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Crocodylomorpha (crocodilian)
Taxonomy
Crocodilomorphi was named by Hay (1930). It is extant. It was considered paraphyletic by Georgi and Krause (2010), monophyletic by Hunt and Lockley (1995) and Turner and Sertich (2010) and Nesbitt (2011).
It was synonymized subjectively with Crocodylia by Haubold (1969); it was corrected as Crocodylomorpha by Walker (1970); it was reranked as the unranked clade Crocodilomorphi by Karl (2007); it was reranked as the unranked clade Crocodylomorpha by Gauthier and Padian (1985), Gauthier et al. (1989), Sereno and Arcucci (1990), Sereno (1991), Parrish (1993), Long and Murry (1995), Hunt and Lockley (1995), Gohlich et al. (2005), Weinbaum and Hungerbühler (2007), Brochu et al. (2009), Turner and Sertich (2010), Georgi and Krause (2010), Nesbitt (2011), Leardi et al. (2017) and Agnolín et al. (2023).
It was assigned to Loricata by Hay (1930); to Archosauria by Walker (1970); to Pseudosuchia by Gauthier and Padian (1985); to Paracrocodylomorpha by Parrish (1993); to Reptilia by Hunt and Lockley (1995); to Paracrocodyliformes by Weinbaum and Hungerbühler (2007); to Suchia by Sereno and Arcucci (1990), Sereno (1991), Long and Murry (1995) and Karl (2007); to Pseudosuchia by Gauthier et al. (1989) and Brochu et al. (2009); to Loricata by Nesbitt (2011); and to Archosauria by Gohlich et al. (2005), Turner and Sertich (2010), Georgi and Krause (2010), Leardi et al. (2017) and Agnolín et al. (2023).
It was synonymized subjectively with Crocodylia by Haubold (1969); it was corrected as Crocodylomorpha by Walker (1970); it was reranked as the unranked clade Crocodilomorphi by Karl (2007); it was reranked as the unranked clade Crocodylomorpha by Gauthier and Padian (1985), Gauthier et al. (1989), Sereno and Arcucci (1990), Sereno (1991), Parrish (1993), Long and Murry (1995), Hunt and Lockley (1995), Gohlich et al. (2005), Weinbaum and Hungerbühler (2007), Brochu et al. (2009), Turner and Sertich (2010), Georgi and Krause (2010), Nesbitt (2011), Leardi et al. (2017) and Agnolín et al. (2023).
It was assigned to Loricata by Hay (1930); to Archosauria by Walker (1970); to Pseudosuchia by Gauthier and Padian (1985); to Paracrocodylomorpha by Parrish (1993); to Reptilia by Hunt and Lockley (1995); to Paracrocodyliformes by Weinbaum and Hungerbühler (2007); to Suchia by Sereno and Arcucci (1990), Sereno (1991), Long and Murry (1995) and Karl (2007); to Pseudosuchia by Gauthier et al. (1989) and Brochu et al. (2009); to Loricata by Nesbitt (2011); and to Archosauria by Gohlich et al. (2005), Turner and Sertich (2010), Georgi and Krause (2010), Leardi et al. (2017) and Agnolín et al. (2023).
