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Aetosauria (aetosaur)
Taxonomy
Aetosauria was named by Lydekker (1889). It was considered monophyletic by Nesbitt (2011).
It was corrected as Aetosauri by Jaekel (1910), Jaekel (1911); it was synonymized subjectively with Pseudosuchia by Charig (1967); it was reranked as the unranked clade Aetosauria by Parker (2007), Schoch (2007), Nesbitt (2011).
It was assigned to Loricata by Hay (1902); to Loricati by Jaekel (1910), Jaekel (1911); to Parasuchia by Osborn (1904), Lull (1915), Lull (1917); to Pseudosuchia by Gauthier and Padian (1985); to Thecodontia by Carroll (1988); to Rauisuchiformes by Parrish (1993); to Crocodylotarsi by Heckert and Lucas (1998), Lucas et al. (1998), Heckert and Lucas (2000); to Pseudosuchia by Parker (2007), Schoch (2007); and to Suchia by Sereno and Arcucci (1990), Sereno (1991), Long and Murry (1995), Weinbaum and Hungerbühler (2007), Nesbitt (2011).
It was corrected as Aetosauri by Jaekel (1910), Jaekel (1911); it was synonymized subjectively with Pseudosuchia by Charig (1967); it was reranked as the unranked clade Aetosauria by Parker (2007), Schoch (2007), Nesbitt (2011).
It was assigned to Loricata by Hay (1902); to Loricati by Jaekel (1910), Jaekel (1911); to Parasuchia by Osborn (1904), Lull (1915), Lull (1917); to Pseudosuchia by Gauthier and Padian (1985); to Thecodontia by Carroll (1988); to Rauisuchiformes by Parrish (1993); to Crocodylotarsi by Heckert and Lucas (1998), Lucas et al. (1998), Heckert and Lucas (2000); to Pseudosuchia by Parker (2007), Schoch (2007); and to Suchia by Sereno and Arcucci (1990), Sereno (1991), Long and Murry (1995), Weinbaum and Hungerbühler (2007), Nesbitt (2011).
Synonyms
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Synonymy list
| Year | Name and author |
|---|---|
| 1887 | Episcoposaurus Cope p. 213 |
| 1889 | Aetosauria Lydekker |
| 1892 | Desmatosuchus haploceras Cope |
| 1892 | Episcoposaurus haplocerus Cope p. 129 |
| 1902 | Aetosauria Hay p. 511 |
| 1902 | Episcoposaurus Hay p. 511 |
| 1902 | Episcoposaurus haplocerus Hay p. 511 |
| 1904 | Aetosauria Osborn p. 100 |
| 1910 | Aetosauri Jaekel p. 340 |
| 1911 | Aetosauri Jaekel p. 164 |
| 1915 | Episcoposaurus Huene p. 492 |
| 1915 | Aetosauria Lull p. 98 |
| 1917 | Aetosauria Lull p. 113 |
| 1922 | Episcoposaurus Huene p. 23 |
| 1930 | Episcoposaurus Hay p. 144 |
| 1930 | Episcoposaurus haplocerus Hay p. 144 |
| 1942 | Episcoposaurus Huene p. 243 |
| 1967 | Desmatosuchus haplocerus Charig p. 709 |
| 1985 | Aetosauria Gauthier and Padian pp. 187-189 fig. 1 |
| 1986 | Desmatosuchus haploceros Parrish and Carpenter p. 152 |
| 1988 | Aetosauria Carroll |
| 1990 | Aetosauria Sereno and Arcucci p. 41 |
| 1991 | Aetosauria Sereno p. 11 |
| 1993 | Aetosauria Parrish p. 290 fig. 1 |
| 1995 | Aetosauria Long and Murry p. 66 |
| 1995 | Desmatosuchus haplocerus Long and Murry p. 89 |
| 1998 | Aetosauria Heckert and Lucas |
| 1998 | Aetosauria Lucas et al. |
| 2000 | Aetosauria Heckert and Lucas |
| 2005 | Desmatosuchus haplocerus Irmis p. 75 |
| 2006 | Desmatosuchus haploceras Spielmann et al. pp. 583-584 |
| 2007 | Aetosauria Parker |
| 2007 | Desmatosuchus haplocerus Parker |
| 2007 | Aetosauria Schoch |
| 2007 | Aetosauria Weinbaum and Hungerbühler p. 141 |
| 2011 | Aetosauria Nesbitt p. 195 fig. 51 |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Unr. †Aetosauria Lydekker 1889 [aetosaur]
