PBDB Taxon

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Protorthoptera

Insecta - Protorthoptera

Taxonomy

Protorthoptera was named by Handlirsch (1906). It is not extant.

It was assigned to Pterygogenea by Handlirsch (1906); to Insecta by Zimmerman (1962); to Orthopteroidea by Martynov (1925), Sharov (1968); to Neoptera by Martynov (1937), Carpenter (1992); and to Polyneoptera by Eades (2012).

Subtaxa

Acridites, Adeloneuridae, Agogoblattina, Anthracomastax, Anthracothremmidae, Apithanidae, Appertiidae, Archaeologus, Archimastax, Asiopompidae, Asiopompus, Atava, Atava ovata, Boutakovia, Cacurgellus, Chrestotella, Chrestotes, Commentrya (syn. Oedischiopsis), Cymbopsidae, Danielsiella, Distasis, Eucaenidae (syn. Teneopteridae), Eucaenus pusillus, Eucaenus rotundatus, Gerarianus, Gerarites, Geraroides, Hadentomidae (syn. Palaeocixiidae), Hadentomoides, Hemeristia, Herdinidae, Homalophlebiidae, Homoeodictyidae, Hyaloptilus, Kelleropteron, Lecopterum, Limburgina, Macrophlebium, Megagnathidae, Megalometer, Melanoblattula, Metacheliphlebia, Notoblattites, Ochetopteron, Orthoneurites, Pachytylopsidae, Palaeocarria, Palaeoedischia, Polyernus, Polyetes, Protoblattinopsidae, Prototettigidae, Prototettix (syn. Lithophasma), Pruvostia, Pseudetoblattina, Pseudogerarus, Pseudopolyernus, Ptenodera, Rigattopteridae, Roomeria, Roomeriidae, Schuchertiella, Schuchertiellidae, Sharovia, Sheimiidae (syn. Visheriferinae), Soyaniidae, Stenaropodidae, Stephanotermopsidae, Thoronysididae

Synonyms

Protoperlaria was named by Tillyard (1928).

It was synonymized subjectively with Protorthoptera by Carpenter (1992).

It was assigned to Neoptera by Martynov (1937); and to Insecta by Bode (1953).
Schuchertiellidae was named by Handlirsch (1911). Its type is Schuchertiella.

It was considered an invalid subgroup of Protorthoptera by Carpenter (1992).

It was assigned to Protorthoptera by Handlirsch (1911), Handlirsch (1922).
Eucaenus pusillus was named by Handlirsch (1911). Its type specimen is YPM 52, an exoskeleton, and it is an impression. Its type locality is Mazon Creek (YPM collection), which is in a Westphalian D deltaic mudstone in the Carbondale Formation of Illinois.

It was considered a nomen dubium by Carpenter and Richardson (1976).
Eucaenus rotundatus was named by Handlirsch (1906). Its type specimen is USNM 38153, a forewing, and it is an impression. Its type locality is Mazon Creek, USNM Lacoe Collection, which is in a Westphalian D interdistributary bay mudstone in the Carbondale Formation of Illinois.

It was considered a nomen dubium by Carpenter and Richardson (1976).
Atava was named by Sellards (1909). Its type is Atava ovata.

It was considered a nomen dubium by Carpenter (1950).

It was assigned to Lepiidae by Handlirsch (1919), Handlirsch (1922); and to Protorthoptera by Carpenter (1992).
Atava ovata was named by Sellards (1909). Its type specimen is Sellards 372, a hindwing, and it is an impression. Its type locality is Elmo, Sellards collection, which is in an Artinskian pond limestone in the Wellington Formation of Kansas. It is the type species of Atava.

It was considered a nomen dubium by Carpenter (1950).
Prototettigidae was named by Handlirsch (1906). Its type is Prototettix.

It was considered an invalid subgroup of Protorthoptera by Carpenter (1992).

It was assigned to Protorthoptera by Handlirsch (1906), Handlirsch (1919), Handlirsch (1922), Guthörl (1934).
Asiopompidae was named by Sharov (1961).

It was considered an invalid subgroup of Protorthoptera by Carpenter (1992).

It was assigned to Protoblattodea by Sharov (1961).
Hadentomoidea was named by Handlirsch (1906).

It was synonymized subjectively with Protorthoptera by Carpenter (1992).

