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Empidoidea

Insecta - Diptera

Taxonomy
Empidoidea was named by Latreille (1804) [Sensu Chvála 1983]. It is extant. Its type is Empis.

It was assigned to Diptera by Peñalver and Arillo (2007); and to Eremoneura by Yeates et al. (2007), Wiegmann et al. (2011), Shin et al. (2018).

Synonymy list
YearName and author
1804Empidoidea Latreille
2007Empidoidea Peñalver and Arillo
2007Empidoidea Yeates et al.
2011Empidoidea Wiegmann et al.
2018Empidoidea Shin et al.

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—EcdysozoaAguinaldo et al. 1997
—Panarthropoda
phylumArthropodaGravenhorst 1843
—Deuteropoda
—Mandibulata
—PancrustaceaZrzavý and Štys 1997
—AltocrustaceaRegier et al. 2010
superclassAllotriocarida
—Athalassocarida
—LabiocaridaSchwentner et al. 2017
superclassHexapodaLatreille 1825
—Cercofilata
classInsecta
RankNameAuthor
—Dicondylia
—Paranotalia
subclassPterygota()
—NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
—Holometabola
—Aparaglossata
—Panorpida
superorderAntliophora
orderDiptera
—NeodipteraMichelsen 1994
suborderBrachyceraZetterstedt 1842
—Heterodactyla
—Eremoneura
superfamilyEmpidoideaLatreille 1804
superfamilyEmpidoideaLatreille 1804

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Superfm. Empidoidea Latreille 1804
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Fm. Atelestidae Hennig 1970
+
Subfm. Atelestinae Hennig 1970
+
G. Alavesia Waters and Arillo 1999
+
Subg. †Alavesia (Pennonalavesia) Stark et al. 2020
+
†Alavesia (Pennonalavesia) longicornuta Sinclair and Grimaldi 2020
†Alavesia (Pennonalavesia) wimpala Stark et al. 2020
†Alavesia angusta Sinclair and Grimaldi 2020
†Alavesia brevipennae Sinclair and Grimaldi 2020
†Alavesia hadroceria Poinar and Vega 2020
†Alavesia lanceolata Sinclair and Grimaldi 2020
†Alavesia latala Sinclair and Grimaldi 2020
†Alavesia magna Sinclair and Grimaldi 2020
†Alavesia myanmarensis Jouault et al. 2020
†Alavesia pankowskiorum Sinclair and Grimaldi 2020
†Alavesia prietoi Peñalver and Arillo 2007
†Alavesia spinosa Sinclair and Grimaldi 2020
†Alavesia subiasi Waters and Arillo 1999
†Alavesia tripudii Zhang and Xie 2020
†Alavesia zigrasi Sinclair and Grimaldi 2020
Invalid names: Neoalavesia Poinar and Vega 2020 [synonym]
G. †Atelestites Grimaldi and Cumming 1999
+
†Atelestites senectus Grimaldi and Cumming 1999
G. †Dianafranksia Coram et al. 2000
+
†Dianafranksia fisheri Coram et al. 2000
G. †Kurnubempis Kaddumi 2007
+
†Kurnubempis palistinicus Kaddumi 2007
G. †Meghyperites Sinclair and Stark 2024
+
†Meghyperites balticus Sinclair and Stark 2024
G. †Promeghyperus Zhang et al. 2024
+
†Promeghyperus hirtus Zhang et al. 2024
†Promeghyperus muricicaudatus Zhang et al. 2024
G. †Trichinites Hennig 1970
+
†Trichinites cretaceus Hennig 1970
Subfm. Nemedininae Sinclair and Cumming 2006
+
G. †Cretodromia Grimaldi and Cumming 1999
+
†Cretodromia glaesa Grimaldi and Cumming 1999
G. Nemedina Chandler 1981
+
†Nemedina eocenica Sinclair and Arnaud 2001
G. †Nemedromia Grimaldi and Cumming 1999
+
†Nemedromia campania Grimaldi and Cumming 1999
†Nemedromia telescopica Grimaldi and Cumming 1999
†Nemedromia turonia Grimaldi and Cumming 1999
G. †Neoturonius Grimaldi and Cumming 1999
+
†Neoturonius asymmetrus Grimaldi and Cumming 1999
†Neoturonius cretatus Grimaldi and Cumming 1999
†Neoturonius vetus Grimaldi and Cumming 1999
G. †Phaetempis Grimaldi and Cumming 1999
+
†Phaetempis lebanensis Grimaldi and Cumming 1999
G. †Prolatomyia Grimaldi and Cumming 1999
+
†Prolatomyia elongata Grimaldi and Cumming 1999
Fm. Dolichopodidae Latreille 1809 [long-legged fly]
+
Subfm. Achalcinae Grootaert and Meuffels 1997
+
G. Achalcus Loew 1857
G. Anepsiomyia Bezzi 1902
+
†Anepsiomyia planipedia Meunier 1907
Invalid names: Anepsius Loew 1857 [replaced]
G. †Archichrysotus Negrobov 1978
+
†Archichrysotus hennigi Negrobov 1978
†Archichrysotus incompletus Grimaldi and Cumming 1999
†Archichrysotus manitobus Grimaldi and Cumming 1999
†Archichrysotus minor Negrobov 1978
G. †Avenaphora Grimaldi and Cumming 1999
+
†Avenaphora gallica Nel et al. 2016
†Avenaphora hispida Grimaldi and Cumming 1999
G. Condylostylus Bigot 1859
G. †Cretomicrophorus Negrobov 1978
+
†Cretomicrophorus novemundus Grimaldi and Cumming 1999
†Cretomicrophorus piolencensis Nel et al. 2016
Subfm. Diaphorinae Schiner 1864
+
G. Argyra Macquart 1834
+
Invalid names: Porphyrops Meigen 1824 [replaced]
G. Chrysotus Meigen 1824
+
†Chrysotus stanciui Parent 1936
Invalid names: Lyroneurus Loew 1857 [synonym]
G. Diaphorus Meigen 1824