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G. †Aetosauroides Casamiquela 1960
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†Aetosauroides scagliai Casamiquela 1960
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Invalid names: Aetosauroides inhamandensis Zacharias 1982 [synonym], Aetosauroides subsulcatus Zacarias 1982 [synonym]
Fm. †Batrachopodidae Lull 1904
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G. †Batrachopus Hitchcock 1845
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†Batrachopus argentinus Lull 1942
†Batrachopus bakoenaorum Ellenberger 1974
†Batrachopus deweyi Hitchcock 1843
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Invalid names: Antipus flexiloquus Hitchcock 1858 [synonym], Arachichnus dehiscens Hitchcock 1858 [synonym], Batrachopus gilberti Lapparent and Montenat 1967 [synonym], Batrachopus parvulus Hitchcock 1837 [synonym], Cheirotheroides pilulatus Hitchcock 1858 [synonym], Chelonoides incedens Hitchcock 1858 [synonym], Exocampe arcta Hitchcock 1858 [synonym], Exocampe minima Hitchcock 1862 [synonym], Exocampe ornata Hitchcock 1858 [synonym], Orthodactylus floriferus Hitchcock 1858 [synonym], Orthodactylus introvergens Hitchcock 1858 [synonym], Orthodactylus linearis Hitchcock 1858 [synonym], Palamopus gracilipes Hitchcock 1858 [synonym], Palamopus palmatus Hitchcock 1845 [synonym], Palamopus rogersi Hitchcock 1843 [synonym], Sustenodactylus curvatus Hitchcock 1858 [synonym]
†Batrachopus dispar Lull 1904
†Batrachopus gainensis Kim et al. 2012
†Batrachopus gracilis Hitchcock 1845
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Invalid names: Batrachopus gracilior Hitchcock 1862 [synonym], Shepardia palmipes Hitchcock 1858 [synonym]
†Batrachopus grandis Kim et al. 2020
†Batrachopus pilosus Ellenberger 1970
†Batrachopus rugosus Ellenberger 1970
†Batrachopus supersaltator Ellenberger 1970
†Batrachopus tenuis Ellenberger 1970
†Batrachopus varians Demathieu 1970
Invalid names: Anisichnus Hitchcock 1871 [synonym], Anisopus Hitchcock 1848 [synonym], Arachichnus Hitchcock 1858 [synonym], Batrachoidichnites Hitchcock 1844 [synonym], Cheirotheroides Hitchcock 1858 [synonym], Chelonoides Agassiz 1857 [synonym], Comptichnus Hitchcock 1865 [synonym], Exocampe Hitchcock 1858 [synonym], Macropterna Hitchcock 1848 [synonym], Nanopodiscus Ellenberger 1970 [synonym], Orthodactylus Hitchcock 1858 [synonym], Palamopus Hitchcock 1845 [synonym], Parabatrachopus Lull 1942 [synonym], Shepardia Hitchcock 1858 [synonym], Suchopus Ellenberger 1974 [synonym], Sustenodactylus Lull 1904 [synonym]
G. †Rigalites Huene 1931
G. †Tarsodactylus Hitchcock 1858
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†Tarsodactylus caudatus Hitchcock 1858
†Tarsodactylus expansus Hitchcock 1866
G. †Revueltosaurus Hunt 1989
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†Revueltosaurus callenderi Hunt 1989
Invalid names: Galtonia Hunt and Lucas 1994 [synonym], Pekinosaurus Hunt and Lucas 1994 [synonym], Pekinosaurus olseni Hunt and Lucas 1994 [nomen dubium], Thecodontosaurus gibbidens Cope 1878 [nomen dubium]
Unr. †Stagonolepididae Lydekker 1887
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Subfm. †Aetosaurinae Baur 1887
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G. †Aetosaurus Fraas 1887
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G. †Coahomasuchus Heckert and Lucas 1999