It was assigned to Polyneoptera by Handlirsch (1906).
Roomeriidae was named by Handlirsch (1919).

It was considered an invalid subgroup of Protorthoptera by Carpenter (1992).

It was assigned to Protorthoptera by Handlirsch (1919).

Synonymy list

Year	Name and author
1906	Protorthoptera Handlirsch p. 128
1906	Prototettigidae Handlirsch p. 135
1906	Eucaenus rotundatus Handlirsch pp. 165-166 figs. Pl XVI fig 22
1906	Hadentomoidea Handlirsch p. 302
1906	Eucaenus rotundatus Handlirsch p. 710 fig. 36
1909	Atava Sellards p. 157
1909	Atava ovata Sellards p. 157 fig. 3
1911	Eucaenus rotundatus Handlirsch
1911	Schuchertiellidae Handlirsch
1911	Eucaenus pusillus Handlirsch p. 362 figs. Fig 43
1919	Prototettigidae Handlirsch p. 541
1919	Atava Handlirsch p. 542
1919	Roomeriidae Handlirsch p. 553
1922	Prototettigidae Handlirsch p. 76
1922	Atava Handlirsch p. 77
1922	Atava ovata Handlirsch p. 77
1922	Schuchertiellidae Handlirsch p. 79
1922	Eucaenus pusillus Handlirsch p. 105
1922	Eucaenus rotundatus Handlirsch p. 105
1925	Protorthoptera Martynov p. 576
1928	Protoperlaria Tillyard
1934	Prototettigidae Guthörl p. 96
1937	Protorthoptera Martynov p. 23
1937	Protoperlaria Martynov p. 29
1953	Protoperlaria Bode
1961	Asiopompidae Sharov p. 160
1962	Protorthoptera Zimmerman p. 1325
1968	Protorthoptera Sharov
1992	Protorthoptera Carpenter p. 97
1992	Atava Carpenter p. 132
2012	Protorthoptera Eades

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Ecdysozoa	Aguinaldo et al. 1997
—	Panarthropoda
phylum	Arthropoda	Latreille 1829
—	Deuteropoda
—	Mandibulata
—	Pancrustacea	Zrzavý and Štys 1997
—	Altocrustacea	Regier et al. 2010
superclass	Allotriocarida
—	Athalassocarida

Rank	Name	Author
—	Labiocarida	Schwentner et al. 2017
superclass	Hexapoda	Latreille 1825
—	Cercofilata
class	Insecta
—	Dicondylia
—	Paranotalia
subclass	Pterygota	()
—	Neopterygota	Crampton 1924
infraclass	Neoptera	Martynov 1923
—	Polyneoptera	Martynov 1938
order	Protorthoptera
order	Protorthoptera

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Or. †Protorthoptera Handlirsch 1906

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G. †Acridites Germar 1842

†Lithosialis carbonaria Germar 1842

Fm. †Adeloneuridae Carpenter 1938

G. †Adeloneura Carpenter 1938

†Adeloneura thompsoni Carpenter 1938

G. †Agogoblattina Handlirsch 1906

†Agogoblattina occidua Scudder 1885

G. †Anthracomastax Handlirsch 1904

Fm. †Anthracothremmidae Handlirsch 1906

G. †Anthracothremma Scudder 1885

†Anthracothremma robusta Scudder 1885

G. †Melinophlebia Handlirsch 1911

†Melinophlebia analis Handlirsch 1911

G. †Pericalyphe Handlirsch 1911

†Pericalyphe longa Handlirsch 1911

G. †Silphion Handlirsch 1911

†Silphion latipenne Handlirsch 1911

Fm. †Apithanidae Handlirsch 1911

G. †Apithanus Handlirsch 1911

†Apithanus jocularis Handlirsch 1911

Fm. †Appertiidae Paulian 1965

G. †Appertia Paulian 1965

†Appertia besairiei Paulian 1965

G. †Archaeologus Handlirsch 1906

†Archaeologus falcatus Handlirsch 1906

G. †Archimastax Handlirsch 1906

†Archimastax americanus Handlirsch 1906

G. †Asiopompus Sharov 1961

†Asiopompus tomicus Sharov 1961

G. †Boutakovia Pruvost 1934

†Boutakovia saleei Pruvost 1934

G. †Cacurgellus Pruvost 1919

†Cacurgellus barryi Pruvost 1919

G. †Chrestotella Handlirsch 1919

†Chrestotella danae Scudder 1868

G. †Chrestotes Scudder 1868

†Chrestotes lapidea Scudder 1868

G. †Commentrya Lameere 1917

†Commentrya maximae Brongniart 1894

Invalid names: Oedischiopsis Handlirsch 1919 [objective synonym]