+
†Diaphorus alsiosus Meunier 1910
†Diaphorus fernandi Meuffels and Grootaert 1999
+
Invalid names: Dolichopus soccatus Meunier 1899 [replaced]
G. †Palaeoargyra Meunier 1895
+
†Palaeoargyra mutabilis Meunier 1908
+
Invalid names: Palaeoargyra dytei Evenhuis 1994 [objective synonym]
†Palaeoargyra planipedia Meunier 1907
G. †Prochrysotus Meunier 1907
+
†Prochrysotus magnus Meunier 1907
Subfm. Dolichopodinae Latreille 1809
+
G. Dolichopus Latreille 1796
+
†Dolichopus georgi Meuffels and Grootaert 1999
+
Invalid names: Dolichopus minutus Statz 1940 [replaced]
†Dolichopus incertus Theobald 1937
†Dolichopus miluus Förster 1891
†Dolichopus scitus Statz 1940
†Dolichopus spinosus Statz 1940
Invalid names: Hygroceleuthus Loew 1857 [synonym]
G. Gymnopternus Loew 1857
+
†Gymnopternus bauckhorni Meunier 1914
†Gymnopternus lacustris Bickel 1995
G. Hercostomus Loew 1857
+
†Hercostomus bauckhorni Meunier 1914
†Hercostomus quievreuxi Parent and Quiévreux 1935
G. †Prohercostomus Grichanov 1997
+
†Prohercostomus bickeli Evenhuis 1994
+
Invalid names: Dolichopus vulgaris Meunier 1907 [replaced]
†Prohercostomus ciliatus Meunier 1907
†Prohercostomus devinctus Meunier 1907
†Prohercostomus gracilis Meunier 1907
†Prohercostomus intremulus Meunier 1907
†Prohercostomus inumbratus Meunier 1907
†Prohercostomus meunierianus Evenhuis 1994
+
Invalid names: Dolichopus notabilis Meunier 1907 [replaced]
†Prohercostomus minutus Meunier 1907
+
Invalid names: Prohercostomus smicrus Meuffels and Grootaert 1999 [objective synonym]
†Prohercostomus monotonus Meunier 1907
†Prohercostomus morbosus Meunier 1907
†Prohercostomus negotiosus Meunier 1907
†Prohercostomus noxialis Meunier 1907
G. †Wangia Hong 2002
+
†Wangia trichopoda Hong 1981
Invalid names: Arpactodolichopodites Hong 2002 [nomen nudum], Convexivertex Hong 2002 [nomen nudum], Eoeuryopterites Hong 2002 [nomen nudum], Orbicapitis Hong 2002 [nomen nudum]
G. †Electrochoreutes Badano et al. 2023
+
†Electrochoreutes electroechinus Badano et al. 2023
†Electrochoreutes falculigerus Badano et al. 2023
†Electrochoreutes furcillatus Badano et al. 2023
†Electrochoreutes hamatus Badano et al. 2023
†Electrochoreutes pankowskii Badano et al. 2023
†Electrochoreutes planitibia Badano et al. 2023
†Electrochoreutes trisetigerus Badano et al. 2023
G. †Gujaratmyia Bickel 2022
+
†Gujaratmyia rotunda Bickel 2022
Subfm. Hydrophorinae Lioy 1864
+
G. Thinophilus Wahlberg 1844
Invalid names: Thinophilinae Aldrich 1905 [synonym]
Subfm. Medeterinae Lioy 1864
+
G. Medetera Fischer von Waldheim 1819
+
†Medetera amissa Bickel 2016
†Medetera totolapa Bickel 2016
G. †Medeterites Grichanov 2010
+
†Medeterites atterraneus Nazarau 1994
†Medeterites gestuosus Meunier 1907
†Medeterites gulosus Meunier 1907
†Medeterites latipennis Meunier 1907
†Medeterites molestus Meunier 1907
G. †Palaeosystenus Grichanov et al. 2014
+
†Palaeosystenus succinorum Meunier 1907
G. †Plesiomedetera Grichanov 2023
+
†Plesiomedetera decora Meunier 1907
†Plesiomedetera elegantula Meunier 1907
†Plesiomedetera flammea Meunier 1907
†Plesiomedetera lasciva Meunier 1907
†Plesiomedetera lepida Meunier 1907
†Plesiomedetera mustela Meunier 1907
†Plesiomedetera vana Meunier 1907
G. †Salishomyia Bickel 2019
+
†Salishomyia eocenica Bickel 2019
G. †Systenites Grichanov et al. 2014
+
†Systenites argutus Meunier 1907
†Systenites inclytus Meunier 1907
†Systenites splendidus Meunier 1907
G. Systenus Loew 1857
Tr. Thrypticini Negrobov 1986
+
G. †Paleothrypticus Ngô-Muller et al. 2020
+
†Paleothrypticus eocenicus Ngô-Muller et al. 2020
G. Thrypticus Gerstacker 1864
G. Medeterus Fischer von Waldheim 1819
Subfm. Microphorinae Collin 1960
+
G. †Curvus Kaddumi 2005
+
†Curvus khuludi Kaddumi 2005
G. †Meghyperiella Meunier 1908
+
G. Microphor Macquart 1827
+
†Microphor defunctus Handlirsch 1910
†Microphor eocenica Meunier 1902
†Microphor rusticus Meunier 1908
G. Schistostoma Becker 1902
+
†Schistostoma burmanicum Brooks et al. 2019
†Schistostoma foliatum Brooks et al. 2019
Invalid names: Microphorini Collin 1960 [empty]
G. †Microphorites Hennig 1971
+
†Microphorites deploegi Nel et al. 2004
†Microphorites erikai Bramuzzo and Nel 2017
†Microphorites extinctus Hennig 1971
†Microphorites magaliae Perrichot and Engel 2014
†Microphorites moravicus Tkoč et al. 2016
†Microphorites oculeus Grimaldi and Cumming 1999
†Microphorites pouilloni Ngô-Muller and Nel 2020
†Microphorites similis Grimaldi and Cumming 1999
†Microphorites utrillensis Arillo et al. 2008
Subfm. Neurigoninae Aldrich 1905
+
G. Neurigona Rondani 1856
+
†Neurigona corcula Meunier 1907
