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Weaniinae

Artiopoda - Proetida - Phillipsiidae

Synonyms
Synonymy list
YearName and author
1983Weaniinae Owens
1998Carbonocoryphinae Yuan and Xiang p. 83
2003Weaniinae Owens p. 383
2020Weaniinae Basse and Lemke p. 223
2025Weaniinae Brezinski p. 143

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—EcdysozoaAguinaldo et al. 1997
—Panarthropoda
phylumArthropodaGravenhorst 1843
RankNameAuthor
classArtiopodaHou and Bergstrom
classTrilobitaWalch 1771
orderProetidaFortey and Owens 1975
superfamilyProetoideaHawle and Corda 1847
familyPhillipsiidaeOehlert 1886
subfamilyWeaniinaeOwens 1983
subfamilyWeaniinaeOwens 1983

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Subfm. †Weaniinae Owens 1983
show all | hide all
G. †Amphiroaspis Müller and Hahn 2018
+
†Amphiroaspis amphiro Hahn et al. 1996
G. †Aprathia Hahn and Brauckmann 1975
+
†Aprathia perlisensis Kobayashi and Hamada 1966
G. †Belgibole Hahn 1963
+
†Belgibole abruptirhachis Richter and Richter 1919
†Belgibole porteri Goldring 1955
G. †Carbonocoryphe Richter and Richter 1950
+
G. †Dechenelloides Gandl 1968
+
†Dechenelloides peoui Engel and Morris 1996
G. †Doublatia Wass and Banks 1971
+
†Doublatia inflata Wass and Banks 1971
†Doublatia matheri Engel and Laurie 1978
†Doublatia pyriforme Wass and Banks 1971
G. †Endops Koizumi 1972
+
†Endops yanagisawai Endo and Matsumoto 1962
G. †Evagena Hahn and Brauckmann 1989
+
†Evagena evae Hahn and Brauckmann 1989
G. †Geigibole Gandl 1968
+
†Geigibole geigensis Gandl 1968
†Geigibole heisingi Hahn and Brauckmann 1988
†Geigibole kohleichensis Müller and Hahn 2022
†Geigibole thomasi Brauckmann 1973
†Geigibole vivax Hahn and Hahn 1971
G. †Gitarra Gandl 1968
G. †Gracemerea Engel and Morris 1989
+
†Gracemerea archeri Engel and Morris 1989
G. †Hassiabole Hahn et al. 1996
G. †Nipponaspis Koizumi 1972
+
†Nipponaspis miosilus Chamberlain 1977
†Nipponaspis takaizumii Koizumi 1972
G. †Phillibolina Gandl 1968
G. †Pseudowaribole Hahn and Hahn 1967
+
†Pseudowaribole octofera Richter and Richter 1926
+
Invalid names: Pseudowaribole altera Osmólska 1962 [synonym]
†Pseudowaribole phacomma Richter and Richter 1926
†Pseudowaribole sardarensis Feist et al. 2003
G. †Schizophillipsia Kobayashi and Hamada 1980
G. †Spergenaspis Brezinski 1987
+
†Spergenaspis easleyi Brezinski 1987
†Spergenaspis mauvaisensis Hessler 1965
†Spergenaspis salemi Brezinski 1987
G. †Thalabaria Engel and Morris 1989
+
†Thalabaria plana Engel and Morris 1980
G. †Weania Campbell and Engel 1963
+
†Weania goldringi Campbell and Engel 1963
†Weania insulana Flick and Shiino 2021
†Weania neso Fey 1983
†Weania praecox Brauckmann 1974
†Weania semicircularis Engel and Morris 1996
G. †Winterbergia Hahn and Brauckmann 1975
+
Subg. †Winterbergia (Winterbergia) Hahn and Brauckman 1975
+
†Winterbergia (Winterbergia) costata Yuan and Xiang 1998
†Winterbergia (Winterbergia) prisca Yuan and Xiang 1998
Invalid names: Winterbergia elegans Engel and Morris 1989 [nomen dubium], Winterbergia keepitensis Engel and Morris 1989 [nomen dubium]
Invalid names: Carbonocoryphinae Yuan and Xiang 1998 [synonym]
Diagnosis
ReferenceDiagnosis
J. Yuan and L.-W. Xiang 1998 (Carbonocoryphinae)A new subfamily of Phillipsiidae (Oehlert, 1886), Hahn, Hahn and Brauckmann, 1980 with the following features: glabella truncated conical, pyriform to cylindrical in outline, slightly constricted between r - r , with 3 or 4 pairs of distinct lateral glabellar furrows, of which the S1 is incised; occipital furrow with or without lateral branches; occipital ring without isolated lateral occipital lobe; preglabellar field commonly present and more or less depressed; anterior border furrow distinct; fixed cheek relatively narrow at E and somewhat depressed opposite to palpebral lobes; elevated banded palpebral lobes large to medium in size; facial sutures with short or without straight section E - S ; thorax of 7-9 segments; pygidium shorter than cephalon, with ill-defined border; axis relatively short, with 7-11 axial rings and a distinct postaxial area, on which a long postaxial ridge is present or not; pleural areas with 5-9 pairs of pleural ribs, of
which anterior bands are more elevated and longer than posterior bands; pleural furrows deep, interpleural furrows shallow; pygidial border more or less demarcated; without distinct border furrow.
Measurements
No measurements are available
Composition: low Mg calcitec
Composition 2: chitinc
Ontogeny: molting, addition of partsc
Environment: marinec
Locomotion: fast-movingc
Life habit: low-level epifaunalc
Diet: deposit feedero
Vision: well-developedf
Created: 2009-12-19 02:34:58
Modified: 2009-12-19 04:34:58
Source: f = family, o = order, c = class
References: Whittington et al. 1997, Fortey and Owens 1999, Aberhan et al. 2004

