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Phacopina

Trilobita - Phacopida

Taxonomy
Phacopina was named by Struve (1959) [in R.C. Moore 1959].

It was assigned to Phacopida by Reijers (1972), Sandford (2004), Adrain (2011), Randolfe and Gass (2024).

Synonymy list
YearName and author
1959Phacopina Struve pp. O461-O462
1972Phacopina Reijers
2004Phacopina Sandford
2011Phacopina Adrain p. 105
2024Phacopina Randolfe and Gass p. 822

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—EcdysozoaAguinaldo et al. 1997
—Panarthropoda
RankNameAuthor
phylumArthropodaLatreille 1829
subphylumArtiopoda(Hou and Bergstrom)
classTrilobitaWalch 1771
orderPhacopidaSalter 1864
suborderPhacopinaStruve 1959
suborderPhacopinaStruve 1959

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Subor. †Phacopina Struve 1959
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Superfm. †Acastoidea Reijers 1972
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Fm. †Acastidae Delo 1935
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G. †Acastava Richter 1954
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†Acastava lerougei van Viersen 2013
Subfm. †Acastavinae Struve 1958
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G. †Acastocephala Shergold 1966
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†Acastocephala macrops Salter 1864
G. †Acaste Goldfuss 1843
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†Acaste andersoni Edgecombe 2007
†Acaste downingiae Murchison 1839
Invalid names: Acastina Reed 1927 [objective synonym]
G. †Acastella Reed 1925
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†Acastella artesia Richter and Richter 1954
†Acastella granulosa Hollard 1963
†Acastella heberti Gosselet 1880
†Acastella heberti elsana Richter and Richter 1954
†Acastella hollardi Gandl 1972
†Acastella jacquemonti Hollard 1963
†Acastella patula Hollard 1963
†Acastella rouaulti Tromelin and Lebesconte 1876
†Acastella spinosa Salter 1864
†Acastella tanzidensis Hollard 1963
†Acastella tiro Richter and Richter 1954
G. †Acastellina Richter and Richter 1975
G. †Acastoides Delo 1935
+
Subg. †Acastoides (Acastoides) Delo 1935
†Acastoides haddadi Chatterton et al. 2006
†Acastoides hollardi Alberti 1966
†Acastoides maurus Alberti 1970
†Acastoides paeckelmanni Richter and Richter 1939
†Acastoides zguilmensis Chatterton et al. 2006
G. †Acastopyge Tomczykowa and Tomczyk 1970
G. †Armorigreenops Lieberman and Kloc 1997
Subfm. †Asteropyginae Delo 1935
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G. †Alcaldops Arbizu 1979
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†Alcaldops alcaldei Arbizu 1977
†Alcaldops camfroutensis Morzadec 1983
†Alcaldops renaudae Morzadec 1983
†Alcaldops troaonensis Morzadec 1983
G. †Asteropyge Hawle and Corda 1847
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Subg. †Asteropyge (Asteropyge) Hawle and Corda 1847
Subg. †Asteropyge (Comura) Richter 1926
†Asteropyge comes Basse 2003
†Asteropyge lauchensis Basse 2003
†Asteropyge perforata Morzadec 1969
†Asteropyge plana Basse 2003
†Asteropyge punctata Steininger 1831
G. †Bradocryphaeus Haas and Mensink 1970
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†Bradocryphaeus briceae Morzadec 1988
†Bradocryphaeus echinatus Van Viersen and Vanherle 2018
†Bradocryphaeus feisti Morzadec 1992
†Bradocryphaeus fiacrensis Morzadec 1976
†Bradocryphaeus kermanensis Morzadec et al. 2002
†Bradocryphaeus laomedeia Van Viersen and Vanherle 2018
