PBDB Collection

Flat Creek 5 (Cretaceous of the United States)

Also known as UCMP V-73087; UWBM C1614; UCMP V73087

Where: Garfield County, Montana (47.5° N, 106.4° W: paleocoordinates 54.2° N, 79.5° W)

• coordinate based on unpublished field data

• small collection-level geographic resolution

When: Upper Member (Hell Creek Formation), Lancian (70.6 - 66.0 Ma)

• bed-level stratigraphic resolution

Environment/lithology: terrestrial; silty sandstone

Size class: mesofossils

Reposited in the UCMP

Collection methods: bulk, surface (float), sieve,

Primary reference: J. D. Archibald. 1982. A study of Mammalia and geology across the Cretaceous-Tertiary boundary in Garfield County, Montana. University of California Publications in Geological Sciences 122:1-286 [J. Alroy/J. Alroy/M. Carrano]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 14549: authorized by John Alroy, entered by John Alroy on 18.02.1993, edited by Matthew Carrano and Patricia Holroyd

Creative Commons license: CC BY (attribution)

Taxonomic list

• this appears to actually post-date Archibald and Clemens 1984, so their "Flat Creek" list should be ignoredupper level of Flat Creek sectionNISP 217 (minimum; maximum 222)

Show authors, comments, and common names

unclassified
-	Galagadon nordquistae¹⁵ Galagadon nordquistae¹⁵ UCMP 287558–287560
Chondrichthyes
Orectolobiformes - Ginglymostomatidae	Cantioscyllium estesi¹⁵ Cantioscyllium estesi¹⁵ nurse shark UCMP 287573
Orectolobiformes - Brachaeluridae	Brachaelurus estesi⁷ Brachaelurus estesi⁷ Herman 1977 carpet shark
Orectolobiformes - Orectolobidae	"Squatirhina americana" = Restesia americana⁷, Restesia monocuspilatus¹⁵ "Squatirhina americana" = Restesia americana⁷ Estes 1964 wobbegong UCMP 194915, one complete tooth; UCMP 194916, one complete tooth; UCMP 194917, one complete tooth; UMCP 128739, 98 complete and fragmented teeth; UCMP 123283, 187 complete and fragmented teeth Restesia monocuspilatus¹⁵ Wynd et al. 2020 wobbegong UCMP 287562, 287563, 287564, 287565, 287566, 287567, 287571, 287575
Rajiformes - Rhinobatidae	Myledaphus pustulosus¹⁵ Myledaphus pustulosus¹⁵ Cook et al. 2014 guitarfish UCMP 128742, 287561
Rajiformes - Sclerorhynchidae	Onchosaurus avonicolus⁷ Onchosaurus avonicolus⁷ ray
Rajiformes -	Squatirhina sp.⁷ Squatirhina sp.⁷ Casier 1947 ray
Batoidea -	Cristomylus leptostemma¹⁵ Cristomylus leptostemma¹⁵ Wynd et al. 2020 ray UCMP 287576
Hybodontiformes - Lonchidiidae	Lonchidion selachos⁷ Lonchidion selachos⁷ Estes 1964 elasmobranch UCMP 194918, fractured tooth; UCMP 194919, fractured tooth; UCMP 123281, 37 fragmented teeth; UCMP 128740, 13 fragmented teeth
Osteichthyes
Acipenseriformes - Acipenseridae	Acipenser sp.⁷ Acipenser sp.⁷ Linnaeus 1758 sturgeon
Mammalia
Multituberculata - Cimolomyidae	Meniscoessus robustus, Cimolomys gracilis, Essonodon browni Meniscoessus robustus Marsh 1889 multituberculate Cimolomys gracilis Marsh 1889 multituberculate Essonodon browni Simpson 1927 multituberculate