G. Oncopygius Loew 1873
+
†Oncopygius gracilior Statz 1940
†Oncopygius venustus Statz 1940
Subfm. Parathalassiinae Chvála 1981
+
G. †Electrophorella Cumming and Brooks 2002
+
†Electrophorella baltica Cumming and Brooks 2002
†Electrophorella grimaldii Cumming and Brooks 2002
G. Microphorella Becker 1909
+
†Microphorella fragilis Cumming and Greenwalt 2022
G. †Retinitus Negrobov 1978
+
†Retinitus nervosus Negrobov 1978
Subfm. Peloropeodinae Robinson 1970
+
G. †Palaeomedeterus Meunier 1895
+
†Palaeomedeterus hirsutus Meunier 1907
†Palaeomedeterus horridus Meunier 1907
+
Invalid names: Sympycnites primaevus Grimaldi and Cumming 1999 [synonym]
†Palaeomedeterus ignavus Meunier 1907
†Palaeomedeterus languidus Meunier 1907
†Palaeomedeterus lassatus Meunier 1908
+
Invalid names: Palaeochrysotus fessus Meunier 1907 [synonym]
†Palaeomedeterus lentus Meunier 1907
†Palaeomedeterus manukyani Grichanov 2025
†Palaeomedeterus meunieri Meuffels and Grootaert 1999
+
Invalid names: Palaeomedeterus culiciformis Meunier 1899 [replaced]
+
Invalid names: Chrysotus concinnus Meunier 1907 [synonym]
†Palaeomedeterus subignavus Grichanov 2025
†Palaeomedeterus sublentus Grichanov 2025
Invalid names: Gheynia Meunier 1899 [synonym], Palaeochrysotus Meunier 1907 [synonym], Sympycnites Grimaldi and Cumming 1999 [synonym]
G. Peloropeodes Wheeler 1890
+
G. †Pristinmicrophor Tang et al. 2019
+
†Pristinmicrophor hukawngensis Tang et al. 2019
G. †Prosystenus Negrobov 1976
+
†Prosystenus zherichini Negrobov 1976
G. †Rajpardia Bickel 2022
+
†Rajpardia grimaldii Bickel 2022
Subfm. Rhaphiinae Bigot 1852
+
G. Nematoproctus Loew 1857
G. Rhaphium Meigen 1803
+
Invalid names: Xiphandrium Loew 1857 [synonym]
Subfm. Sciapodinae Becker 1917
+
G. Amblypsilopus Bigot 1888
+
†Amblypsilopus monicae Bickel 2016
G. Mesorhaga Schiner 1868
+
G. Sciapus Zeller 1842
+
†Sciapus combaluzieri Timon-David 1944
†Sciapus erasmius Meuffels and Grootaert 1999
+
Invalid names: Sciapus amabilis Statz 1940 [replaced]
†Sciapus rottensis Statz 1940
Invalid names: Psilopus Meigen 1824 [replaced]
G. †Wheelerenomyia Meunier 1907
+
†Wheelerenomyia bickeli Negrobov and Selivanova 2003
†Wheelerenomyia corcula Meunier 1908
†Wheelerenomyia eocenica Meunier 1907
†Wheelerenomyia longicerca Negrobov and Selivanova 2003
†Wheelerenomyia originaria Meunier 1907
†Wheelerenomyia pacata Meunier 1907
†Wheelerenomyia parca Meunier 1907
†Wheelerenomyia parva Meunier 1907
†Wheelerenomyia parvula Meunier 1907
†Wheelerenomyia pellucida Meunier 1907
†Wheelerenomyia perattica Meunier 1907
†Wheelerenomyia quadrispinosa Negrobov and Selivanova 2003
†Wheelerenomyia subparva Meunier 1916
†Wheelerenomyia vladimiri Negrobov and Selivanova 2003
Invalid names: Agonosominae Aldrich 1905 [synonym]
Subfm. Sympycninae Aldrich 1905
+
G. Campsicnemus Haliday 1832
G. Chaetogonopteron de Meijere 1914
+
†Chaetogonopteron sobrium Meunier 1910
G. Neoparentia Robinson 1967
+
†Neoparentia chiapensis Bickel 2016
G. Teuchophorus Loew 1857
Invalid names: Campsicneminae Becker 1917 [synonym]
Subfm. Xanthochlorinae Aldrich 1905
+
G. Xanthochlorus Loew 1857
+
†Xanthochlorus tenellus Statz 1940
Invalid names: Argyra debellata Meunier 1907 [nomen dubium], Argyra debilis Meunier 1907 [nomen dubium], Argyra deceptoria Meunier 1907 [nomen dubium], Chrysotus setosus Giebel 1856 [nomen dubium], Eodolichopoditinae Hong 2002 [invalid subgroup], Hygroceleuthus titanus Meunier 1907 [nomen dubium], Lyroneurus venustus Meunier 1907 [nomen dubium], Medetera frauenfeldi Giebel 1856 [nomen dubium], Sciara pusilla Meunier 1899 [nomen nudum], Systenus ciliatus Meunier 1907 [nomen dubium], Thinophilus piraticus Meunier 1907 [nomen dubium], Thrypticus sobrius Meunier 1907 [nomen dubium]
G. †Ecommocydromia Schlüter 1978
+
†Ecommocydromia difficilis Schlüter 1978
Fm. Empididae Latreille 1804 [dagger fly]
+
G. Acallomyia Melander 1928
+
†Acallomyia probolaea Melander 1949
G. †Basilorhagio Ren 1995
+
Subfm. Brachystomatinae Melander 1908
+
Tr. Brachystomatini Melander 1908
+
G. Brachystoma Meigen 1822
+
Invalid names: Brachystoma spinulosa Loew 1850 [nomen nudum]
Tr. Trichopezini Vaillant 1981
+
G. Apalocnemis Philippi 1865
+
†Apalocnemis canadambris Grimaldi and Cumming 1999
†Apalocnemis gracilis Meunier 1908
Invalid names: Timalphes Melander 1928 [synonym]
G. Gloma Meigen 1822
+
†Gloma hirta Loew 1850
Invalid names: Gloma acuticornis Loew 1850 [nomen nudum], Gloma palpata Loew 1850 [nomen nudum]
G. Heleodromia Haliday 1833
+
+
Invalid names: Heleodromia (Illiesiella) Wagner 1985 [replaced]