†Bradocryphaeus maillieuxi Richter and Richter 1926
†Bradocryphaeus mosanus Richter and Richter 1926
†Bradocryphaeus neptuni Van Viersen and Bignon 2011
†Bradocryphaeus nodosa Morzadec 1983
†Bradocryphaeus vanherlei Van Viersen and Bignon 2011
†Bradocryphaeus zornensis Morzadec 1983
G. †Destombesina Morzadec 1997
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†Destombesina schumacherorum Basse and Mueller 2016
G. †Dunopyge Struve 1998
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†Dunopyge drevermanni Richter 1909
†Dunopyge lebanzaensis Smeenk 1983
G. †Gandlops Bignon and Crônier 2013
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†Gandlops brevispinosus Gandl 1972
†Gandlops ogivalis Morzadec 1976
†Gandlops tamnrherta Chatterton et al. 2006
G. †Ganetops Bignon et al. 2014
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†Ganetops djemelensis Morzadec 1997
†Ganetops ebbae Richter and Richter 1954
†Ganetops gerrinensis Timm 1978
†Ganetops lata Timm 1978
G. †Heliopyge Haas and Mensink 1970
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†Heliopyge helios Richter and Richter 1926
†Heliopyge sharudensis Pillet 1974
G. †Jaylopygiella Basse 2003
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†Jaylopygiella debruni van Viersen 2013
†Jaylopygiella medusa Basse 2003
G. †Metacanthina Pillet 1954
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†Metacanthina asnoensis Gandl 1972
†Metacanthina barrandei de Tromelin and Lebesconte 1876
†Metacanthina issoumourensis Morzadec 2001
G. †Minicryphaeus Bignon and Crônier 2014
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†Minicryphaeus giganteus Bignon et al. 2014
†Minicryphaeus minimus Morzadec 2001
†Minicryphaeus sarirus Morzadec 2001
G. †Morzadecops Bignon and Crônier 2014
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†Morzadecops matutina Gandl 1972
†Morzadecops zguidensis Morzadec 2001
G. †Pennarbedops Bignon and Crônier 2014
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†Pennarbedops babini Morzadec 1983
G. †Platykardiapyge Van Viersen and Kloc 2022
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†Platykardiapyge maderensis Morzadec 2001
†Platykardiapyge wallacei Termier and Termier 1950
G. †Quadratispina Gandl 1972
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†Quadratispina excelsa Van Viersen and Vanherle 2018
†Quadratispina hispanica Richter and Richter 1926
G. †Radiopyge Farsan 1981
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†Radiopyge hamedii Morzadec et al. 2002
†Radiopyge heratensis Farsan 1981
G. †Rhenops Richter and Richter 1943
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†Rhenops hammerschmidti Richter and Richter 1939
†Rhenops odremani Carvalho and Moody 2000
G. †Baniaspis Destombes 1972
G. †Bellacartwrightia Lieberman and Kloc 1997
G. †Braunops Lieberman and Kloc 1997
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†Braunops champagnensis Morzadec 1971
†Braunops djazairensis Morzadec 1997
†Braunops obsoletus Gandl 1972
G. †Breizhops Morzadec 1983
G. †Centauropyge Haas 1968
G. †Chimaerastella Gandl 1972
G. †Coltraneia Lieberman and Kloc 1997
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†Coltraneia celtica Morzadec 1969
†Coltraneia effelesa Chatterton et al. 2006
G. †Cryphina Oehlert 1889
G. †Delocare Struve 1958
G. †Deloops Lieberman and Kloc 1997
G. †Echinopyge Haas 1968
G. †Feruminops Haas 1986
G. †Gourdonia Pillet 1954