Multituberculata - Cimolodontidae	Cimolodon nitidus Cimolodon nitidus Marsh 1889 multituberculate
Multituberculata - Neoplagiaulacidae	Mesodma cf. formosa, Mesodma thompsoni, Mesodma hensleighi, ? Neoplagiaulax burgessi n. sp. Mesodma cf. formosa Marsh 1889 multituberculate Mesodma thompsoni Clemens 1964 multituberculate Mesodma hensleighi Lillegraven 1969 multituberculate ? Neoplagiaulax burgessi n. sp. Archibald 1982 multituberculate
Multituberculata -	Paracimexomys priscus Paracimexomys priscus Lillegraven 1969 multituberculate
Leptictida - Gypsonictopidae	Gypsonictops cf. illuminatus Gypsonictops cf. illuminatus Lillegraven 1969 eutherian
Cimolesta - Cimolestidae	Batodon tenuis, "Cimolestes propalaeoryctes" = Scollardius propalaeoryctes, Cimolestes incisus, Cimolestes stirtoni Batodon tenuis Marsh 1892 eutherian "Cimolestes propalaeoryctes" = Scollardius propalaeoryctes Lillegraven 1969 eutherian Cimolestes incisus Marsh 1889 eutherian Cimolestes stirtoni Clemens 1973 eutherian
Tribosphenida - Alphadontidae	Alphadon marshi, Turgidodon rhaister¹ Alphadon marshi Simpson 1927 metatherian includes "A. wilsoni": (32 + 23) Turgidodon rhaister¹ Clemens 1966 metatherian
Tribosphenida - Pediomyidae	"Pediomys krejcii" = Leptalestes krejcii, "Pediomys cf. cooki" = Leptalestes cooki, Pediomys elegans, "Pediomys florencae" = Protolambda florencae, "Pediomys hatcheri" = Protolambda hatcheri "Pediomys krejcii" = Leptalestes krejcii Clemens 1966 metatherian "Pediomys cf. cooki" = Leptalestes cooki Clemens 1966 metatherian Pediomys elegans Marsh 1889 metatherian "Pediomys florencae" = Protolambda florencae Clemens 1966 metatherian "Pediomys hatcheri" = Protolambda hatcheri Osborn 1898 metatherian
Tribosphenida -	Protalphadon foxi n. sp.⁹ Protalphadon foxi n. sp.⁹ Johanson 1996 metatherian
Tribosphenida - Stagodontidae	Didelphodon vorax Didelphodon vorax Marsh 1889 metatherian (minimum; maximum 6)
Polydolopimorphia - Glasbiidae	Glasbius twitchelli n. sp. Glasbius twitchelli n. sp. Archibald 1982 metatherian
Tribosphenida - Herpetotheriidae	"Alphadon jasoni" = Nortedelphys jasoni⁹ "Alphadon jasoni" = Nortedelphys jasoni⁹ Storer 1991 marsupial most of the A. marshi/wilsoni sample
Reptilia
Loricata -	"Leidyosuchus sternbergi" = Borealosuchus sternbergii⁷, Crocodylia indet.³ "Leidyosuchus sternbergi" = Borealosuchus sternbergii⁷ Gilmore 1910 crocodilian Crocodylia indet.³ crocodilian
Crocodylia - Alligatoridae	Brachychampsa sp.⁷, Brachychampsa montana⁷ Brachychampsa sp.⁷ Gilmore 1911 crocodilian Brachychampsa montana⁷ Gilmore 1911 crocodilian
Ornithischia -	Thescelosaurus sp.⁷ Thescelosaurus sp.⁷ Gilmore 1913 ornithopod
Ornithischia - Hadrosauridae	"Anatosaurus sp." = Edmontosaurus⁷ "Anatosaurus sp." = Edmontosaurus⁷ Lambe 1917 hadrosaurine
Theropoda -	Theropoda indet.⁶ Theropoda indet.⁶ Marsh 1881 theropod
Aves
-	Aves indet.¹³ Aves indet.¹³ Linnaeus 1758 bird UCMP 186826, 214056-58
Reptilia