Subtaxa
Carnufex, Crocodilus humilis, Crocodilus kutchensis, Crocodilus proavus, Crocodilus vetustus, Dibothrosuchus, Diplosaurus nanus, Dromicosuchus, Eopneumatosuchus, Erythrochampsa, Erythrochampsa longipes, Gavialiformes, Gondwanasuchia, Goniopholidiformes, Goniopholis minor, Hallopodidae, Hallopus, Hesperosuchus, Kayentasuchus, Litargosuchus, Lukousaurus, Mycomorphoolithus, Paracrocodylia, Paratetrasauropus, Pedeticosaurus, Phyllodontosuchus, Pseudhesperosuchus, Redondavenator, Saltoposuchus, Solidocrania, Sphenosuchia, Sphenosuchidae, Sphenosuchus, Stegomosuchus, Stegomus longipes, Terrestrisuchus, Thalattosuchia (syn. Mesosauria), Thoracosaurus bahiensis, Trialestes (syn. Triassolestes)
Synonyms
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Synonymy list
Year | Name and author |
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1856 | Crocodilus humilis Leidy p. 73 |
1857 | Crocodilus humilis Leidy p. 90 |
1860 | Crocodilus humilis Leidy p. 146 |
1869 | Crocodilus vetustus Gervais p. 215 |
1869 | Thoracosaurus bahiensis Marsh |
1869 | Crocodilus vetustus Matheron p. 371 |
1881 | Crocodilus proavus Seeley p. 685 |
1887 | Goniopholis minor Koken p. 24 |
1895 | Diplosaurus nanus Marsh p. 405 |
1897 | Diplosaurus nanus Marsh p. 485 |
1897 | Crocodilus humilus Marsh p. 527 |
1902 | Crocodylus humilis Hay p. 513 |
1902 | Crocodilus humilis Lambe p. 47 |
1904 | Stegomus longipes Emerson and Loomis |
1915 | Stegomus longipes Lull p. 102 |
1917 | Stegomus longipes Lull p. 110 |
1922 | Stegomosuchus Huene pp. 22-23 |
1922 | Stegomosuchus longipes Huene pp. 22-23 |
1924 | Erythrochampsa Haughton |
1924 | Erythrochampsa longipes Haughton p. 367 |
1928 | Erythrochampsa Nopcsa p. 186 |
1930 | Stegomosuchus Hay p. 139 |
1930 | Stegomosuchus longipes Hay p. 139 |
1930 | Crocodilomorphi Hay p. 145 |
1930 | Goniopholis nanus Hay p. 146 |
1930 | Crocodylus humilis Hay p. 153 |
1942 | Stegomosuchus Huene p. 244 |
1970 | Erythrochampsa Ellenberger p. 347 |
1970 | Crocodylomorpha Walker p. 368 |
1970 | Erythrochampsa Walker p. 369 |
1970 | Stegomosuchus Walker p. 369 |
1971 | Crocodilus kutchensis Tandon |
1985 | Crocodylomorpha Gauthier and Padian pp. 187-189 fig. 1 |
1988 | Erythrochampsa Carroll |
1988 | Sphenosuchia Carroll |
1988 | Stegomosuchus Carroll |
1989 | Crocodylomorpha Gauthier et al. p. 348 |
1990 | Crocodylomorpha Sereno and Arcucci p. 41 |
1991 | Crocodylomorpha Sereno p. 11 |
1993 | Crocodylomorpha Parrish p. 290 fig. 1 |
1995 | Crocodylomorpha Hunt and Lockley |
1995 | Crocodylomorpha Long and Murry p. 164 |
1995 | Sphenosuchia Long and Murry p. 164 |
2005 | Crocodylomorpha Gohlich et al. |
2007 | Crocodilomorphi Karl p. 59 |
2007 | Crocodylomorpha Weinbaum and Hungerbühler p. 141 |
2009 | Crocodylomorpha Brochu et al. p. 538 |
2010 | Crocodylomorpha Georgi and Krause |
2010 | Crocodylomorpha Turner and Sertich |
2011 | Crocodylomorpha Nesbitt p. 203 fig. 51 |
2017 | Crocodylomorpha Leardi et al. p. 3 |
2023 | Crocodylomorpha Agnolín et al. |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Unr. Crocodylomorpha Hay 1930 [crocodilian]
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G. †Dibothrosuchus Simmons 1965
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Unr. †Gondwanasuchia Carvalho et al. 2004
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Unr. †Notosuchimorpha Carvalho et al. 2004
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Unr. †Notosuchiformes Carvalho et al. 2004
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Invalid names: Terriasuchia Carvalho et al. 2004 [empty]
Invalid names: Chimaerasuchidae Carvalho et al. 2004 [empty]