†Heleodromia delicata Meunier 1908
†Heleodromia dolosa Meunier 1908
†Heleodromia starki Hoffeins et al. 1997
G. †Palaeoparamesia Meunier 1902
+
†Palaeoparamesia proosti Meunier 1902
G. Trichopeza Rondani 1856
+
†Trichopeza sucina Melander 1928
+
Invalid names: Trichopeza longicornis Meunier 1908 [replaced]
G. †Burmitempis Cockerell 1917
+
†Burmitempis halteralis Cockerell 1917
Subfm. Clinocerinae Schiner 1862
+
G. Proclinopyga Melander 1928
+
†Proclinopyga ulrichi Sinclair 2010
Invalid names: Fushunempites Hong 2002 [nomen nudum]
Subfm. Empidinae Latreille 1804
+
Tr. Chelipodini Hendel 1936
+
G. Chelipoda Macquart 1823
+
†Chelipoda vaga Meunier 1908
Invalid names: Lepidomya Bigot 1857 [synonym]
G. Phyllodromia Zetterstedt 1837
+
†Phyllodromia inclusa Shelford 1911
Tr. Empidini Latreille 1804
+
G. Empis Linnaeus 1758
+
Subg. Empis (Acallomyia) Melander 1928
†Empis carbonum Germar 1837
†Empis exilis Meunier 1908
†Empis florissantana Cockerell 1915
†Empis infossa Melander 1949
†Empis inscita Meunier 1908
†Empis macilenta Meunier 1908
†Empis macrophthalma Förster 1891
†Empis mala Meunier 1908
†Empis malefica Meunier 1908
†Empis melia von Heyden 1870
†Empis miocenica Cockerell 1915
†Empis mordax Meunier 1908
†Empis morosella Melander 1928
+
Invalid names: Empis morosa Meunier 1908 [replaced]
†Empis orapaensis Waters 1989
†Empis perdita Cockerell 1916
†Empis personata Meunier 1908
†Empis poeppigi Giebel 1856
†Empis pulvillata Loew 1850
†Empis spinifera Statz 1940
†Empis tritava Melander 1928
+
Invalid names: Empis tristis Meunier 1908 [replaced]
†Empis umbonata Statz 1940
Invalid names: Empis bulbirostris Meunier 1899 [nomen nudum], Empis stilicornis Meunier 1899 [nomen nudum], Empis tibialis Meunier 1899 [nomen nudum]
G. Rhamphomyia Meigen 1822
+
†Rhamphomyia ablata Meunier 1908
†Rhamphomyia aeterna Melander 1949
†Rhamphomyia angusta Meunier 1908
†Rhamphomyia corrupta Meunier 1908
†Rhamphomyia craterae Melander 1949
†Rhamphomyia distans Loew 1850
†Rhamphomyia enena Cockerell 1921
†Rhamphomyia errabunda Meunier 1908
†Rhamphomyia fossa Melander 1949
†Rhamphomyia fulvescens Statz 1940
†Rhamphomyia hypolitha Cockerell 1917
†Rhamphomyia inanimata Melander 1949
†Rhamphomyia infernalis Melander 1949
†Rhamphomyia insolita Meunier 1908
†Rhamphomyia interita Melander 1949
†Rhamphomyia involuta Meunier 1908
†Rhamphomyia media Meunier 1908
†Rhamphomyia morticina Melander 1949
†Rhamphomyia obtusa Meunier 1908
†Rhamphomyia oedaloides Meunier 1908
†Rhamphomyia polymorpha Meunier 1899
†Rhamphomyia porrecta Meunier 1908
†Rhamphomyia rottensis Statz 1940
†Rhamphomyia senecta Melander 1949
†Rhamphomyia sepulta Cockerell 1916
†Rhamphomyia spodites Melander 1949
†Rhamphomyia tumulata Melander 1949
†Rhamphomyia ungulina Loew 1850
Invalid names: Rhamphomyia antipedalis Loew 1850 [nomen nudum], Rhamphomyia formosa Loew 1850 [nomen nudum], Rhamphomyia pteropa Loew 1850 [nomen nudum], Rhamphomyia ptilopa Loew 1850 [nomen nudum], Rhamphomyia remitarsis Loew 1850 [nomen nudum]
G. †Emplita Grimaldi and Cumming 1999
+
†Emplita casei Grimaldi and Cumming 1999
Tr. Hemerodromiini Schiner 1862
+
G. Chelifera Macquart 1823
+
†Chelifera detestata Meunier 1908
G. Hemerodromia Meigen 1822
+
†Hemerodromia grootaerti Plant and Shamshev 2011
Tr. Hilarini Collin 1961
+
G. Hilara Meigen 1822
+
†Hilara heerii Meunier 1915
†Hilara litigiosa Meunier 1908
†Hilara royoi Gil Collado 1926
†Hilara tarda Meunier 1908
G. Hilarempis Bezzi 1905
+
†Hilarempis otagoensis Kaulfuss and Sinclair 2024
G. †Turonempis Grimaldi and Cumming 1999
+
†Turonempis styx Grimaldi and Cumming 1999
Invalid names: Hemerodromiinae Schiner 1862 [synonym]
G. †Hasmona Giebel 1856
+
†Hasmona leo Giebel 1856
G. †Hilarites Heer 1856
+
†Hilarites bellus Heer 1856
G. †Oustaletimyia Meunier 1893
+
†Oustaletimyia succinorum Meunier 1893
G. †Progloma James 1937
+
†Progloma rohweri James 1937
Subfm. †Protempidinae Ussatchov 1968
+
G. †Homalocnemimus Mostovski 1999
+
†Homalocnemimus abjugatus Mostovski 1999
G. †Protempis Ussatchov 1968
+
†Protempis antennata Ussatchov 1968
†Protempis minuta Ren 1998
G. †Protoreogeton Mostovski 1999
+
†Protoreogeton admirabilis Mostovski 1999
G. †Protoedalea Cockerell 1921
+
†Protoedalea brachystoma Cockerell 1921
G. †Thirza Giebel 1856
+
†Thirza naumanni Giebel 1856
Invalid names: Brachystomatidae Melander 1908 [synonym], Tachydromia stilpon Meunier 1899 [nomen dubium]
Fm. Hybotidae Meigen 1820 [dance fly]
+
Subfm. Bicellariinae Sinclair and Cumming 2006
+
G. Hoplocyrtoma Melander 1928
+
†Hoplocyrtoma eocenica Sinclair and Greenwalt 2022