Fm. †Cymbopsidae Kukalová 1965

G. †Cymbopsis Kukalová 1965

†Cymbopsis excelsa Kukalová 1965

G. †Danielsiella Cockerell 1916

†Danielsiella priscula Cockerell 1916

G. †Distasis Handlirsch 1904

†Distasis rhipiphora Handlirsch 1904

Fm. †Eucaenidae Handlirsch 1906

G. †Eucaenus Scudder 1885

†Eucaenus ovalis Scudder 1885

Invalid names: Eucaenus attenuatus Melander 1903 [synonym], Eucaenus mazonus Melander 1903 [synonym], Eucaenus minor Handlirsch 1911 [synonym], Teneopteron mirabile Carpenter 1943 [synonym]

Invalid names: Teneopteron Carpenter 1943 [synonym]

Invalid names: Teneopteridae Richardson 1956 [synonym]

G. †Gerarianus Handlirsch 1919

†Gerarianus commentryanus Handlirsch 1919

G. †Gerarites Handlirsch 1919

†Gerarites commentryi Brongniart 1894

G. †Geraroides Handlirsch 1906

†Geraroides maximus Melander 1903

Fm. †Hadentomidae Handlirsch 1906

G. †Hadentomum Handlirsch 1906

†Hadentomum americanum Handlirsch 1906

Invalid names: Palaeocixiidae Handlirsch 1919 [synonym]

G. †Hadentomoides Riek 1974

†Hadentomoides dwykensis Riek 1974

G. †Hemeristia Dana 1864

†Hemeristia occidentalis Dana 1864

Fm. †Herdinidae Carpenter and Richardson 1971

G. †Herdina Carpenter and Richardson 1971

†Herdina mirificus Carpenter and Richardson 1971

Fm. †Homalophlebiidae Handlirsch 1906

G. †Homalophlebia Brongniart 1894

†Homalophlebia finoti Brongniart 1894

G. †Parahomalophlebia Handlirsch 1906

†Parahomalophlebia courtini Brongniart 1894

Fm. †Homoeodictyidae Martynov 1937

G. †Homoeodictyon Martynov 1937

†Homoeodictyon elongatum Martynov 1937

G. †Hyaloptilus Lameere 1917

†Hyaloptilus minimus Lameere 1917

G. †Kelleropteron Brauckmann and Hahn 1980

†Kelleropteron kaelberbergense Brauckmann and Hahn 1980

G. †Lecopterum Sellards 1909

†Lecopterum delicosum Sellards 1909

G. †Limburgina Laurentiaux 1950

†Limburgina antiqua Laurentiaux 1950

G. †Macrophlebium Goldenberg 1869

†Macrophlebium hollebeni Goldenberg 1869

Fm. †Megagnathidae Tillyard 1918

G. †Megagnatha Bolton 1917

†Megagnatha odonatiformis Bolton 1917

G. †Megalometer Handlirsch 1906

†Megalometer lata Handlirsch 1906

G. †Melanoblattula Cockerell 1927

†Melanoblattula nigrescens Cockerell 1927

G. †Metacheliphlebia Handlirsch 1906

†Metacheliphlebia elongata Scudder 1885

G. †Notoblattites Tillyard 1916

†Notoblattites mitchelli Tillyard 1918

†Notoblattites subcostalis Tillyard 1916

†Notoblattites wianamattensis Tillyard 1918

G. †Ochetopteron Cockerell 1927

†Ochetopteron canaliculatum Cockerell 1927

G. †Orthoneurites Martynov 1928

†Orthoneurites regularis Martynov 1928

Fm. †Pachytylopsidae Handlirsch 1904

G. †Pachytylopsis de Borre 1875

†Pachytylopsis deriodoti Pruvost 1930

†Pachytylopsis melas Handlirsch 1904

†Pachytylopsis persenairei de Borre 1875

†Pachytylopsis stainieri Pruvost 1930

Invalid names: Palorthopteron Handlirsch 1904 [synonym]