Theropoda - Dromaeosauridae	Dromaeosauridae indet.⁶, Saurornitholestinae indet.¹⁰, Richardoestesia sp.⁷, Richardoestesia cf. gilmorei¹⁰, Richardoestesia cf. isosceles¹³ Dromaeosauridae indet.⁶ Colbert and Russell 1969 maniraptoran Saurornitholestinae indet.¹⁰ Longrich and Currie 2009 maniraptoran UCMP 124987, 187041 Richardoestesia sp.⁷ Currie et al. 1990 maniraptoran Richardoestesia cf. gilmorei¹⁰ Currie et al. 1990 maniraptoran UCMP 187163 Richardoestesia cf. isosceles¹³ Sankey 2001 maniraptoran UCMP 213922–214006
Theropoda - Troodontidae	cf. Pectinodon sp.¹⁰ cf. Pectinodon sp.¹⁰ Carpenter 1982 maniraptoran UCMP 186885, 214060
Theropoda -	Paronychodon sp.⁷, Paronychodon lacustris⁶, cf. Paronychodon lacustris¹³ Paronychodon sp.⁷ Cope 1876 maniraptoran Paronychodon lacustris⁶ Cope 1876 maniraptoran cf. Paronychodon lacustris¹³ Cope 1876 maniraptoran UCMP 187159, 123342, 187156, 187162, 1871464
Theropoda - Tyrannosauridae	Aublysodon mirandus⁶ Aublysodon mirandus⁶ Leidy 1868 tyrannosaurid
Choristodera -	Champsosaurus sp.⁷, Champsosaurus natator⁷ Champsosaurus sp.⁷ Cope 1876 choristodere Champsosaurus natator⁷ Parks 1933 choristodere
Squamata - Anguidae	Odaxosaurus piger⁷ Odaxosaurus piger⁷ Gilmore 1928 squamates
Squamata - Xenosauridae	Exostinus sp.⁷, Exostinus lancensis⁷ Exostinus sp.⁷ Cope 1873 squamates Exostinus lancensis⁷ Gilmore 1928 squamates
Squamata - Chamopsiidae	Peneteius aquilonius¹², Chamops segnis⁷, Haptosphenus placodon⁷, Leptochamops denticulatus⁷ Peneteius aquilonius¹² Estes 1969 squamates teiid lizard; UCMP 123325 9left maxilla), 124744 (right dentary) Chamops segnis⁷ Marsh 1892 squamates Haptosphenus placodon⁷ Estes 1964 squamates Leptochamops denticulatus⁷ Gilmore 1928 squamates
Squamata - Globauridae	Contogenys sloani⁷ Contogenys sloani⁷ Estes 1969 squamates UCMP 130732
Testudinata -	Compsemys victa⁷ Compsemys victa⁷ Leidy 1856 turtle
Testudines - Adocidae	Adocus sp.⁷ Adocus sp.⁷ Cope 1868 turtle
Testudines - Trionychidae	Trionychidae indet.² Trionychidae indet.² Gray 1825 softshell turtle
Amphibia
Caudata - Urodela	Habrosaurus sp.⁷, Habrosaurus dilatus⁷, Lisserpeton bairdi⁷, Scapherpeton tectum⁷ Habrosaurus sp.⁷ Gilmore 1928 siren Habrosaurus dilatus⁷ Gilmore 1928 siren Lisserpeton bairdi⁷ Estes 1965 salamander Scapherpeton tectum⁷ Cope 1876 salamander
Proteida - Batrachosauroididae	Opisthotriton kayi⁷ Opisthotriton kayi⁷ Auffenberg 1961 salamander
Salientia -	Scotiophryne sp.⁷ Scotiophryne sp.⁷ Estes 1969 frog
Temnospondyli - Albanerpetontidae	Albanerpetontidae indet.⁸ Albanerpetontidae indet.⁸ Fox and Naylor 1982 tetrapod
Actinopteri
Lepisosteiformes - Lepisosteidae	Lepisosteus sp.⁷, Lepisosteus occidentalis⁷ Lepisosteus sp.⁷ Agassiz 1843 gar Lepisosteus occidentalis⁷ Leidy 1856 gar
Amiiformes - Amiidae	Amia fragosa⁷, Melvius thomasi⁵ Amia fragosa⁷ Jordan 1927 bowfin Melvius thomasi⁵ Bryant 1988 bowfin UCMP 123289, 150+ tooth crowns; UCMP 130386, dentary fragment