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†Coahomasuchus chathamensis Heckert et al. 2017
†Coahomasuchus kahleorum Heckert and Lucas 1999
Unr. †Typothoracisinae Parker 2007
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Unr. †Paratypothoracisini Parker 2007
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G. †Kocurypelta Czepiński et al. 2021
G. †Paratypothorax Long and Ballew 1985
G. †Tecovasuchus Martz and Small 2006
G. †Venkatasuchus Haldar et al. 2023
G. †Redondasuchus Hunt and Lucas 1991
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†Redondasuchus reseri Hunt and Lucas 1991
†Redondasuchus rineharti Spielmann et al. 2006
G. †Typothorax Cope 1875
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†Typothorax coccinarum Cope 1875
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Invalid names: Episcoposaurus horridus Cope 1887 [synonym], Typothorax antiquum Lucas et al. 2002 [synonym]
†Typothorax meadei Sawin 1947
Invalid names: Argentinosuchus Casamiquela 1960 [nomen dubium], Argentinosuchus bonapartei Casamiquela 1960 [nomen dubium]
Subor. †Desmatosuchia Case 1920
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Subfm. †Desmatosuchinae Huene 1942
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G. †Calyptosuchus Long and Ballew 1985
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†Calyptosuchus wellesi Long and Ballew 1985
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Invalid names: Acompsosaurus wingatensis Mehl 1915 [synonym]
Invalid names: Acompsosaurus Mehl 1915 [synonym]
Unr. †Desmatosuchini Parker 2016
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G. †Desmatosuchus Case 1920
G. †Gorgetosuchus Heckert et al. 2015
G. †Longosuchus Hunt and Lucas 1990
G. †Lucasuchus Long and Murry 1995
G. †Sierritasuchus Parker et al. 2008
Subfm. †Stagonolepininae Heckert and Lucas 2000
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G. †Polesinesuchus Roberto-da-Silva et al. 2014
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†Polesinesuchus aurelioi Roberto-da-Silva et al. 2014
G. †Stagonolepis Agassiz 1844
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†Stagonolepis olenkae Sulej 2010
Invalid names: Ebrachosaurus Kuhn 1936 [synonym]
Unr. †Stagonolepidoidea Parker 2018
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G. †Typlothorax
Invalid names: Episcoposaurinae Huene 1936 [empty], Stegominae Nopcsa 1923 [empty]
Invalid names: Episcoposaurus Cope 1887 [nomen dubium], Episcoposaurus haplocerus Cope 1892 [nomen dubium]
Diagnosis
| Reference | Diagnosis | |
|---|---|---|
| S. J. Nesbitt 2011 | The most inclusive clade containing Aetosaurus ferratus Fraas, 1877, and Desmatosuchus haplocerus Cope, 1892, than to Rutiodon carolinensis Emmons, 1856, Postosuchus kirkpatricki Chatterjee, 1985, Prestosuchus chiniquensis Huene, 1942, Poposaurus gracilis Mehl, 1915, Crocodylus niloticus Laurenti, 1768, Gracilisuchus stipanicicorum Romer, 1972c, and Revueltosaurus callenderi Hunt, 1989 |
Measurements
No measurements are available
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| Source: subo = suborder, subc = subclass, c = class, subp = subphylum | |||||
| References: Benton 1983, Hendy et al. 2009, Carroll 1988 | |||||
Age range
Maximum range based only on fossils: base of the Olenekian to the top of the Late/Upper Aptian or 251.20000 to 113.00000 Ma