G. †Greenops Delo 1935
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Subg. †Greenops (Neometacanthus) Richter 1984
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†Greenops boothi Green 1837
†Greenops perijaensis Carvalho and Moody 2000
Invalid names: Cryphaeus Green 1837 [replaced]
G. †Gudralisium Basse 1998
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†Gudralisium viator Basse 1998
G. †Hallandclarkeops Lieberman and Kloc 1997
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†Hallandclarkeops cantarmoricus Morzadec and Arbizu 1978
†Hallandclarkeops renaudae Morzadec 1969
G. †Harringtonacaste Baldis and Blasco 1976
G. †Hexacosta Farsan 1981
G. †Hollardops Morzadec 1997
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†Hollardops aithassainorum Chatterton et al. 2006
†Hollardops mesocristata Le Maitre 1952
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Invalid names: Hollardops hyfinkeli Lieberman and Kloc 1997 [synonym]
Invalid names: Modellops Lieberman and Kloc 1997 [synonym], Philipsmithiana Lieberman and Kloc 1997 [synonym]
G. †Kayserops Delo 1935
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†Kayserops kochi Kayser 1883
G. †Kennacryphaeus Lieberman and Kloc 1997
G. †Kloucekia Delo 1935
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Subg. †Kloucekia (Kloucekia) Delo 1935
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†Kloucekia phillipsi Barrande 1846
Subg. †Kloucekia (Phacopidina) Bancroft 1949
†Kloucekia apiculata McCoy 1851
G. †Llandovacaste Edgecombe 1993
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G. †Mimocryphaeus Gandl 1972
G. †Morocconites Boucot and McClure 1989
G. †Neocalmonia Pillet 1969
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Subg. †Neocalmonia (Radiopyge) Farsan 1981
†Neocalmonia batillifera Haas and Mensink 1969
†Neocalmonia chitralensis Feist et al. 2001
†Neocalmonia lutensis Morzadec 2002
†Neocalmonia quadricosta Pillet 1969
†Neocalmonia yazdii Morzadec 2002
G. †Paracryphaeus Gandl 1972
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†Paracryphaeus jonesi Oehlert 1877
G. †Pelitlina Haas 1968
G. †Phacopidina Bancroft and Lamont 1949
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†Phacopidina harnagensis Bancroft 1949
†Phacopidina micheli Tromelin 1877
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†Phacopidina micheli micheli Tromelin 1877
G. †Pilletina Haas 1970
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†Pilletina oehlerti Pillet 1953
†Pilletina primitiae Gandl 1972
G. †Protacanthina Gandl 1972
G. †Pseudocryphaeus Pillet 1954
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†Pseudocryphaeus michelini Rouault 1851
†Pseudocryphaeus munieri Oehlert 1877
G. †Psychopyge Termier and Termier 1950
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†Psychopyge elegans Termier and Termier 1950
†Psychopyge hammerorum Chatterton et al. 2006
†Psychopyge praestans Morzadec 2001
G. †Rheicops Basse 1998
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G. †Sanidopyge Haas 1968
G. †Scotiella Delo 1935
G. †Sokhretia Hupe 1956
G. †Stummiana Lieberman and Kloc 1997
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†Stummiana osismorum Morzadec 1983
G. †Tolkienia Lieberman and Kloc 1997
G. †Treveropyge Struve 1958
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†Treveropyge acrifrons Gandl 1972
†Treveropyge iberica Gandl 1972
†Treveropyge lips Richter and Richter 1943
