PBDB Taxon

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Paoliidae

Insecta - Paoliida - Paoliidae

Taxonomy

Paoliidae was named by Handlirsch (1906). Its type is Paolia.

It was assigned to Palaeodictyoptera by Handlirsch (1906), Handlirsch (1919), Handlirsch (1922); to Cacurgoidea by Kukalová (1958); to Protorthoptera by Brauckmann (1984), Brauckmann et al. (1985), Carpenter (1992); and to Paoliida by Prokop et al. (2012), Prokop et al. (2014), Prokop et al. (2014), Nel et al. (2023).

Synonyms

Aliculidae was named by Storozhenko (1997).

It was replaced with Permulidae by Aristov and Storozhenko (2011); it was synonymized subjectively with Permotermopsidae by Aristov (2015).

It was assigned to Protoperlina by Storozhenko (1997), Storozhenko (1998), Aristov (2008), Aristov (2009), Aristov (2009).
Protoblattinidae was named by Meunier (1909).

It was synonymized subjectively with Paoliidae by Prokop et al. (2014).

It was assigned to Grylloblattida by Béthoux et al. (2005).
Kortshakoliidae was named by Storozhenko (1997).

It was synonymized subjectively with Permotermopsidae by Aristov (2015).

It was assigned to Grylloblattina by Storozhenko (1997), Storozhenko (1998), Aristov (2009).
Permotermopsidae was named by Martynov (1937).

It was synonymized subjectively with Ideliidae by Storozhenko (1992); it was synonymized subjectively with Paoliidae by Nel and Poschmann (2021).

It was assigned to Protorthoptera by Carpenter (1992); to Grylloblattina by Storozhenko (1997), Storozhenko (1998), Aristov (2009), Aristov (2009); and to Eoblattida by Aristov (2015), Aristov (2015).
Idelidae was named by Zalessky (1929).

It was corrected as Ideliidae by Riek (1954), Carpenter (1992), Storozhenko (1997), Storozhenko (1998), Aristov (2004), Aristov (2009), Aristov (2015); it was synonymized subjectively with Paoliidae by Prokop et al. (2014), Nel and Poschmann (2021).

It was assigned to Paraplecoptera by Martynov (1940); to Protorthoptera by Zalessky (1929), Martynov (1937), Riek (1954), Carpenter (1992); to Grylloblattina by Storozhenko (1997), Storozhenko (1998), Aristov (2004), Aristov (2009); and to Eoblattida by Aristov (2015).
Permulidae was named by Aristov and Storozhenko (2011).

It was synonymized subjectively with Permotermopsidae by Aristov (2015); it was synonymized subjectively with Paoliidae by Nel and Poschmann (2021).

It was assigned to Protoperlina by Aristov and Storozhenko (2011).
Herbstialidae was named by Schmidt (1952). Its type is Herbstiala.

It was synonymized subjectively with Paoliidae by Prokop et al. (2014).

It was assigned to Protorthoptera by Carpenter (1992).
Rachimentomonidae was named by Zalessky (1939). Its type is Rachimentomon.

It was synonymized subjectively with Ideliidae by Carpenter (1992), Storozhenko (1997), Storozhenko (1998).

It was assigned to Protoblattoidea by Zalessky (1939).