G. †Protopachytylopsis Laurentiaux and Laurentiaux-Vieira 1981

†Protopachytylopsis leckwycki Laurentiaux and Laurentiaux-Vieira 1981

G. †Symballophlebia Handlirsch 1904

†Symballophlebia latipennis Handlirsch 1904

G. †Palaeocarria Cockerell 1917

†Palaeocarria ornata Cockerell 1917

G. †Palaeoedischia Meunier 1914

†Palaeoedischia boulei Meunier 1914

G. †Polyernus Scudder 1885

†Polyernus complanatus Scudder 1885

G. †Polyetes Handlirsch 1906

†Polyetes furcifer Handlirsch 1906

†Polyetes problematica Bolton 1934

G. †Prototettix Giebel 1856

†Prototettix lithanthraca Goldenberg 1854

Invalid names: Lithophasma Brongniart 1885 [objective synonym]

G. †Pruvostia Bolton 1921

†Pruvostia spectabilis Bolton 1921

G. †Pseudetoblattina Handlirsch 1906

†Pseudetoblattina reliqua Scudder 1893

G. †Pseudogerarus Handlirsch 1906

†Pseudogerarus scudderi Handlirsch 1906

G. †Pseudopolyernus Handlirsch 1906

†Pseudopolyernus laminarum Scudder 1885

G. †Ptenodera Bolton 1922

†Ptenodera dubius Bolton 1922

Fm. †Rigattopteridae Pinto 1996

G. †Rigattoptera Pinto 1996

†Rigattoptera ornellasae Pinto 1996

G. †Roomeria Meunier 1914

†Roomeria carbonaria Meunier 1914

G. †Schuchertiella Handlirsch 1911

†Schuchertiella gracilis Handlirsch 1911

G. †Sharovia Pinto and de Ornellas 1978

†Sharovia permiafricana Pinto and de Ornellas 1978

Fm. †Sheimiidae Martynova 1958

G. †Magniscapa Shcherbakov 2015

†Magniscapa alligans Aristov and Rasnitsyn 2008

G. †Parasheimia Aristov 2004

†Parasheimia kazanica Aristov and Rasnitsyn 2008

†Parasheimia rotundata Aristov 2004

†Parasheimia truncata Aristov 2004

G. †Pseudosheimia Aristov 2004

†Pseudosheimia caudata Aristov 2004

†Pseudosheimia caudissima Aristov and Rasnitsyn 2008

G. †Sheimia Martynova 1958

†Sheimia sojanensis Martynova 1958

†Sheimia tshekardensis Aristov 2004

G. †Soyanembia Aristov and Rasnitsyn 2008

†Soyanembia sharovi Aristov and Rasnitsyn 2008

G. †Visherifera Novokshonov et al. 2002

†Visherifera camura Novokshonov et al. 2002

†Visherifera sylvaensis Aristov 2004

Invalid names: Visheriferinae Novokshonov et al. 2002 [synonym]

Fm. †Soyaniidae Shcherbakov 2015

G. †Soyania Aristov and Rasnitsyn 2008

†Soyania logica Aristov and Rasnitsyn 2008

Fm. †Stephanotermopsidae Laurentiaux 1966

G. †Stephanotermopsis Laurentiaux 1966

†Stephanotermopsis rodendorfi Laurentiaux 1966

Fm. †Thoronysididae Handlirsch 1919

G. †Thoronysis Handlirsch 1906

†Thoronysis ingbertensis von Ammon 1903

Invalid names: Asiopompidae Sharov 1961 [invalid subgroup], Atava Sellards 1909 [nomen dubium], Atava ovata Sellards 1909 [nomen dubium], Eucaenus pusillus Handlirsch 1911 [nomen dubium], Eucaenus rotundatus Handlirsch 1906 [nomen dubium], Hadentomoidea Handlirsch 1906 [synonym], Protoblattinopsidae Laurentiaux 1966 [empty], Protoperlaria Tillyard 1928 [synonym], Prototettigidae Handlirsch 1906 [invalid subgroup], Roomeriidae Handlirsch 1919 [invalid subgroup], Schuchertiellidae Handlirsch 1911 [invalid subgroup], Stenaropodidae Handlirsch 1906 [empty]

Diagnosis

No diagnoses are available