G. †Grimaldipeza Solórzano-Kraemer et al. 2023
+
†Grimaldipeza coelica Solórzano-Kraemer et al. 2023
Subfm. Hybotinae Meigen 1820
+
G. †Eternia Martins-Neto et al. 1992
+
†Eternia papaveroi Martins-Neto et al. 1992
G. †Euthyneuriella Meunier 1908
+
G. Hybos Meigen 1803
+
†Hybos exilis Meunier 1908
†Hybos tenuis Meunier 1908
G. †Palaeoleptopeza Meunier 1908
+
†Palaeoleptopeza gracilis Meunier 1908
G. †Pseudoacarterus Waters 1989
+
G. †Syneproctus Solórzano-Kraemer et al. 2020
+
†Syneproctus caridadi Solórzano-Kraemer et al. 2020
G. †Tremembella Martins-Neto et al. 1992
+
†Tremembella gracilis Martins-Neto et al. 1992
Invalid names: Hybotini Meigen 1820 [invalid subgroup]
Subfm. Ocydromiinae Schiner 1862
+
G. Leptopeza Macquart 1827
+
†Leptopeza clavipes Loew 1850
†Leptopeza concinna Meunier 1908
Invalid names: Leptopeza spinigera Meunier 1899 [nomen nudum]
Subfm. Oedaleinae Chvala 1983
+
G. Oedalea Meigen 1820
+
†Oedalea putida Meunier 1908
†Oedalea robusta Meunier 1908
G. †Palaeoedalea Meunier 1902
+
†Palaeoedalea samlandica Meunier 1902
G. †Pouillonhybos Ngô-Muller et al. 2020
+
†Pouillonhybos venator Ngô-Muller et al. 2020
Subfm. Tachydromiinae Meigen 1822
+
G. †Archaeodrapetiops Martins-Neto et al. 1992
+
†Archaeodrapetiops elongata Martins-Neto et al. 1992
†Archaeodrapetiops mezzalirai Martins-Neto et al. 1992
†Archaeodrapetiops nefera Martins-Neto et al. 1992
†Archaeodrapetiops transversa Martins-Neto et al. 1992
G. †Archiplatypalpus Kovalev 1974
+
G. †Cretoplatypalpus Kovalev 1978
+
†Cretoplatypalpus americanus Grimaldi and Cumming 1999
†Cretoplatypalpus archaeus Kovalev 1978
G. †Drapetiella Meunier 1908
+
†Drapetiella definitum Evenhuis 1994
Tr. Drapetini Collin 1961
+
G. Chaetodromia Chillcott 1983
+
†Chaetodromia pulchra Solórzano Kraemer et al. 2005
G. Chersodromia Walker 1849
+
†Chersodromia dominicana Solórzano Kraemer et al. 2005
G. Drapetis Meigen 1822
+
†Drapetis adelomedos Greenwalt 2019
†Drapetis brevis Meunier 1908
†Drapetis decolorata Meunier 1908
†Drapetis decoratum Meunier 1908
†Drapetis dissentis Solórzano Kraemer et al. 2005
†Drapetis mortuum Meunier 1908
†Drapetis vitiosum Meunier 1908
G. Elaphropeza Macquart 1827
G. †Eodromyia Myskowiak et al. 2018
+
†Eodromyia pumilio Myskowiak et al. 2018
†Eodromyia rovnoensis Shamshev 2025
G. Micrempis Melander 1928
+
†Micrempis brevis Solórzano Kraemer et al. 2005
†Micrempis eocenica Meunier 1908
†Micrempis suspiciosa Meunier 1908
G. Stilpon Loew 1859
+
†Stilpon mexicanus Solórzano Kraemer et al. 2005
G. †Electrocyrtoma Cockerell 1917
+
†Electrocyrtoma burmanica Cockerell 1917
G. †Mesoplatypalpus Grimaldi and Cumming 1999
+
†Mesoplatypalpus carpenteri Grimaldi and Cumming 1999
Tr. Symballophthalmini Sinclair and Cumming 2006
+
G. Symballophthalmus Becker 1889
+
Tr. Tachydromiini Meigen 1822
+
G. Dysaletria Loew 1860
+
†Dysaletria diabolica Meunier 1908
G. Platypalpus Macquart 1827
+
†Platypalpus concitatus Meunier 1908
†Platypalpus eversoris Meunier 1908
†Platypalpus interfectoris Meunier 1908
†Platypalpus predatoris Meunier 1908
G. Tachydromia Meigen 1803
+
Subg. †Tachydromia (Tachypona) Keilbach 1982
†Tachydromia egelata Meunier 1908
†Tachydromia theobaldi Timon-David 1944
†Tachydromia voracis Meunier 1908
Invalid names: Phoneutisca Loew 1863 [synonym]
G. Tachypeza Meigen 1830
+
†Tachypeza primitiva Melander 1949
Invalid names: Corynetinae Kertesz 1909 [synonym]
G. †Megeana Meunier 1899
Fm. Ragadidae Sinclair 2016
+
Subfm. Ragadinae Sinclair 2016
+
G. Ragas Walker 1837
+
†Ragas baltica Sinclair and Hoffeins 2013
†Ragas bizarra Hoffeins et al. 2015
†Ragas electrica Sinclair and Hoffeins 2013
†Ragas eocenica Sinclair and Hoffeins 2013
†Ragas generosa Meunier 1908
†Ragas succinea Sinclair and Hoffeins 2013
†Ragas ulrichi Sinclair and Hoffeins 2013
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Kiessling 2004, Bush and Bambach 2015

Age range

Maximum range based only on fossils: base of the Bajocian to the top of the Holocene or 170.90000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 165.3 Ma

Collections (132 total)


Time interval Ma Country or state Original ID and collection number
Bajocian - Bathonian170.9 - 165.3Mongolia (Bayan-Khongor) Protoreogeton admirabilis (114615)
Kimmeridgian154.8 - 149.2Kazakhstan Homalocnemimus abjugatus (105970) Homalocnemimus abjugatus, Protempis antennata (106101) Homalocnemimus sp. (105971) Homalocnemimus sp., Homalocnemimus abjugatus (105426)
Early/Lower Berriasian143.1 - 141.0United Kingdom (England) Dianafranksia fisheri (125985)