Synonymy list

Year	Name and author
1906	Paoliidae Handlirsch p. 114
1909	Protoblattinidae Meunier p. 48
1919	Paoliidae Handlirsch p. 532
1922	Paoliidae Handlirsch p. 49
1929	Idelidae Zalessky p. 21
1937	Idelidae Martynov p. 23
1937	Permotermopsidae Martynov p. 84
1939	Rachimentomonidae Zalessky p. 55
1940	Idelidae Martynov p. 19
1952	Herbstialidae Schmidt p. 165
1954	Ideliidae Riek p. 161
1958	Paoliidae Kukalová p. 935
1984	Paoliidae Brauckmann
1985	Paoliidae Brauckmann et al. p. 60
1992	Herbstialidae Carpenter
1992	Paoliidae Carpenter p. 101
1992	Ideliidae Carpenter p. 118
1992	Permotermopsidae Carpenter p. 124
1997	Aliculidae Storozhenko p. 8
1997	Ideliidae Storozhenko p. 11
1997	Kortshakoliidae Storozhenko p. 12
1997	Permotermopsidae Storozhenko p. 16
1998	Aliculidae Storozhenko p. 45
1998	Ideliidae Storozhenko p. 45
1998	Kortshakoliidae Storozhenko p. 45
1998	Permotermopsidae Storozhenko p. 46
2004	Ideliidae Aristov
2005	Protoblattinidae Béthoux et al. p. 127
2008	Aliculidae Aristov
2009	Aliculidae Aristov p. 178
2009	Ideliidae Aristov p. 178
2009	Kortshakoliidae Aristov p. 178
2009	Permotermopsidae Aristov p. 178
2009	Aliculidae Aristov p. 643
2009	Permotermopsidae Aristov p. 645
2011	Permulidae Aristov and Storozhenko p. 113
2012	Paoliidae Prokop et al.
2014	Paoliidae Prokop et al. p. 602
2014	Paoliidae Prokop et al. p. 159
2015	Ideliidae Aristov p. 21
2015	Permotermopsidae Aristov p. 25
2023	Paoliidae Nel et al. p. 306

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Ecdysozoa	Aguinaldo et al. 1997
—	Panarthropoda
phylum	Arthropoda	Latreille 1829
—	Deuteropoda
—	Mandibulata
—	Pancrustacea	Zrzavý and Štys 1997
—	Altocrustacea	Regier et al. 2010
superclass	Allotriocarida
—	Athalassocarida

Rank	Name	Author
—	Labiocarida	Schwentner et al. 2017
superclass	Hexapoda	Latreille 1825
—	Cercofilata
class	Insecta
—	Dicondylia
—	Paranotalia
subclass	Pterygota	()
—	Neopterygota	Crampton 1924
infraclass	Neoptera	Martynov 1923
—	Polyneoptera	Martynov 1938
order	Paoliida
family	Paoliidae
family	Paoliidae

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Fm. †Paoliidae Handlirsch 1906

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G. †Acropermula Aristov and Storozhenko 2011

†Acropermula acra Kukalová 1964

G. †Aenigmidelia Sharov 1961

†Aenigmidelia incredibilis Sharov 1961

G. †Archidelia Sharov 1961

†Archidelia elongata Sharov 1961

†Archidelia ovata Sharov 1961

G. †Avionpaolia Boderau et al. 2024

†Avionpaolia amansfossilia Boderau et al. 2024

G. †Carbonidelia Nel et al. 2023

†Carbonidelia gallica Nel et al. 2023

G. †Darekia Prokop et al. 2012

†Darekia sanguinea Prokop et al. 2012

G. †Herbstiala Schmidt 1952

†Herbstiala herbsti Schmidt 1952

G. †Holasicia Kukalová 1958

†Holasicia rasnitsyni Brauckmann 1984

†Holasicia vetula Kukalová 1958

G. †Idelina Storozhenko 1992

†Idelina kamensis Storozhenko 1992

†Idelina mala Aristov 2015

G. †Kemperala Brauckmann 1984

†Kemperala hagenensis Brauckmann 1984

G. †Khosaridelia Storozhenko 1992

†Khosaridelia martynovi Storozhenko 1992

G. †Kochopteron Brauckmann 1984

†Kochopteron hoffmannorum Brauckmann 1984

G. †Kortshakolia Sharov 1961

†Kortshakolia ideliformis Sharov 1961

G. †Mertovia Prokop and Nel 2007

†Mertovia sustai Kukalová 1958

G. †Micaidelia Aristov 2004

†Micaidelia minutissima Aristov 2004

G. †Mongoloidelia Storozhenko 1992

†Mongoloidelia gorochovi Storozhenko 1992

G. †Olinka Kukalová 1958

†Olinka modica Kukalová 1958

G. †Paolekia Riek 1976

†Paolekia perditae Riek 1976

G. †Paolia Smith 1871

†Paolia vetusta Smith 1871

G. †Paoliola Handlirsch 1919

†Paoliola gurleyi Scudder 1885

G. †Permotermopsis Martynov 1937

†Permotermopsis pectinata Martynov 1937

†Permotermopsis roseni Martynov 1937

G. †Permula Handlirsch 1919

†Permula lebachensis von Schlechtendal 1913

Invalid names: Permula lebachensis Handlirsch 1919 [objective synonym]