Minimum age of oldest fossil (stem group age): 247.2 Ma
Minimum age of oldest fossil (stem group age): 247.2 Ma
Collections (325 total)
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| Olenekian | United Kingdom (England) | Aetosauripus sp. (184045) | |
| Anisian | Netherlands (Gelderland) | Sustenodactylus hollandicus (96875) | |
| Anisian - Ladinian | France (Rhône-Alpes) | Batrachopus sp. (199929) | |
| Anisian - Ladinian | France (Auvergne-Rhône-Alpes) | Batrachopus varians (31310) | |
| Carnian | Argentina (San Juan) | Stagonolepis sp., Argentinosuchus bonapartei (92317) | |
| Carnian | Brazil (Rio Grande do Sul) | Polesinesuchus aurelioi (175578) | |
| Carnian | Morocco (Marrakech-Safi) | Longosuchus meadei (58618) | |
| Carnian | USA (North Carolina) | Desmatosuchinae indet., Lucasuchus sp., Gorgetosuchus pekinensis, Coahomasuchus chathamensis (169982) Desmatosuchus sp. (93563) Ornithischia indet. (74205) Typothorax sp. (46567) | |
| Carnian | USA (Wyoming) | Desmatosuchus sp. (91590) | |
| Carnian | USA (Texas) | Stagonolepis wellesi, Desmatosuchus haplocerus (84271) | |
| Carnian | Zimbabwe (Mashonaland Central) | Aetosauria indet. (228332) | |
| Carnian | Germany (Bayern) | Aetosauripus schlauersbachensis (139738) Ebrachosaurus singularis (97736) | |
| Carnian | Canada (Nova Scotia) | Aetosauridae indet. (107286) | |
| Carnian - Norian | Argentina (San Juan) | Aetosauroides scagliai (92315) Aetosauroides scagliai, Argentinosuchus bonapartei, Stagonolepis sp. (92316) Stagonolepididae indet., Aetosauroides scagliai (92318) | |
| Carnian - Norian | USA (North Carolina) | Stegomus sp. (32343) Stegomus sp., Stegomus arcuatus (89149) | |
| Carnian - Norian | Poland (Opole) | Stagonolepis sp. (61448 82992) | |
| Carnian - Norian | USA (Texas) | Aetosauria indet. (12942) Coahomasuchus kahleorum (92093) Coahomasuchus kahleorum, Lucasuchus hunti, Desmatosuchus sp. (22706) Desmatosuchus haplocerus (71982 85485) Desmatosuchus spurensis, Paratypothorax sp. (71977) Episcoposaurus haplocerus (135653) Longosuchus meadi, Desmatosuchus haplocerus (98605) Lucasuchus hunti, Desmatosuchus sp. (22707) Parasuchia indet. (71976) Paratypothorax sp. (105766) Phytosauria indet. (135662) Stagonolepis wellesi, Desmatosuchus sp. (98596) Tecovasuchus sp., Paratypothorax sp. (105769) Theropoda indet. (71979) | |
| Carnian - Norian | Brazil (Rio Grande do Sul) | Hoplitosuchus raui (72086 72090) Stagonolepis robertsoni (92320) Stagonolepis robertsoni, Aetosauria new species (92319) | |
| Carnian - Norian | India (Telangana) | Typothorax sp. (112285) | |
| Carnian - Norian | United Kingdom (Scotland) | Stagonolepis robertsoni (57948 89005) | |
| Late/Upper Triassic | Algeria (Illizi) | Desmatosuchus sp., Paratypothorax sp. (107214) | |
| Late/Upper Triassic | USA (New Jersey) | Anisichnus gracilis (91694) | |
| Late/Upper Triassic | Chile (Antofagasta) | Aetosauria indet. (170677) Chilenosuchus forttae (170676) | |
| Tuvalian | Poland (Opole) | Stagonolepis sp. (61449) | |
| Tuvalian | Argentina (San Juan) | Aetosauroides sp. (24877) | |
| Norian | Switzerland (Aargau) | Aetosaurus sp. (22680) | |
| Norian | USA (New Jersey) | Batrachopus sp. (187287) Stegomus arcuatus (89131 92082) | |