†Treveropyge prorotundifrons Richter and Richter 1943
†Treveropyge wiltziana Basse 2003
G. †Turcopyge Richter 1939
G. †Walliserops Morzadec 2001
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†Walliserops hammii Chatterton et al. 2006
†Walliserops lindoei Chatterton and Gibb 2010
†Walliserops tridens Chatterton et al. 2006
†Walliserops trifurcatus Morzadec 2001
Fm. †Calmoniidae Delo 1935
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G. †Anchiopella Reed 1907
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Invalid names: Bainella Rennie 1930 [synonym]
G. †Andinacaste Eldredge and Ormiston 1979
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†Andinacaste talebensis Hollard 1963
G. †Australoacaste Benedetto and Martel 1976
G. †Australops Baldis 1972
G. †Bouleia Kozlowski 1923
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Invalid names: Dereimsia Kozlowski 1923 [synonym]
G. †Calmonia Clarke 1913
G. †Chiarumanipyge Branisa and Vanek 1973
G. †Clarkeaspis Lieberman 1993
G. †Colombianaspis Morzadec et al. 2015
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†Colombianaspis carvalhoae Morzadec et al. 2015
G. †Cryphaeoides Delo 1935
G. †Deltacephalaspis Eldredge and Ormiston 1979
G. †Eldredgeia Lieberman 1993
G. †Hadrorachus Delo 1935
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†Hadrorachus australis Clarke 1913
Invalid names: Hadrorakos Delo 1940 [objective synonym]
G. †Jujuyops Baldis and Blasco 1976
G. †Kozlowskiaspis Branisa and Vanek 1973
G. †Malvinella Wolart 1968
G. †Malvinocooperella Lieberman 1993
G. †Metacryphaeus Reed 1907
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†Metacryphaeus kegeli Carvalho et al. 1997
†Metacryphaeus meloi Carvalho et al. 1997
†Metacryphaeus tuberculatus Kozlowski 1923
G. †Oosthuizenella Cooper 1982
G. †Palpebrops Lieberman and Edgecombe 1991
G. †Parabouleia Eldredge 1972
G. †Paracalmonia Struve 1958
G. †Pennaia Clarke 1913
G. †Phacopina Clarke 1913
G. †Plesioconvexa Lieberman 1993
G. †Plesiomalvinella Lieberman and Edgecombe 1991
G. †Probolops Delo 1935
G. †Punillaspis Baldis and Longobucco 1979
G. †Renniella Cooper 1982
G. †Schizostylus Delo 1935
G. †Tarijactinoides Suárez-Soruco 1971
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Invalid names: Bolivianaspis Branisa and Vanek 1973 [synonym]
G. †Tibagya Struve 1958
G. †Typhloniscus Salter 1856
G. †Vogesina Wolfart 1968
G. †Wolfartaspis Cooper 1982
Superfm. †Dalmanitoidea Vogdes 1890
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Fm. †Dalmanitidae Vogdes 1890
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G. †Anchiopsis Delo 1935
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†Anchiopsis anchiops Green 1832
G. †Andreaspis Struve 1962
G. †Argentopyge Baldis 1972
G. †Bessazoon Curtis and Lane 1998
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†Bessazoon gibbae Chatterton and Ludvigsen 2004
G. †Blanodalmanites Kobayashi and Hamada 1972
G. †Chattiaspis Struve 1958
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†Chattiaspis almadenensis Hammann 1972
G. †Coronura Hall and Clarke 1888
G. †Corycephalus Hall and Clarke 1888
G. †Crozonaspis Henry 1968
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†Crozonaspis armata Hammann 1972
†Crozonaspis chauveli Henry 1980
†Crozonaspis dujardini Rouault 1847
†Crozonaspis incerta Deslongchamps 1825
†Crozonaspis kerfornei Clarkson and Henry 1969
†Crozonaspis mayensis Clarkson and Henry 1972
†Crozonaspis morenensis Hammann 1972