Middle Berriasian141.0 - 139.1United Kingdom (England) Empididae indet. (104008)
Early/Lower Barremian125.77 - 121.4Lebanon Atelestites senectus, Phaetempis lebanensis, Avenaphora hispida (123017) Microphorites similis, Microphorites oculeus (122359) Phaetempis lebanensis (123001) Trichinites cretaceus, Microphorites extinctus (124276)
Late/Upper Barremian125.77 - 121.4China (Liaoning) Protempis minuta (138436)
Late/Upper Barremian125.77 - 121.4China (Hebei) Basilorhagio venustus (107001)
Aptian121.4 - 113.2Mongolia (Govi-Altai) Empididae indet., Hybotidae indet. (122332)
Aptian121.4 - 113.2Russian Federation (Buryatia) Empididae indet. (113349)
Early/Lower Albian113.2 - 110.1Spain Grimaldipeza sp. 1, Grimaldipeza coelica (109438)
Early/Lower Albian113.2 - 110.1Ecuador Microphorites sp. (242350)
Albian113.2 - 100.5Jordan Curvus khuludi, Kurnubempis palistinicus (124847)
Late/Upper Albian106.3 - 100.5France (Poitou-Charentes) Microphorites deploegi (99114)
Late/Upper Albian106.3 - 100.5Spain (Aragón) Microphorites sp. (91626)
Late/Upper Albian106.3 - 100.5Spain (Álava) Alavesia subiasi (108769)
Late/Upper Albian106.3 - 100.5Spain (Asturias) Alavesia prietoi (107510)
Early/Lower Cenomanian100.5 - 93.9Spain (Castellón) Grimaldipeza sp. 2 (228868)
Early/Lower Cenomanian100.5 - 93.9Myanmar (Kachin) Alavesia sp., Alavesia longicornuta, Alavesia zigrasi (123113) Neoalavesia hadroceria (104053)
Early/Lower Cenomanian100.5 - 93.9France Ecommocydromia difficilis (123019)
Early/Lower Cenomanian100.5 - 93.9France (Poitou-Charentes) Dolichopodidae indet. (65422)
Early/Lower Cenomanian100.5 - 93.9Myanmar Alavesia longistylata, Alavesia myanmarensis, Promeghyperus muricicaudatus, Promeghyperus hirtus (165561) Alavesia myanmarensis (179227) Alavesia sp., Alavesia longistylata, Alavesia longicornuta, Alavesia angusta, Alavesia lanceolata, Alavesia magna, Alavesia pankowskiorum, Electrochoreutes pankowskii (205729) Alavesia sp., Schistostoma burmanicum, Schistostoma foliatum, Alavesia myanmarensis, Alavesia brevipennae, Alavesia latala, Alavesia magna, Alavesia pankowskiorum, Alavesia spinosa, Alavesia zigrasi (111688) Alavesia tripudii (216019) Alavesia wimpala (228876) Burmitempis halteralis, Electrocyrtoma burmanica (123020) Microphorites pouilloni (189531) Microphorites pouilloni, Electrochoreutes sp., Electrochoreutes trisetigerus, Electrochoreutes electroechinus, Electrochoreutes falculigerus, Electrochoreutes furcillatus, Electrochoreutes hamatus, Electrochoreutes pankowskii (161726) Pouillonhybos venator (212258) Pristinmicrophor hukawngensis (201881)
Late/Upper Cenomanian100.5 - 93.9Russian Federation Cretoplatypalpus archaeus (35051)
Turonian93.9 - 89.8France Microphorites magaliae (164386)
Turonian93.9 - 89.8USA (New Jersey) Neoturonius asymmetrus (123016) Turonempis styx, Emplita casei, Nemedromia turonia, Neoturonius cretatus, Neoturonius vetus, Cretomicrophorus novemundus, Archichrysotus incompletus (110895)
Turonian93.9 - 89.8Botswana Empis orapaensis, Pseudoacarterus orapaensis (123430)
Early/Lower Santonian85.7 - 83.6France Avenaphora gallica, Cretomicrophorus piolencensis (152462)
Late/Upper Santonian85.7 - 83.6Russian Federation Archichrysotus hennigi, Archiplatypalpus cretaceus (122590) Cretomicrophorus rohdendorfi, Archichrysotus hennigi, Archichrysotus minor, Retinitus nervosus (35056)
Late/Upper Campanian83.6 - 72.2Canada (Manitoba) Nemedromia campania, Prolatomyia elongata, Cretodromia glaesa, Cretoplatypalpus americanus, Mesoplatypalpus carpenteri, Apalocnemis canadambris, Archichrysotus manitobus (122354)
Middle Campanian83.6 - 72.2Canada (Alberta) Nemedromia campania, Nemedromia telescopica (122384)
Paleocene - Middle Eocene66.0 - 37.71Czechia Microphorites moravicus (154314)
Early/Lower Eocene56.0 - 48.07USA (Colorado) Protoedalea brachystoma (121194)
Ypresian56.0 - 48.07India (Gujarat) Dolichopodidae indet., Diaphorinae indet., Palaeomedeterus cambayensis (228738) Dolichopodidae indet., Sympycninae indet., Medeterinae indet., Diaphorinae indet., Palaeomedeterus sp., Palaeomedeterus cambayensis, Gujaratmyia rotunda, Rajpardia grimaldii (124463) Sympycninae indet., Palaeomedeterus sp. (183742)
Ypresian56.0 - 48.07France (Oise) Microphorites erikai, Eodromyia pumilio, Paleothrypticus eocenicus (123850)
Ypresian56.0 - 48.07Canada (British Columbia) Microphorus defunctus (119864)