| Norian | USA (New Mexico) | Acompsosaurus wingatensis (93559) Desmatosuchus haplocerus (93618) Desmatosuchus sp. (49499 93570) Desmatosuchus sp., Typothorax sp. (93604) Episcoposaurus horridus (94150) Neoaetosauroides sp. (93672) Paratypothorax sp. (93603) Paratypothorax sp., Typothorax coccinarum (52503) Paratypothorax sp., Typothorax sp. (80926) Phytosauridae indet. (94100) Redondasuchus reseri (83737) Redondasuchus rineharti (89969) Revueltosaurus callenderi (223082) Stagonolepidae indet. (75661 75662) Stagonolepididae indet. (55248 93607 93608 93649) Stagonolepididae indet., Calyptosuchus sp., Desmatosuchus sp. (93611) Stagonolepididae indet., Desmatosuchus sp. (93605) Stagonolepididae indet., Longosuchus meadei (93565) Stagonolepididae indet., Typothorax antiquum (93584 93585 93589) Stagonolepididae indet., Typothorax coccinarum (93583 93590 179601) Stagonolepididae indet., Typothorax sp. (83738) Stagonolepis sp. (75663 93576) Stagonolepis sp., Desmatosuchus sp., Stagonolepidae indet. (75651) Stagonolepis wellesi (179082) Tecovasuchus sp., Paratypothorax sp. (75653) Typothorax antiquum (93580 93586) Typothorax coccinarum (47126 80925 93591 93592 93593 93594 94151) Typothorax coccinarum, Revueltosaurus callenderi, Stagonolepididae indet. (48516) Typothorax coccinarum, Rioarribasuchus chamaensis (74202) Typothorax sp. (54007 84272 89256 93581 179083 223030) Typothorax sp., Typothorax coccinarum (93588) | |
| Norian | USA (Arizona) | Acaenasuchus geoffreyi (196662 215309) Acaenasuchus geoffreyi, Desmatosuchus haplocerus, Fabrosauridae indet. (52502) Acaenasuchus geoffreyi, Desmatosuchus spurensis, Aetosauria indet. (223390) Acaenasuchus geoffreyi, Stagonolepis wellesi, Desmatosuchus haplocerus, Tecovasuchus sp. (52499) Acaenasuchus geoffreyi, Stagonolepis wellesi, Revueltosaurus sp. (92850) Acaenasuchus geoffreyi, Stagonolepis wellesi, Tecovasuchus chatterjeei, Desmatosuchus sp. (39219) Adamanasuchus eisenhardtae (93478) Aetosauria indet. (48547) Desmatosuchus haplocerus (93352 93353 93355 93359 93451 218744) Desmatosuchus haplocerus, Paratypothorax sp. (84265) Desmatosuchus spurensis (94497) Paratypothorax sp. (93112 93462) Paratypothorax sp., Desmatosuchus haplocerus, Stagonolepis wellesi (52501) Paratypothorax sp., Stagonolepis wellesi (93232) Paratypothorax sp., Typothorax coccinarum (57686 84266 93430 93431 93432 93460) Paratypothorax sp., Typothorax coccinarum, Desmatosuchus chamaensis (93443) Revueltosaurus callenderi (53513 57687 92811 92827) Revueltosaurus callenderi, Acaenasuchus geoffreyi, Stagonolepis wellesi, Tecovasuchus sp., Tecovasuchus chatterjeei, Desmatosuchus haplocerus (39160) Scutarx deltatylus (182617 182618) Stagonolepididae indet. (93404 93571) Stagonolepis sp., Desmatosuchus sp. (22705) Stagonolepis sp., Paratypothorax sp. (93230) Stagonolepis wellesi (84264 92824 92912 93113 93115 93117 93118 93119 93120 93123 93126 93127 93129 93135 93226 93227 93228 93354 93356) Stagonolepis wellesi, Desmatosuchus haplocerus (52500 57683 84268 92322 92821 92922 92923 93116 93131 93229 93541) Typothorax coccinarum (48421 48423 48546 57684 84017 93393 93394 93395 93396 93400 93412 93415 93416 93423 93424 93425 93426 93427 93436 93439 93441 93450 93453 93455 93456 93458 93459 93463 93465 93466 93469 218116) Typothorax coccinarum, Prosauropoda indet., Ornithischia indet. (49500) Typothorax coccinarum, Revueltosaurus callenderi (57685) Typothorax coccinarum, Stagonolepis wellesi (92819) Typothorax sp. (88338) | |
| Norian | Germany (Baden-Württemberg) | Aetosauria indet. (22683) Paratypothorax andressorum (106363) Paratypothorax sp., Aetosauria indet. (105834) | |