Age range: base of the Late/Upper Famennian to the top of the Capitanian or 365.20000 to 259.51000 Ma

Collections (122 total)


Time interval Ma Country or state Original ID and collection number
Late/Upper Devonian382.31 - 358.86United Kingdom Belgibole abruptirhachis (47471)
Famennian372.15 - 358.86Malaysia (Langkawi Islands) Waribole perlisense (218988)
Famennian372.15 - 358.86Germany Belgibole abruptirhachis (47467)
Famennian372.15 - 358.86Germany (Nordrhein-Westfalen) Pseudowaribole quaesita (38295 38296 38298 38299 38300 38302)
Famennian372.15 - 358.86Poland Cyrtosymbole octofera, Cyrtosymbole phacomma (217139) Cyrtosymbole phacomma, Cyrtosymbole octofera (217138)
Famennian372.15 - 358.86China (Guangxi) Pseudocyrtosymbole acutangularis (229268)
Late/Upper Famennian365.2 - 358.86Iran Pseudowaribole sardarensis (223428)
Late/Upper Famennian365.2 - 358.86Germany (Nordrhein-Westfalen) Belgibole abruptirhachis, Semiproetus funirepus (229359) Pseudowaribole quaesita (38304 38306)
Late/Upper Famennian365.2 - 358.86Germany Belgibole abruptirhachis (47468 47469 47470) Belgibole abruptirhachis, Semiproetus funirepa (218980) Eowinterbergia effenbergensis (229398) Pseudowaribole conifera (218983 218984) Pseudowaribole sp. (229399) Pseudowaribole sp., Pseudowaribole octofera (229263)
Late/Upper Famennian365.2 - 358.86Austria Belgibole abruptirhachis, Semiproetus funirepa (38608 42407)
Late/Upper Famennian365.2 - 358.86Czechia Cytosymbole superna (218975) Winterbergia superna (234673)
Late/Upper Famennian365.2 - 358.86China (Guangxi) Pseudowaribole sp., Pseudocyrtosymbole sp., Pseudocyrtosymbole yishanensis (222338)
Late/Upper Famennian365.2 - 358.86France Archegonus pauli (235124) Pseudowaribole sp., Pseudowaribole octofera, Pseudowaribole conifera (38641)
Late/Upper Famennian365.2 - 358.86China (Guizhou) Pseudowaribole macrops (218959 218960 218961 218962 218963 218964 218965 218966 234698)
Late/Upper Famennian365.2 - 358.86Poland (Swietokrzyskie) Cyrtosymbole conifera (223416)
Late/Upper Famennian365.2 - 358.86United Kingdom (England) Cyrtosymbole conifera (234936)
Hastarian358.86 - 353.7China (Guangxi) Belgibole abruptirhachis (222330 222331) Winterbergia conica (222342) Winterbergia conica, Pseudocyrtosymbole granifera (222341)
Hastarian358.86 - 353.7France Belgibole abruptirhachis, Semiproetus funirepa (38642)
Hastarian358.86 - 353.7United Kingdom (England) Archegonus porteri (38448)
Hastarian358.86 - 353.7Poland Carbonocoryphe hercynica (218771)
Hastarian358.86 - 353.7Austria Semiproetus funirepa (38610 38612 38613 42408 42409 42410 42411 42412 42413)
Hastarian358.86 - 353.7Germany (Nordrhein-Westfalen) Belgibole abruptirhachis (38646 229361)
Hastarian358.86 - 353.7Australia (New South Wales) Weania goldringi (75621)