†Crozonaspis morenensis mayensis Clarkson and Henry 1973
†Crozonaspis struvei Henry 1968
G. †Dalmanites Barrande 1852
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Subg. †Dalmanites (Synphoria) Clarke 1900
†Dalmanites caudatus Brünnich 1781
†Dalmanites jupiterensis Twenhofel 1928
†Dalmanites meeki Walcott 1884
†Dalmanites oriens Barrande 1852
†Dalmanites socialis Barrande 1852
†Dalmanites townsendae Raymond 1931
Invalid names: Hausmannia Etheridge and Mitchell 1895 [synonym], Heliocephalus Delo 1935 [synonym], Ommokris Holloway 1981 [synonym]
G. †Dalmanitina Reed 1905
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Subg. †Dalmanitina (Eodalmanitina) Henry 1965
Subg. †Dalmanitina (Struveria) Rickards 1964
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†Struveria simrica Hede 1915
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Invalid names: Struveria howgillensis Rickards 1965 [synonym]
†Dalmanitina socialis Barrande 1846
Subfm. †Dalmanitinae Vodges 1890
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G. †Waukeshaaspis Randolfe and Gass 2024
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†Waukeshaaspis eatonae Randolfe and Gass 2024
G. †Dalmaniturus Chernysheva 1937
G. †Daytonia Holloway 1981
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†Daytonia jollieti Chatterton and Ludvigsen 2004
G. †Delops Rickards 1965
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†Delops obtusicaudatus Salter 1849
G. †Dreyfussina Hupé 1956
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†Dreyfussina exophtalma Dreyfuss 1948
†Dreyfussina struvei Destombes 1963
G. †Duftonia Dean 1959
G. †Fenestraspis Branisa and Vanek 1973
G. †Forillonaria Lesperance 1975
G. †Francovichia Branisa 1960
G. †Furacopyge Aribazu 1978
G. †Gamonedaspis Branisa and Vanek 1973
G. †Glyptambon Holloway 1981
G. †Huntoniatonia Campbell
+
Invalid names: Huntonia Vandel 1973 [replaced]
G. †Kazachstania Maksimova 1972
G. †Lygdozoon Holloway 1981
G. †Malladaia Oehlert 1896
G. †Morgatia Hammann 1972
+
†Morgatia hupei Nion and Henry 1966
†Morgatia primitiva Hammann 1972
G. †Mucronaspis Destombes 1963
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Subg. †Mucronaspis (Songxites) Lin 1981
+
+
Invalid names: Mucronaspis malayensis Kobayashi and Hamada 1964 [synonym]
†Mucronaspis chillonensis Hammann 1972
†Mucronaspis destombesi Henry 1965
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†Mucronaspis ganabina Snajdr 1987
†Mucronaspis grandis Barrande 1852
†Mucronaspis macrophtalma Brongniart 1822
†Mucronaspis olini Temple 1952
Invalid names: Eodalmanitina Henry 1965 [synonym]
G. †Neoprobolium Struve 1985
+
Invalid names: Probolium Oehlert 1889 [replaced]
G. †Odontocephalus Conrad 1840
G. †Odontochile Hawle and Corda 1847
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Subg. †Odontochile (Erbenochile) Alberti 1981
†Odontochile cristata Hawle and Corda 1847
†Odontochile hausmani Brongniart 1822
Invalid names: Hausmannia Hall and Clark 1888 [objective synonym]
G. †Ormathops Delo 1935
+
†Ormathops hupei Destombes 1972
†Ormathops inflatus Zelzko 1922
†Ormathops nicholsoni Salter 1866
G. †Phalangocephalus Campbell 1977
G. †Preodontochile Degardin and Pillet 1984
+
†Preodontochile camprodonensis Degardin and Pillet 1984
†Preodontochile springfieldensis Sanford and Holloway 2008
G. †Prodontochile Kobayashi and Hamada 1971
G. †Reussiana Snajdr 1987