Teleostei -	Teleostei "indet. U-4"⁴ Teleostei "indet. U-4"⁴ Müller 1846 UCMP 230626, one abdominal centrum; UCMP 230627, one abdominal centrum; UCMP 230628, two abdominal centra
Acanthomorphata -	"Acanthomorpha "indet. HC-1"" = Acanthomorphata⁴, "Acanthomorpha "indet. HC-2"" = Acanthomorphata⁴, "Acanthomorpha "indet. HC-4"" = Acanthomorphata⁴, "Acanthomorpha "indet. HC-5"" = Acanthomorphata⁴, "Acanthomorpha "indet. HC-6"" = Acanthomorphata⁴ "Acanthomorpha "indet. HC-1"" = Acanthomorphata⁴ Rosen 1973 UCMP 230601, five first abdominal centra; UCMP 230591, four abdominal centra; UCMP 230599, two abdominal centra "Acanthomorpha "indet. HC-2"" = Acanthomorphata⁴ Rosen 1973 UCMP 230593, abdominal centra "Acanthomorpha "indet. HC-4"" = Acanthomorphata⁴ Rosen 1973 "Acanthomorpha "indet. HC-5"" = Acanthomorphata⁴ Rosen 1973 UCMP 230597, first abdominal centrum "Acanthomorpha "indet. HC-6"" = Acanthomorphata⁴ Rosen 1973 UCMP 198876, one abdominal centrum
Acanthomorphata - Sciaenidae	Platacodon nanus⁷ Platacodon nanus⁷ Marsh 1889 UCMP 198873, tooth plate; UCMP 198872, teeth
Acanthomorphata - Moronidae	? Moronidae indet.⁴ ? Moronidae indet.⁴ temperate bass UCMP 198873, dentary
Teleostei -	Percopsiformes indet.⁴ Percopsiformes indet.⁴ Berg 1940 UCMP 198875, dentary
Esociformes - Esocidae	Estesesox foxi¹⁴ Estesesox foxi¹⁴ Wilson et al. 1992 UCMP 230608, centrum type NvC; UCMP 230606, centrum type NvD
Salmoniformes -	Esocoidei indet.¹⁴ Esocoidei indet.¹⁴ Berg 1936
Gonorhynchiformes - Gonorhynchidae	Notogoneus sp.⁴ Notogoneus sp.⁴ Cope 1885
Ostariophysi -	"Otophysi "indet. U-3/BvD"" = Otophysa⁴ "Otophysi "indet. U-3/BvD"" = Otophysa⁴ UCMP 230603, one anterior Weberian centrum (centrum type U3); UCMP 230604, one anterior Weberian centrum (centrum type U3); UCMP 230620, a third Weberian centrum (centrum type BvC); UCMP 230621, a third Weberian centrum (centrum type BvC)
Teleostei -	Osteoglossomorpha indet.⁴ Osteoglossomorpha indet.⁴ Greenwood et al. 1966 = Coriops/Lopadichthys; UCMP 230612, posterior abdominal centrum
Osteoglossiformes -	Wilsonichthys sp.¹¹, ? Wilsonichthys sp.⁴, Wilsonichthys aridinsulensis⁴ Wilsonichthys sp.¹¹ Murray et al. 2016 ? Wilsonichthys sp.⁴ Murray et al. 2016 UCMP 230619, centra type BvA Wilsonichthys aridinsulensis⁴ Murray et al. 2016
Hiodontiformes - Hiodontidae	Hiodontidae "indet. small morph"⁴, Hiodontidae "indet. large morph"⁴ Hiodontidae "indet. small morph"⁴ UCMP 230617, four centra of small size, including a first abdominal centrum Hiodontidae "indet. large morph"⁴ UCMP 230616, two centra of large size, including a first abdominal centrum
Aspidorhynchiformes - Aspidorhynchidae	Belonostomus longirostris⁷ Belonostomus longirostris⁷ Lambe 1902

¹Alroy 2002; ²Alroy 2006; ³Alroy 2007; ⁴Brinkman et al. 2021; ⁵Bryant 1988; ⁶Carpenter 1982; ⁷Database 2005; ⁸Gardner and DeMar 2013; ⁹Johanson 1996; ¹⁰Larson and Currie 2013; ¹¹Murray et al. 2016; ¹²Nydam et al. 2000; ¹³Sankey 2008; ¹⁴Wilson et al. 1992; ¹⁵Wynd et al. 2020