G. †Mycomorphoolithus Moreno-Azanza et al. 2015
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†Mycomorphoolithus kohringi Moreno-Azanza et al. 2015
G. †Saltoposuchus von Huene 1921
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†Saltoposuchus connectens Huene 1921
†Saltoposuchus longipes Huene 1921
Unr. †Solidocrania Ruebenstahl et al. 2022
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Subor. Crocodyliformes Hay 1930
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Fm. Krokolithidae Kohring and Hirsch 1996 [crocodile eggshell]
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G. †Krokolithes Hirsch 1985
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†Krokolithes helleri Kohring and Hirsch 1996
†Krokolithes wilsoni Hirsch 1985
Unr. †Mesoeucrocodylia Whetstone and Whybrow 1983
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Unr. Neosuchia Clark 1988
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Unr. †Atoposauridae Gervais 1871
G. †Bernissartia Dollo 1883
Fm. †Bernissartiidae Dollo 1883
G. †Burkesuchus Novas et al. 2021
Unr. †Coelognathosuchia Martin et al. 2013
Subor. Eusuchia Huxley 1875
G. †Gilchristosuchus Wu and Brinkman 1993
G. †Khoratosuchus Lauprasert et al. 2009
G. †Montsecosuchus Buscalioni and Sanz 1988
Unr. †Paluxysuchidae Drumheller et al. 2021
Fm. †Stomatosuchidae Stromer 1925
G. †Wahasuchus Saber et al. 2018
Invalid names: Alligatorium franconicum Ammon 1906 [nomen dubium], Brillanceausuchidae Michard et al. 1990 [empty], Goniopholis phuwiangensis Buffetaut and Ingavat 1983 [nomen dubium], Heterosuchus Seeley 1887 [nomen dubium], Ischyrochampsa Vasse 1995 [nomen dubium], Ischyrochampsa meridionalis Vasse 1995 [nomen dubium], Shamosuchus karakalpakensis Nesov et al. 1989 [nomen dubium], Shamosuchus occidentalis Efimov 1982 [nomen dubium], Susisuchidae Salisbury et al. 2003 [empty]
Infraor. †Notosuchia Gasparini 1971
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G. †Coringasuchus Kellner et al. 2009
G. †Eptalofosuchus Marinho et al. 2022
Unr. †Eunotosuchia Ruiz et al. 2021
G. †Llanosuchus Fiorelli et al. 2016
Unr. †Peirosaurimorpha Carvalho et al. 2004
Unr. †Ziphosuchia Ortega et al. 2000
Invalid names: Comahuesuchidae Bonaparte 1991 [empty], Libycosuchidae Stromer 1914 [empty], Trematochampsa Buffetaut 1974 [nomen dubium], Trematochampsa taqueti Buffetaut 1974 [nomen dubium]
Invalid names: Cunampaia Rusconi 1946 [nomen dubium], Cunampaia simplex Rusconi 1946 [nomen dubium], Cunampaiidae [invalid subgroup], Hsisosuchidae Young and Chow 1953 [empty], Mesosuchia Huxley 1875 [invalid subgroup]
G. †Neuquensuchus Fiorelli and Calvo 2007 [Neuquen crocodile]
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†Neuquensuchus universitas Fiorelli and Calvo 2007
Fm. †Protosuchidae Brown 1934
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G. †Protosuchus Brown 1934
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†Protosuchus haughtoni Busbey III and Gow 1984
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Invalid names: Lesothosuchus charigi Whetstone and Whybrow 1983 [synonym]
†Protosuchus micmac Sues et al. 1996
†Protosuchus richardsoni Brown 1933
Invalid names: Archaeosuchus Brown 1933 [replaced], Lesothosuchus Whetstone and Whybrow 1983 [synonym]
Unr. †Shartegosuchoidea Dollman et al. 2018
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G. †Shantungosuchus Young 1961
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†Shantungosuchus chuhsienensis Young 1961
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Invalid names: Shantungosuchus brachycephalus Young 1982 [synonym]
