ Berg 1936
Aspidorhynchiformes - Aspidorhynchidae	Belonostomus longirostris⁶ Belonostomus longirostris⁶ Lambe 1902
Osteichthyes
Acipenseriformes - Acipenseridae	Acipenser sp.⁶ Acipenser sp.⁶ Linnaeus 1758 sturgeon
Chondrichthyes
Batoidea -	Cristomylus leptostemma¹⁴ Cristomylus leptostemma¹⁴ Wynd et al. 2020 ray UCMP 287576
Rajiformes - Sclerorhynchidae	Onchosaurus avonicolus⁶ Onchosaurus avonicolus⁶ ray
Rajiformes - Rhinobatidae	Myledaphus pustulosus¹⁴ Myledaphus pustulosus¹⁴ Cook et al. 2014 guitarfish UCMP 128742, 287561
Rajiformes -	Squatirhina sp.⁶ Squatirhina sp.⁶ Casier 1947 ray
Orectolobiformes - Orectolobidae	"Squatirhina americana" = Restesia americana⁶, Restesia monocuspilatus¹⁴ "Squatirhina americana" = Restesia americana⁶ Estes 1964 wobbegong UCMP 194915, one complete tooth; UCMP 194916, one complete tooth; UCMP 194917, one complete tooth; UMCP 128739, 98 complete and fragmented teeth; UCMP 123283, 187 complete and fragmented teeth Restesia monocuspilatus¹⁴ Wynd et al. 2020 wobbegong UCMP 287562, 287563, 287564, 287565, 287566, 287567, 287571, 287575
Orectolobiformes - Ginglymostomatidae	Cantioscyllium estesi¹⁴ Cantioscyllium estesi¹⁴ nurse shark UCMP 287573
Orectolobiformes - Brachaeluridae	Brachaelurus estesi⁶ Brachaelurus estesi⁶ Herman 1977 carpet shark
Hybodontiformes - Lonchidiidae	Lonchidion selachos⁶ Lonchidion selachos⁶ Estes 1964 elasmobranch UCMP 194918, fractured tooth; UCMP 194919, fractured tooth; UCMP 123281, 37 fragmented teeth; UCMP 128740, 13 fragmented teeth

¹Alroy 2002; ²Alroy 2006; ³Alroy 2007; ⁴Brinkman et al. 2021; ⁵Carpenter 1982; ⁶Database 2005; ⁷Gardner and DeMar 2013; ⁸Johanson 1996; ⁹Larson and Currie 2013; ¹⁰Murray et al. 2016; ¹¹Nydam et al. 2000; ¹²Sankey 2008; ¹³Wilson et al. 1992; ¹⁴Wynd et al. 2020