†Permula simpliciradius Poschmann and Nel 2020

Invalid names: Alicula von Schlechtendal 1913 [replaced], Permomertovia Poschmann and Nel 2020 [synonym]

G. †Protoblattina Meunier 1909

†Protoblattina bouvieri Meunier 1909

G. †Pseudofouquea Handlirsch 1906

†Pseudofouquea cambrensis Allen 1901

G. †Silesiapteron Prokop et al. 2014

†Silesiapteron jarmilae Prokop 2014

G. †Sojanidelia Storozhenko 1992

†Sojanidelia fasciata Storozhenko 1992

†Sojanidelia floralis Rasnitsyn 1996

†Sojanidelia florea Rasnitsyn 1996

†Sojanidelia fluctuosa Storozhenko 1992

†Sojanidelia kostinae Sharov 1961

†Sojanidelia lineata Storozhenko 1992

†Sojanidelia longula Storozhenko 1992

†Sojanidelia maculosa Storozhenko 1992

†Sojanidelia makarkini Storozhenko 1992

†Sojanidelia multimediana Storozhenko 1992

†Sojanidelia nana Storozhenko 1992

†Sojanidelia parvula Storozhenko 1992

†Sojanidelia striata Storozhenko 1992

†Sojanidelia vorkutensis Storozhenko 1992

G. †Stenaropodites Martynov 1928

†Stenaropodites magna Storozhenko 1992

†Stenaropodites mistshenkoi Storozhenko 1992

†Stenaropodites permiakovae Zalessky 1929

†Stenaropodites reticulata Martynov 1928

†Stenaropodites sojanensis Storozhenko 1992

Invalid names: Idelia Zalessky 1929 [synonym]

G. †Sustaia Kukalová 1958

†Sustaia impar Kukalová 1958

G. †Sylvidelia Martynov 1940

†Sylvidelia latipennis Martynov 1940

G. †Zdenekia Kukalová 1958

†Zdenekia grandis Kukalová 1958

†Zdenekia occidentalis Laurentiaux-Vieira and Laurentiaux 1986

†Zdenekia silesiensis Prokop et al. 2012

Invalid names: Herbstialidae Schmidt 1952 [synonym], Ideliidae Zalessky 1929 [synonym], Permotermopsidae Martynov 1937 [synonym], Permulidae Aristov and Storozhenko 2011 [synonym], Protoblattinidae Meunier 1909 [synonym]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

chitinc

Environment:

terrestrialc

Locomotion:

actively mobilep

Created:

2017-04-17 09:56:18

Modified:

2017-04-17 09:56:18

Source: c = class, p = phylum

References: Bush and Bambach 2015, Kiessling 2004

Age range: base of the Marsdenian to the top of the Norian or 321.30000 to 205.70000 Ma

Collections (50 total)