†Shantungosuchus hangjinensis Wu et al. 1994
Fm. †Shartegosuchidae Efimov 1988
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G. †Adzhosuchus Efimov et al. 2000
G. †Fruitachampsa Clark 2011
G. †Hsisosuchus Young and Chow 1953
G. †Kyasuchus Efimov and Leshchinskiy 2000
G. †Nominosuchus Efimov 1996
G. †Shartegosuchus Efimov 1988
G. †Sichuanosuchus Peng 1995
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†Sichuanosuchus huidongensis Peng 1995
†Sichuanosuchus shuhanensis Wu et al. 1997
Invalid names: Crocodilus blavieri Gray 1831 [nomen dubium], Crocodilus provencialis Giebel 1847 [nomen dubium], Kemkemia Cau and Maganuco 2009 [nomen dubium], Kemkemia auditorei Cau and Maganuco 2009 [nomen dubium], Lisboasaurus Seiffert 1975 [nomen dubium], Lisboasaurus estesi Seiffert 1975 [nomen dubium], Metasuchia Benton and Clark 1988 [empty], Microchampsa Young 1951 [nomen dubium], Microchampsa scutata Young 1951 [nomen dubium]
Infraor. †Thalattosuchia Fraas 1901 [marine crocodile]
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G. †Mystriosaurus Munster 1834
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†Mystriosaurus laurillardi Kaup 1837
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Invalid names: Steneosaurus brevior Tate and Blake 1876 [synonym]
Invalid names: Engyommasaurus Kaup 1834 [synonym], Macrospondylus Meyer 1830 [synonym]
Unr. †Neothalattosuchia Young et al. 2024
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Superfm. †Teleosauroidea Saint-Hilaire 1831
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G. †Charitomenosuchus Johnson et al. 2020
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†Charitomenosuchus leedsi Andrews 1909
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Invalid names: Stenosaurus teleosauroides Auer 1909 [synonym]
G. †Plagiophthalmosuchus Johnson et al. 2020
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†Plagiophthalmosuchus gracilirostris Westphal 1961
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Invalid names: Teleosaurus chapmanni Buckland 1836 [synonym]
Fm. †Teleosauridae Geoffroy 1831
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Tr. †Aeolodontini Johnson et al. 2022
G. †Crocodilaemus Jourdan 1857
G. †Haematosaurus Sauvage 1874
G. †Indosinosuchus Martin et al. 2018
G. †Steneosaurus Saint-Hilaire 1825
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Invalid names: Glaphyrorhynchus Meyer 1842 [synonym], Leptocranius Bronn 1837 [synonym]
Subfm. †Teleosaurinae Saint-Hilaire 1831
Invalid names: Aeolodontinae Johnson et al. 2020 [empty], Megalosaurus mersensis Lapparent 1955 [nomen dubium]
G. †Pelagosaurus Bronn 1841
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†Pelagosaurus typus Bronn 1841
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Invalid names: Crocodilus temporalis de Blainville 1835 [synonym], Mosellaesaurus rostrominor Monard 1846 [synonym]
Invalid names: Mosellaesaurus Monard 1846 [synonym]
G. †Teleidosaurus Eudes-Deslongchamps 1869
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†Teleidosaurus calvadosii Eudes-Deslongchamps 1866
†Teleidosaurus joberti Eudes-Deslongchamps 1867
Invalid names: Teleidosaurus bathonicus Mercier 1933 [nomen dubium]
Invalid names: Mesosauria Giebel 1847 [synonym], Steneosaurus gerthi Huene 1927 [nomen dubium]
G. †Trialestes Bonaparte 1982
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†Trialestes romeri Reig 1963
Invalid names: Triassolestes Reig 1963 [replaced]