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†Reussiana brevispicula Hörbinger 2000
†Reussiana reussi Barrande 1846
Invalid names: Reussia Maksimova 1972 [replaced]
G. †Roncellia Lesperance and Bourque 1971
G. †Schoharia Lesperance 1975
G. †Synphoria Clarke 1900
+
†Synphoria stemmata Clarke 1900
Invalid names: Eocorycephalus Reed 1925 [objective synonym], Neosynphoria Pillet 1954 [objective synonym]
Subfm. †Synphoriinae Delo 1935
+
G. †Rickardsia Storey et al. 2016
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†Rickardsia mobergi Hede 1915
†Rickardsia nobilis Thomas 1900
G. †Synphoroides Delo 1940
G. †Toletanaspis Rabano 1989
G. †Trypaulites Delo 1935
+
†Trypaulites calypso Hall 1861
Subfm. †Zeliszkellinae Delo 1935
+
G. †Eudolatites Delo 1935
+
Subg. †Eudolatites (Deloites) Destombes 1972
†Eudolatites angelini Barrande 1852
†Eudolatites dubia Barrande 1846
†Eudolatites inflatus Destombes 1972
Invalid names: Banilatites Destombes 1972 [synonym], Destombesites Snadjr 1987 [synonym]
G. †Guichenia Henry 1968
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†Guichenia dufouri Tromelin and Lebesconte 1876
G. †Retamaspis Hammann 1972
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†Retamaspis melendezi Hammann 1972
G. †Zeliszkella Delo 1935
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Subg. †Zeliszkella (Mytocephala) Struve 1958
Subg. †Zeliszkella (Zeliszkella) Delo 1935
†Zeliszkella deshayesi Barrande 1846
G. †Zlichovaspis Pribyl and Vanek 1971
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Subg. †Zlichovaspis (Zlichovaspis) Pribyl and Vanek 1971
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†Zlichovaspis (Zlichovaspis) rugosa Hawle and Corda 1847
Invalid names: Devonodontochile Snajdr 1985 [synonym], Spinodontochile Snajdr 1985 [synonym]
Invalid names: Acastinae Delo 1935 [empty], Dalmanitininae Destombes 1972 [empty]
Fm. †Diaphanometopidae Jaanusson 1959 [trilobite]
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G. †Diaphanometopus Schmidt 1881
G. †Gyrometopus Jaanusson 1975
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†Gyrometopus lineatus Angelin 1854
G. †Prodalmanitina Chugaeva 1968
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†Prodalmanitina nikolayevi Chugaeva 1968
Superfm. †Phacopoidea Hupe 1953
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Fm. †Phacopidae Hawle and Corda 1847
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G. †Acernaspis Campbell 1967
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Subg. †Acernaspis (Eskaspis) Clarkson et al. 1977
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†Acernaspis (Eskaspis) woodburnensis Clarkson et al. 1977
†Acernaspis copperi Chatterton and Ludvigsen 2004
†Acernaspis elliptifrons Esmark 1833
†Acernaspis nanus Maksimova 1962
†Acernaspis orestes Billings 1860
†Acernaspis xynon Howells 1982
Invalid names: Murphycops Lespérance 1968 [synonym]
G. †Acuticryphops Cronier and Feist 2000
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†Acuticryphops acuticeps Kayser 1889
†Acuticryphops klapperi Feist et al. 2009
†Acuticryphops prorotundus Feist and Klapper 2022
G. †Adastocephalum Mitchell 1919
G. †Afrops Alberti 1983
G. †Altaesajania Maksimova 1978
G. †Ananaspis Campbell 1967
+
†Ananaspis crossleii Etheridge and Mitchell 1895
†Ananaspis fecunda Barrande 1852
†Ananaspis macdonaldi Fletcher 1950
G. †Babinops Feist and Becker 1997
+
†Babinops minor Feist et al. 2009
†Babinops planiventer Feist and Becker 1997