Hastarian358.86 - 353.7China (Guizhou) Belgibole abruptirhachis (218953) Dechenelloides sp., Winterbergia transversa, Winterbergia australis, Winterbergia lata (218945) Winterbergia australis (218947) Winterbergia costata (218952) Winterbergia cylindrica (218970 218972) Winterbergia cylindrica, Winterbergia conica (218948) Winterbergia prisca, Winterbergia costata (218977) Winterbergia prisca, Winterbergia subcylindrica, Winterbergia costata, Pseudowaribole sp. (218976) Winterbergia rugosa, Winterbergia cylindrica (218969 218971) Winterbergia rugosa, Winterbergia cylindrica, Winterbergia conica (218973) Winterbergia rugosa, Winterbergia prisca, Winterbergia subcylindrica, Winterbergia conica (218974) Winterbergia sp. (218949 218950) Winterbergia subcylindrica, Winterbergia costata (218951) Winterbergia subcylindrica, Winterbergia superna (218979) Winterbergia superna (218954) Winterbergia transversa, Winterbergia australis, Winterbergia lata (218946)
Kinderhookian358.86 - 351.9USA (Texas) Griffithidella depressus (229265)
Kinderhookian358.86 - 351.9USA (Missouri) Spergenaspis easleyi (229176)
Hastarian - Ivorian358.86 - 346.7Germany (Nordrhein-Westfalen) Pseudowaribole conifera (229364)
Tournaisian358.86 - 346.7Australia (Queensland) Phillipsia rockhamptonensis (217065) Winterbergia elegans, Gracemerea archeri (217040)
Tournaisian358.86 - 346.7Germany (Nordrhein-Westfalen) Belgibole abruptirhachis (38644)
Tournaisian358.86 - 346.7Australia (New South Wales) Cyrtoproetus planus (217055) Winterbergia keepitensis (217073)
Kinderhookian - Osagean358.86 - 342.9USA (Texas) Griffithidella depressa (5)
Ivorian353.7 - 346.7Germany (Nordrhein-Westfalen) Winterbergia parahahnorum, Amphiroaspis amphiro (229351)
Ivorian353.7 - 346.7Belgium Carbonocoryphe andrasta (42758) Carbonocoryphe modavensis (37127)
Ivorian353.7 - 346.7Czechia Carbonocoryphe bindemanni (138585) Carbonocoryphe egregia (67220)
Ivorian353.7 - 346.7Spain (León) Pseudowaribole sajambrensis (67109)
Ivorian353.7 - 346.7Italy Carbonocoryphe suprahercynica, Pseudowaribole inusitata (37221)
Ivorian353.7 - 346.7Germany Carbonocoryphe hercynica (37192) Carbonocoryphe suprahercynica, Pseudowaribole inusitata (42417) Hassiabole plexaura (37187) Hassiabole plexaura, Carbonocoryphe laedata, Carbonocoryphe amphiro (37189)
Chadian346.7 - 339.9United Kingdom (England) Phillibolina worsawensis (5962) Phillibolina worsawensis, Weania feltrimensis, Winterbergia hahnorum (5963) Winterbergia sp. (5964)
Chadian346.7 - 339.9Germany Belgibole elektra, Belgibole sp., Carbonocoryphe kelaino, Weania neso, Pseudowaribole steedenensis (37185)
Visean346.7 - 330.3Belgium Carbonocoryphe ogmios (37125)
Visean346.7 - 330.3Germany (Nordrhein Westfalen) Geigibole heisingi, Evagena evae (229490) Weania preacox (229491)
Visean346.7 - 330.3Japan (Okayama-ken) Carbonocoryphe orientalis (79495)
Visean346.7 - 330.3Australia (New South Wales) Aprathia semicircularis (218772) Dechenelloides peoui (229177)
Meramecian342.9 - 335.4USA (Indiana) Spergenaspis salemi (229175)
Asbian339.9 - 335.9United Kingdom (England) Weania anglica, Winterbergia sp. (5966)
Moscovian315.2 - 307.0Canada (Nunavut) Nipponaspis miosilus (62696)
Asselian298.9 - 293.52Thailand Paraphillipsia sp. (133250)
Sakmarian293.52 - 290.1Australia (New South Wales) Doublatia matheri (107626)
Artinskian290.1 - 283.3Australia (Tasmania) Doublatia pyriforme (120242) Doublatia sp., Doublatia pyriforme (120244)
Roadian274.4 - 266.9Australia (New South Wales) Doublatia inflata (120243)
Wordian266.9 - 264.28Japan Weania insulana (224525 224526)
Capitanian264.28 - 259.51Japan Nipponaspis takaizumii, Endops yanagisawai (118971)