Invalid names: Crocodilus humilis Leidy 1856 [nomen dubium], Crocodilus kutchensis Tandon 1971 [nomen dubium], Crocodilus proavus Seeley 1881 [nomen dubium], Crocodilus vetustus Matheron 1869 [nomen dubium], Diplosaurus nanus Marsh 1895 [nomen dubium], Erythrochampsa Haughton 1924 [nomen dubium], Erythrochampsa longipes Haughton 1924 [nomen dubium], Gavialiformes Hay 1930 [empty], Goniopholidiformes Hay 1930 [empty], Goniopholis minor Koken 1887 [nomen dubium], Sphenosuchia [invalid subgroup], Sphenosuchidae Huene 1922 [empty], Stegomosuchus Huene 1922 [nomen dubium], Stegomus longipes Emerson and Loomis 1904 [nomen dubium], Thoracosaurus bahiensis Marsh 1869 [nomen dubium]
Diagnosis
Reference | Diagnosis | |
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S. J. Nesbitt 2011 | The most inclusive clade containing Crocodylus niloticus Laurenti, 1768, but not Rauisuchus tiradentes Huene, 1942, Poposaurus gracilis Mehl, 1915, Gracilisuchus stipanicicorum Romer, 1972c, Prestosuchus chiniquensis Huene, 1942, or Aetosaurus ferratus Fraas, 1877 |
Measurements
No measurements are available
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Source: subo = suborder, subc = subclass, c = class, subp = subphylum | |||||
References: Hendy et al. 2009, Benton 1983, Carroll 1988 |
Age range
Maximum range based only on fossils: base of the Tuvalian to the top of the Holocene or 232.00000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 228.0 Ma
Minimum age of oldest fossil (stem group age): 228.0 Ma
Collections (3943 total)
Oldest occurrences | |||
Time interval | Ma | Country or state | Original ID and collection number |
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Carnian | Slovenia | Walteria sp. (66930) | |
Carnian | United Kingdom (Wales) | Crocodilia indet. (27012) | |
Carnian | USA (North Carolina) | Carnufex carolinensis (169982) | |
Carnian | Lesotho (Maseru) | Dijaquesopus obliquus (206549) | |
Carnian | United Kingdom (England) | Saltoposuchus sp. (13173 13174 13182) Terrestrisuchus sp. (25298 46889) Thecodontia indet. (13175 13176 13183) | |
Carnian | Argentina (San Juan) | Crocodylomorpha indet. (222095) Crocodylomorpha n. sp. (131436) Triassolestes romeri (52747) | |
Carnian - Norian | Thailand (Chaiyaphum) | Crocodyliformes indet. (219817) | |
Carnian - Norian | USA (North Carolina) | Dromicosuchus grallator, Crocodylomorpha indet. (32343) Protosuchia indet., Sphenosuchia indet. (89149) | |
Carnian - Norian | USA (Texas) | Crocodylomorpha indet. (22706) | |
Late/Upper Triassic - Early/Lower Jurassic | USA (Utah) | Sphenosuchia indet. A, Sphenosuchia indet. B (183201) | |
Late/Upper Triassic - Early/Lower Jurassic | Ethiopia | Mesoeucrocodylia indet. (48830) | |
Tuvalian | Argentina (La Rioja) | Trialestes romeri (26452) | |
Tuvalian | Italy (Trentino-Alto Adigo) | Crocodylomorpha indet. (205945) | |
Norian | Argentina (San Juan) | Crocodylomorpha indet. (182613 182614) | |
Norian | Switzerland (Aargau) | Saltoposuchus sp. (22680) | |
Norian | USA (Arizona) | Crocodylomorpha indet. (88329) Hesperosuchus agilis (39219) Hesperosuchus agilis, Sphenosuchia indet. (39160) Hesperosuchus sp. (57685 92850) Reptilia indet. (93551) Sphenosuchia indet. (22705 48547 93361) Sphenosuchidae indet. (48423 84017) Sphenosuchus sp. (49500) | |
Norian | USA (Texas) | Sphenosuchidae indet. (13273) | |
Norian | USA (Utah) | Crocodylomorpha indet. (92808) | |
Norian | Argentina (La Rioja) | Coloradisuchus abelini (200293) Hemiprotosuchus leali, Pseudhesperosuchus jachaleri, Crocodylomorpha n. sp. (105691) | |
Norian | USA (New Mexico) | Hesperosuchus sp. (83738) Redondavenator quayensis, Sphenosuchia indet. (89969) Sphenosuchia indet. (188759) Sphenosuchidae indet., Sphenosuchia indet. (48516) |