G. †Boeckops Chlupac 1972
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†Boeckops boecki Hawle and Corda 1847
†Boeckops delphinoides Chlupac 1972
†Boeckops slimanensis Alberti 1981
†Boeckops stelcki McKellar and Chatterton 2009
†Boeckops zenonis Chlupac 1977
†Boeckops zinkeni Roemer 1843
G. †Burtonops Struve 1990
G. †Denckmannites Wedekind 1911
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†Denckmannites miser Barrande 1852
G. †Dianops Richter 1923
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†Dianops algeriensis Cronier et al. 2013
†Dianops anophthalmus Frech 1894
†Dianops kaufmannii Basse and Lemke 2023
†Dianops limbatus Richter 1848
†Dianops quizhouensis Xiang 1981
†Dianops typhlops Gürich 1896
†Dianops vicarius Chlupac 1961
G. †Dienstina Richter 1931
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†Dienstina diensti Richter and Richter 1923
G. †Ductina Richter 1931
Subfm. †Echinophacopinae Zhou and Campbell 1990
+
G. †Echinophacops Zhou 1983
G. †Toxophacops Zhiquiang and Campbell 1990
+
Subg. †Toxophacops (Atopophacops) Zhou and Campbell 1990
G. †Eldredgeops Struve 1990
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†Eldredgeops rana Green 1832
+
†Phacops rana milleri Stewart 1927
†Phacops rana bellensis Stumm 1953
G. †Enigmapyge Feist et al. 2016
G. †Eophacops Delo 1935
+
Invalid names: Bullicephalus Pribyl and Vanek 1971 [synonym]
G. †Illaenula Chlupac 1977
G. †Inexpectatops Basse 2003
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G. †Kainops Ramsköld and Werdelin 1991
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†Kainops microps Chatterton et al. 1979
†Kainops veles Chlupác 1972
G. †Liolophops Struve 1972
+
†Liolophops sublevatus Struve 1970
†Liolophops syringis Struve 1992
G. †Lochkovella Chlupac 1972
+
†Lochkovella deckeri Delo 1935
G. †Nephranomma Erben 1952
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†Nephranomma drepanomma Erben 1952
†Nephranomma modestus Barrande 1872
G. †Nyterops Struve 1972
+
†Nyterops hollandi Van Viersen 2007
†Nyterops nyter Struve 1970
G. †Paciphacops Maksimova 1971
+
Subg. †Paciphacops (Paciphacops) Maksimova 1972
†Paciphacops altaicus Tschernyschev 1893
†Paciphacops argentinus Thomas 1905
†Paciphacops claviger Haas 1969
†Paciphacops edgecombei Sandford 2005
†Paciphacops latigenalis Etheridge and Mitchell 1896
†Paciphacops logani Hall 1860
†Paciphacops mansfieldensis Etheridge and Mitchell 1896
†Paciphacops waisfeldae Edgecombe and Ramsköld 1994
G. †Phacopidella Reed 1905
+
Subg. †Phacopidella (Prephacopidella) Destombes 1963
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†Phacopidella (Prephacopidella) hupei Nion and Henry 1966
Subg. †Phacopidella (Struveaspis) Alberti 1966
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†Struveaspis fugitiva Barrande 1872
†Struveaspis maroccanica Alberti 1970
†Struveaspis micromma Roemer 1852
†Struveaspis prantli Ruzicka 1946
†Phacopidella glockeri Barrande 1846
Invalid names: Glockeria Wedekind 1914 [objective synonym]
Subfm. †Phacopinae Hawle and Corda 1847
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G. †Austerops McKellar and Chatterton 2009
+
†Austerops kermiti McKellar and Chatterton 2009
†Austerops salamandar McKellar and Chatterton 2009
†Austerops salamandaroides Crônier et al. 2018
†Austerops smoothops Chatterton et al. 2006
†Austerops speculator Alberti 1970
G. †Chlupacops Feist et al. 2016
+
†Chlupacops cryphoides Richter and Richter 1926