Ypresian56.0 - 48.07China (Liaoning) Fushunempites furvis, Eocorynetites guchengziensis, Orbicapitis borealis, Eoeuryopterites wanghuacunensis, Convexivertex viridulis, Arpactodolichopodites eocenicus, Eodolichopodites floricopulatus, Eodolichopodites fushunensis, Haodolichopodites fasciatus, Longilabia aroliaris, Sinodolichopodites liaoningensis, Laticopulus arcidorsatus, Paradolichopodites xilutianensis, Sunodolichopodites longus, Columnocorna latilabrosa, Bicercites fushunensis (124820) Lochmocola osterosa, Symballophthalmus clavilabrosus, Dolichopus longiflagellatus, Septocellula trichopoda, Septocellula asiatica, Septocellula fera (124530)
Ypresian56.0 - 48.07USA (Colorado) Rhamphomyia enena (105004)
Bridgerian51.1 - 45.9USA (Wyoming) Dolichopus sp. (121322)
Lutetian48.07 - 41.03USA (Montana) Drapetis adelomedos (207021) Hoplocyrtoma eocenica (203699) Salishomyia eocenica, Microphorella fragilis (135666)
Lutetian48.07 - 41.03Russian Federation Prosystenus zherichini (107340)
Bartonian41.03 - 37.71India (Gujarat) Sympycninae indet., Medeterinae indet., Neurigoninae indet. (228739)
Priabonian37.71 - 33.9Poland Rhamphomyia ungulina, Empis pulvillata, Gloma hirta, Leptopeza clavipes, Rhamphomyia distans, Tachydromia postposita (109625)
Priabonian37.71 - 33.9Lithuania Amesorhaga bickeli, Amesorhaga longicerca, Amesorhaga quadrispinosa, Amesorhaga vladimiri, Systenites inclytus, Palaeosystenus succinorum (113470)
Priabonian37.71 - 33.9Belarus Anepsiomyia atterraneus (173158)
Priabonian37.71 - 33.9Russian Federation Heleodromia starki, Heleodromia foveata, Ragas baltica, Ragas eocenica, Ragas succinea, Ragas ulrichi, Ragas bizarra (123419) Nemedina eocenica (163130)
Priabonian37.71 - 33.9Latvia Medeterites molestus (173420)
Priabonian37.71 - 33.9Ukraine Hemerodromia grootaerti (122997) Meghyperiella porphyropsoides (185682 209304) Prohercostomus noxialis, Prohercostomus meunierianus, Eodromyia rovnoensis (113621)
Priabonian37.71 - 33.9Romania Chrysotus stanciui (163213)
Priabonian37.71 - 33.9Russian Federation (Kaliningrad) Chrysotus setosus, Empis poeppigi, Thirza naumanni (147344) Dolichopodidae indet. (201682) Dolichopus noxialis, Dolichopus vulgaris, Gymnopternus interceptus, Gymnopternus intremulus, Dolichopus monotonus, Dolichopus negotiosus, Phyllodromia rustica, Oustaletimyia succinorum, Drapetis brevis, Drapetis decolorata, Drapetis decoratum, Drapetis vitiosum, Phoneutisca eocenica, Phoneutisca suspiciosa, Tachydromia egelata, Tachydromia voracis, Elaphropeza diabolica, Platypalpus concitatus, Platypalpus predatoris, Platypalpus interfectoris, Platypalpus eversoris, Hemerodromia detestata, Phyllodromia dolosa, Phyllodromia delicata, Lepidomyia vaga, Parathalassiella problematica, Palaeoparamesia proosti, Trichopeza longicornis, Hybos tenuis, Hybos exilis, Oedalea robusta, Leptopeza concinna, Palaeoleptopeza gracilis, Meghyperiella porphyropsoides, Euthyneuriella longirostris, Brachystoma gracilis, Rhamphomyia corrupta, Rhamphomyia ablata, Rhamphomyia involuta, Rhamphomyia ungulina, Rhamphomyia crinitarsis, Rhamphomyia errabunda, Rhamphomyia obtusa, Rhamphomyia insolita, Rhamphomyia angusta, Rhamphomyia media, Hilara litigiosa, Hilara tarda, Hilara macilenta, Empis exilis, Empis inscita, Empis personata, Empis mordax, Empis tristis, Empis pulvillata, Empis morosa, Ragas generosa, Microphorus putidus, Holoclera eocenica, Drapetiella definitum, Palaeochrysotus fessus, Palaeochrysotus lassatus, Gheynius bifurcatus, Diaphorus tertiarius, Nematoproctus originarius, Nematoproctus pacatus, Nematoproctus parcus, Nematoproctus parvus, Nematoproctus parvulus, Psilopus pellucidus, Psilopus perastutulus, Psilopus peratticus, Neurigona corculus, Argyra debellata, Argyra debilis, Argyra deceptoria, Argyra mutabilis, Dolichopus notabilis, Campsicnemus gracilis, Thinophilus piraticus, Wheelerenomyia eocenica, Anepsius planipedius, Nematoproctus subparvus, Palaeoedalea samlandica, Porphyrops succinorum, Porphyrops inclytus, Porphyrops argutus, Achalcus latipennis, Thrypticus sobrius, Prochrysotus magnus, Thrypticus molestus, Thrypticus gestuosus, Thrypticus gulosus, Medeterus elegantulus, Medeterus flammeus, Medeterus lascivus, Medeterus mustela, Chrysotus decorus, Chrysotus concinnus, Palaeochrysotus horridus, Palaeochrysotus hirsutus, Palaeochrysotus ignavus, Palaeochrysotus lentus, Palaeochrysotus languidus, Hygroceleuthus minuta, Gymnopternus devinctus, Gymnopternus inumbratus, Dolichopus morbosus, Systenus ciliatus, Lyroneurus venustus, Brachystoma spinulosa, Rhamphomyia polymorpha, Tachydromia stilpon, Dolichopus soccatus, Palaeomedeterus culiciforme, Palaeoargyra sp. (123927) Dolichopus vulgaris, Phyllodromia sp., Drapetis mortuum, Tachydromia voracis, Platypalpus