| Norian | Argentina (La Rioja) | Neoaetosauroides engaeus (101991 105691 105692 105695 105696 105697) | |
| Norian | Germany (Baden-Württenburg) | Aetosaurus ferratus (92087 92088) | |
| Norian | Germany (Baden-Wurttemberg) | Paratypothorax andressi (47785) Paratypothorax sp. (138914) | |
| Norian | USA (Pennsylvania) | Anisichnus gracilis (55413) Batrachopus sp. (84111) Thecodontosaurus gibbidens (59371) | |
| Norian | USA (Texas) | Coahomasuchus sp. (194471) Desmatosuchus haplocerus (84269) Paratypothoracisini indet. (218151) Paratypothoracisini indet., Scutarx deltatylus (218145) Rioarribasuchus sp., Garzapelta muelleri (79947) Stagonolepididae indet., Paratypothorax sp., Calyptosuchus wellesi, Typothorax coccinarum, Desmatosuchus haplocerus (13273) Stagonolepididae indet., Pekinosaurus olseni (13266 48424) Stagonolepis sp., Revueltosaurus callenderi, Aetosauria indet. (218126) Stagonolepis wellesi, Desmatosuchus haplocerus (94493) Typothorax coccinarum (218143) Typothorax coccinarum, Aetosaurinae indet. (218144) Typothorax coccinarum, Desmatosuchinae indet. (218149) Typothorax sp. (196537) | |
| Norian | USA (North Carolina) | Revueltosaurus sp., Galtonia gibbidens, Revueltosaurus sp. (99541) | |
| Norian | Greenland | Aetosaurus ferratus, Paratypothorax andressi (26998) | |
| Norian | Poland (Opole) | Kocurypelta silvestris (140077) | |
| Norian | USA (Colorado) | Stenomyti huangae (182593) | |
| Norian | Argentina (Mendoza) | Chirotherium sp. (206089) | |
| Norian | USA (Utah) | Stagonolepis sp. (92321) | |
| Norian - Rhaetian | USA (New Jersey) | Batrachopus sp. (187321) | |
| Norian - Rhaetian | Lesotho (Mohale's Hoek) | Comptichnus sp. (56020) | |
| Norian - Rhaetian | USA (Colorado) | Aetosauria indet., Aetosaurus ferratus (92089) | |
| Norian - Rhaetian | Poland (Śląskie) | Aetosauria indet. (132933) | |
| Norian - Rhaetian | India | Venkatasuchus armatum (131840) | |
| Norian - Rhaetian | USA (Pennsylvania) | Orthodactylus howelli (68705) | |
| Lacian - Alaunian | USA (Virginia) | Aetosauria indet. (68889) | |
| Lacian - Alaunian | USA (New Mexico) | Typothorax sp., Aetosaurus arcuatus, Aetosaurus sp. (92090) | |
| Lacian - Sevatian | USA (New Mexico) | Typothorax coccinarum, Desmatosuchus chamaensis, Procolophonidae indet. (62311) | |
| Alaunian | Germany (Baden-Württemberg) | Aetosaurus crassicauda (47800) | |
| Alaunian | USA (Connecticut) | Stegomus arcuatus (91750) | |
| Alaunian | Italy (Lombardia) | Aetosaurus ferratus (59639) | |
| Alaunian - Sevatian | USA (Arizona) | Stagonolepis sp. (93480) | |
| Sevatian | USA (Arizona) | Typothorax coccinarum (93392) | |
| Rhaetian | France (Pays de la Loire) | Pseudosuchia indet. (70801) | |
| Rhaetian | USA (New Jersey) | Batrachopus deweyi, Batrachopus bellus (194093) | |
| Rhaetian | USA (Pennsylvania) | Batrachopus sp. (62768) | |
| Rhaetian | USA (Arizona) | Typothorax coccinarum (84009 84012 84013 84014 84015 84016 198618) | |
| Rhaetian - Hettangian | USA (Arizona) | Batrachopus deweyi (137291) | |
| Rhaetian - Hettangian | USA (Utah) | Batrachopus sp. (57065 223706) | |
| Early/Lower Jurassic | China (Nei Mongol) | Batrachopus sp. (107973) | |
| Jurassic | Colombia (Santander) | Batrachopus sp. (55322) | |
| Hettangian | France (Pays de la Loire) | Pseudosuchia indet. (70802) | |