Time interval	Ma	Country or state	Original ID and collection number
Marsdenian	321.3 - 320.0	Germany (Ruhr)	Kochopteron hoffmannorum (155261) Paoliidae indet., Kemperala hagenensis, Holasicia rasnitsyni (155260)
Marsdenian - Yeadonian	321.3 - 318.7	Germany	Kochopteron hoffmannorum (118176)
Yeadonian	320.0 - 318.7	USA (Indiana)	Paolia gurleyi (114466) Paolia vetusta (114465)
Westphalian A	318.7 - 316.9	Netherlands	Paolia vetusta (157406)
Westphalian A	318.7 - 316.9	Czech Republic	Holasicia sustai (124364) Paoliidae indet., Holasicia vetula, Olinka modica, Zdenekia grandis (114184) Sustaia impar (157231)
Westphalian A	318.7 - 316.9	Germany	Herbstiala herbsti (139801)
Westphalian A	318.7 - 316.9	Belgium	Zdenekia occidentalis (157028)
Westphalian A	318.7 - 316.9	United Kingdom (Lancashire)	Pseudofouquea sp. (172765)
Westphalian A	318.7 - 316.9	Poland	Paoliidae indet., Zdenekia silesiensis, Darekia sanguinea, Silesiapteron jarmilae (107805)
Westphalian B	316.9 - 314.6	United Kingdom (Wales)	Fouquea cambrensis (123946)
Westphalian C - Westphalian D	314.6 - 307.4	France	Paoliidae indet., Paolia sp., Avionpaolia amansfossilia (122504)
Gzhelian	303.7 - 298.9	France	Carbonidelia gallica (228605) Protoblattina bouvieri (121979)
Asselian	298.9 - 293.52	Germany	Alicula lebachensis (138375) Permomertovia simpliciradius (112604) Permula lebachensis (222678)
Asselian	298.9 - 293.52	Czech Republic	Permula acra (114186)
Kungurian	283.3 - 274.4	Russian Federation (Kemerovo)	Kortshakolia ideliformis (124597)
Kungurian	283.3 - 274.4	Russian Federation (Suksun)	Sylvidelia latipennis (113379) Sylvidelia latipennis, Micaidelia minutissima, Sojanidelia florea (108206)
Kungurian	283.3 - 274.4	Russian Federation (Komi Republic)	Khosaridelia martynovi, Sojanidelia multimediana, Sojanidelia vorkutensis (151567)
Kungurian	283.3 - 274.4	Russian Federation	Ideliidae indet. (115214) Tshekardelia media (114207)
Kungurian	283.3 - 274.4	France	Paoliidae indet. (174413)
Roadian	274.4 - 266.9	Russian Federation	Idelia permiakovi (122871) Idelina kamensis (155230) Mezenalicula connata, Stenaropodites sojanensis, Sojanidelia nana, Sojanidelia makarkini, Sojanidelia fluctuosa, Sojanidelia striata, Permotermopsis pectinata (96860) Stenaropodites reticulata (116456)
Roadian	274.4 - 266.9	Russian Federation (Kemerovo)	Archidelia elongata, Archidelia ovata, Aenigmidelia incredibilis (104704) Archidelia ovata (122569 122571) Metidelia kostinae (122581)
Roadian	274.4 - 266.9	Russian Federation (Arkangelsk)	Permotermopsis roseni, Permotermopsis pectinata (116874) Protoblattinidae indet., Stenaropodites sojanensis, Stenaropodites mistshenkoi, Sojanidelia maculosa, Sojanidelia fasciata, Sojanidelia parvula, Sojanidelia nana, Sojanidelia makarkini, Sojanidelia fluctuosa, Sojanidelia lineata, Permotermopsis roseni, Permotermopsis pectinata (104992) Sojanidelia maculosa, Sojanidelia nana (104993) Sojanidelia striata (116933) Stenaropodites sojanensis, Stenaropodites mistshenkoi, Sojanidelia longula, Sojanidelia parvula, Sojanidelia makarkini, Sojanidelia fluctuosa, Sojanidelia lineata, Permotermopsis roseni, Permotermopsis pectinata (104716)
Roadian	274.4 - 266.9	Russian Federation (Kirov)	Stenaropodites magna, Idelina mala (105058)
Capitanian	264.28 - 259.51	Russian Federation (Orenburg)	Metidelia kargalensis (116607)
Wuchiapingian	259.51 - 254.14	Mongolia	Mongoloidelia gorochovi (97602)
Changhsingian	254.14 - 251.902	South Africa (Natal)	Paolekia perditae (124428)
Ladinian	241.464 - 237.0	Kyrgyzstan (Osh)	Ideliopsina nana, Ideliopsina ornata (106088) Ideliopsina nana, Ideliopsina ruginosa, Ideliopsina stupenda, Ideliopsina ornata (110763) Ideliopsina nana, Ideliopsina stupenda, Ideliopsina ornata (105007)
Carnian	237.0 - 227.3	Japan	Ideliopsina aristovi (230243)
Norian	227.3 - 205.7	Kazakhstan	Ideliopsina kenderlykensis (105055)