concitatus, Platypalpus predatoris, Platypalpus interfectoris, Leptopeza concinna, Meghyperiella porphyropsoides, Euthyneuriella longirostris, Rhamphomyia ablata, Rhamphomyia ungulina, Rhamphomyia crinitarsis, Rhamphomyia errabunda, Rhamphomyia porrecta, Rhamphomyia oedaloides, Rhamphomyia insolita, Rhamphomyia media, Empis malefica, Empis mala, Empis morosa, Palaeochrysotus fessus, Palaeochrysotus lassatus, Gheynius bifurcatus, Psilopus perastutulus, Neurigona corculus, Dolichopus notabilis, Xiphandrium splendidum, Porphyrops succinorum, Thrypticus sobrius, Thrypticus molestus, Thrypticus gestuosus, Medeterus vanus, Chrysotus lepidus, Palaeochrysotus hirsutus, Palaeochrysotus ignavus, Palaeochrysotus lentus, Palaeochrysotus languidus, Hygroceleuthus titana, Poecilobothrus ciliatus, Gymnopternus inumbratus (124073) Heleodromia starki (154618) Medeterus frauenfeldi (147348) Meghyperites balticus (110735) Palaeomedeterus ignavus, Palaeomedeterus lassatus, Palaeomedeterus bifurcatus (124849) Plesiomedetera mustela, Plesiomedetera flammea, Medeterites atterraneus, Palaeomedeterus manukyani, Palaeomedeterus subignavus, Palaeomedeterus sublentus (192258) Proclinopyga ulrichi, Gheynia bifurcata, Electrophorella baltica, Ragas electrica, Ragas ulrichi (125512) Sympycnites primaevus, Electrophorella baltica, Electrophorella grimaldii (123493) Wheelerenomyia corcula (124003)
Priabonian37.71 - 33.9Canada (British Columbia) Dolichopus sp. (121480)
Priabonian37.71 - 33.9Germany (Halle) Heleodromia starki (124273)
Chadronian37.0 - 33.9USA (Colorado) Acallomyia probolaea, Rhamphomyia aeterna, Rhamphomyia interita, Rhamphomyia spodites, Rhamphomyia tumulata (106381) Empis miocenica, Empis florissantana (113749) Empis perdita (124692) Gymnopternus lacustris (151374) Progloma rohweri, Rhamphomyia sepulta (106379) Rhamphomyia hypolitha, Empis infossa, Rhamphomyia craterae, Rhamphomyia fossa, Rhamphomyia inanimata, Rhamphomyia infernalis, Tachypeza primitiva (124624) Rhamphomyia morticina (124680) Rhamphomyia senecta (123119)
Early/Lower Oligocene33.9 - 27.3France Dolichopus miluus (121474) Empis macrophthalma (121473)
Early/Lower Oligocene33.9 - 27.3Germany Dolichopus sp. (113964)
MP 2333.9 - 27.3France Hilara heerii (122535)
Rupelian33.9 - 27.3France Hercostomus sp., Hercostomus quievreuxi (146907)
Rupelian33.9 - 27.3Germany Dolichopus incertus (122407)
Chattian27.3 - 23.04Brazil Archaeodrapetiops nefera, Archaeodrapetiops mezzalirai, Archaeodrapetiops transversa, Archaeodrapetiops elongata, Tremembella gracilis, Eternia papaveroi (153054)
Chattian27.3 - 23.04Germany (North Rhine-Westphalia) Empis spinifera, Empis umbonata, Rhamphomyia fulvescens, Rhamphomyia latipennata, Rhamphomyia rottensis, Dolichopus spinosus, Dolichopus minutus, Dolichopus scitus, Oncopygius oligocaenicus, Oncopygius gracilior, Oncopygius venustus, Xanthochlorus tenellus, Sciapus rottensis, Sciapus amabilis (122010)
Chattian27.3 - 23.04France (Bouches-Du-Rhone) Dolichopodidae indet. (157194) Hilara heerii (113712) Hilarites bellus (122876)
Chattian27.3 - 23.04France (Bouches-du-Rhone) Tachydromia theobaldi, Sciapus combaluzieri (155869)
MP 3027.3 - 23.04Germany (Nordrhein-Westfalen) Gymnopternus bankhornius, Empis melia (138696)
MP 3027.3 - 23.04Germany Empis melia (123945)
Aquitanian23.04 - 20.45New Zealand Hilarempis otagoensis (50035)
Early/Lower Miocene23.04 - 15.98Mexico (Chiapas) Chaetodromia pulchra, Peloropeodes paleomexicana (140369) Chrysotus sp. (151865) Diaphorinae indet. (140260) Drapetis dissentis, Diaphorinae indet. (152002) Hercostomus sp., Medetera sp. (140208) Medetera amissa (140312) Medetera sp. (178483) Medetera totolapa (178482)
Early/Lower Miocene23.04 - 15.98Mexico Sympycninae indet., Micrempis brevis, Stilpon mexicanus, Neoparentia chiapensis, Medetera sp., Mesorhaga sp. 1, Mesorhaga sp. 2, Condylostylus sp. 2, Condylostylus sp. 3, Condylostylus sp. 4, Condylostylus sp. 1 (21977)
Early/Lower Miocene23.04 - 15.98Japan Empididae indet., Dolichopodidae indet. (114003)
Early/Lower Miocene23.04 - 15.98Spain (Castellon) Hilara royoi (122583 152716)
Neogene23.04 - 2.58Australia (Queensland) Chaetogonopteron bethnorrisae (139940)
Burdigalian - Langhian20.45 - 13.82Dominican Republic Chersodromia dominicana (152001) Syneproctus caridadi (124429 152045)
Late/Upper Miocene11.63 - 5.333Germany Empis carbonum (117341)
Late/Upper Pleistocene0.129 - 0.0117Ukraine Chrysotus sp. (153476)
Holocene0.0117 - 0.0Tanzania Nematoproctus alsiosus, Teuchophorus sobrius (140788) Phyllodromia inclusa (123991)