| Hettangian | USA (Connecticut) | Batrachopus deweyi (51257) Batrachopus dispar (55741) Shepardia palmipes (51256) | |
| Hettangian | USA (Utah) | Batrachopus sp. (121305 121321) Batrachopus sp., Batrachopus sp. (56131) | |
| Hettangian | France (Region Midi-Pyrenees) | Batrachopus deweyi (31306) | |
| Hettangian | USA (Massachusetts) | Apatichnus bellus (53030) Shepardia palmipes, Anisopus sp., Anisopus gracilis, Anisopus gracilior, Anisopus deweyanus, Antipus flexiloquus, Comptichnus obesus, Exocampe sp., Tarsodactylus caudatus, Chelonoides incedens, Macropterna sp., Macropterna divaricans, Macropterna gracilipes, Macropterna vulgaris, Exocampe arcta, Exocampe ornata, Orthodactylus floriferus, Orthodactylus introvergens, Arachichnus sp., Apatichnus bellus, Batrachopus dispar (52986) | |
| Hettangian | USA (Colorado) | Batrachopus sp. (193011) | |
| Early/Lower Hettangian | USA (Connecticut) | Batrachopus sp. (192863) | |
| Early/Lower Hettangian | Canada (Nova Scotia) | Batrachopus sp. (138976 138978 227835) Batrachopus spp. (138977) | |
| Middle Hettangian | France (Occitanie) | Batrachopus sp. (190122) | |
| Hettangian - Sinemurian | USA (Connecticut) | Anisopus barrattii, Anisopus gracilis, Anisopus deweyanus (53033) Comptichnus sp., Exocampe ornata (53000) Macropterna divaricans, Macropterna rhynchosauroidea, Macropterna vulgaris, Exocampe ornata, Arachichnus dehiscens, Palamopus rogersi (53023) | |
| Hettangian - Sinemurian | Romania (Caraș-Severin) | Batrachopus deweyi (238761) | |
| Hettangian - Sinemurian | USA (Massachusetts) | Anisopus deweyanus (52994) Anisopus gracilis, Macropterna divaricans, Macropterna rhynchosauroidea, Macropterna vulgaris, Sauroidichnites palmatus, Ornithoidichnites rogersi (52985) Anisopus gracilis, Orthodactylus sp., Anisopus deweyanus, Orthodactylus linearis, Stenodactylus curvatus, Arachichnus dehiscens, Apatichnus bellus, Exocampe minima (52987) Anisopus sp., Anisopus gracilis, Orthodactylus sp., Tarsodactylus sp., Exocampe ornata (52998) Batrachopus deweyi, Anisopus sp. (52999) Batrachopus deweyi, Macropterna vulgaris (52995) Shepardia palmipes, Sauroidichnites deweyi, Anisopus gracilis, Macropterna vuigaris, Macropterna divaricans, Palamopus gracilipes, Macropterna vulgaris, Exocampe arcta, Exocampe ornata, Palamopus palmatus, Palamopus rogersi (52996) Tarsodactylus expansus (91578) | |
| Hettangian - Sinemurian | Lesotho (Quthing) | Nanopodiscus tenuis, Plateotetrapodiscus rugosus (84064) Suchopus bakoenaorum (84072) | |
| Sinemurian | France (Pays de la Loire) | Batrachopus gilberti (200113) | |
| Sinemurian - Pliensbachian | USA (Utah) | Batrachopus sp. (223699) | |
| Pliensbachian | USA (Utah) | Batrachopus sp. (229818) | |
| Pliensbachian | Lesotho (Leribe) | Theromorpha indet. (56001) | |
| Pliensbachian - Toarcian | USA (Idaho) | Batrachopus sp. (195189) | |
| Pliensbachian - Toarcian | USA (Utah) | Batrachopus sp. (201625) | |
| Berriasian - Hauterivian | Brazil (Paraíba) | Batrachopodidae indet. (58790) | |
| Neocomian | Argentina (San Luis) | Parabatrachopus argentina (55479) | |
| Aptian | South Korea (Gyeongsangnam-do) | Batrachopus grandis (220038 220039 220040) | |
| Aptian - Albian | South Korea (Gyeongsangnam-do) | Haenamichnus gainensis (59177 220049) | |
| Late/Upper Aptian | Angola (Lunda Sul) | Angolaichnus adamanticus (190121) | |
| Early/Lower Aptian | United Kingdom (England) | Heliocanthus sp. (235315) |