†Chlupacops koeneni Holzapfel 1895
†Chlupacops reichi Kegel 1931
†Chlupacops spectabilis Meischner 1965
G. †Chotecops Chlupac 1971
+
†Chotecops auspex Chlupác 1971
†Chotecops breviceps Barrande 1846
†Chotecops chlupaci Sandford 2005
†Chotecops ferdinandi Kayser 1880
†Chotecops glabrens Chlupác 1977
†Chotecops hoseri Hawle and Corda 1847
†Chotecops zizensis Alberti 1983
Invalid names: Cordapeltis Pribyl and Vanek 1971 [synonym]
G. †Cronierella Feist and Klapper 2022
+
†Cronierella expansa Feist and Klapper 2022
†Cronierella postera Feist and Klapper 2022
†Cronierella zofiae Chlupáč 1993
G. †Cryphops Richter and Richter 1926
+
†Cryphops cryptophthalmus Richter and Richter 1926
G. †Echidnops Sandford 2002 [trilobite]
+
†Echidnops serratus Foerste 1888
†Echidnops taphomimus Rustan and Balseiro 2016 [trilobite]
†Echidnops wrighti Sandford 2002
G. †Eocryphops Richter 1931
+
†Eocryphops kayseri Hermann 1911
†Eocryphops termieri Richter and Richter 1943
G. †Feistops Crônier et al. 2021
+
†Feistops mongoliensis Crônier et al. 2021
Tr. †Geesopini Struve 1984
+
G. †Arduennops Struve 1972
G. †Drotops Struve 1990
G. †Hypsipariops Struve 1982
G. †Geesops Struve 1972
+
†Geesops brunopauli Struve 1982
†Geesops schlotheimi Bronn 1825
†Geesops sparsinodosus Struve 1970
G. †Girardina Feist and Klapper 2022
+
†Girardina consimilis Feist and Klapper 2022
†Girardina konradbartzschi Feist and Klapper 2022
†Girardina liopyga Richter 1863
G. †Houseops Feist et al. 2009
+
†Houseops canningensis Feist et al. 2009
†Houseops miserrimus Drevermann 1901
†Houseops nalivkini Maksimova 1955
†Houseops ocellatus Perna 1915
†Houseops olonbulagensis Crônier et al. 2021
G. †Magreanops Van Viersen and Vanherle 2018
+
†Magreanops monachus Van Viersen and Vanherle 2018
†Magreanops renateae Van Viersen and Vanherle 2018
G. †Morocops Basse 2006
+
†Morocops struvei Schraut 2000
Invalid names: Barrandeops McKellar and Chatterton 2009 [synonym]
G. †Occitanella Feist and Klapper 2022
+
†Occitanella hexagonalis Feist and Klapper 2022
†Occitanella linguiformis Feist et al. 2009
†Occitanella postinflexa Feist and Klapper 2022
G. †Omegops Struve 1976
+
†Omegops accipitrinus Phillips 1841
†Omegops cornelius Richter and Richter 1933
†Omegops junggariensis Crônier and Waters 2023
†Omegops maretiolensis Richter and Richter 1933
†Omegops tilabadensis Ghobadi Pour et al. 2018
†Omegops xiangi Zong 2023
Invalid names: Clarksonops Crônier and Waters 2023 [synonym]
G. †Pedinopariops Struve 1972
+
†Pedinopariops brongniarti Steininger 1831
†Pedinopariops degener Barrande 1852
†Pedinopariops hoelleri Van Viersen et al. 2009
†Pedinopariops insequens Chlupác 1983
†Pedinopariops lentigifer Struve 1970
†Pedinopariops major Barrande 1852
†Pedinopariops proniceps Struve 1970
†Pedinopariops regius Chlupáč 1971
†Pedinopariops superstes Barrande 1852
G. †Prokops Chlupac 1971
+
†Prokops benziregensis Alberti 1983
†Prokops hoeninghausi Barrande 1846
†Prokops moorei Sandford 2004
†Prokops prokopi Chlupáč 1971
G. †Reedops Richter and Richter 1925
+
†Reedops albertii Sandford 2005
†Reedops bronni Barrande 1846
†Reedops cephalotes Hawle and Corda 1847
†Reedops decorus Hawle and Corda 1847
†Reedops intermedius Barrande 1846
†Reedops limespragensis Chlupác 1986
†Reedops maurulus Alberti 1969
†Reedops pembertoni McKellar and Chatterton 2009
†Reedops platilegnotor Alberti 1970
†Reedops prospicens Chlupác 1977
†Reedops sternbergi Hawle and Corda 1847
†Reedops struveianus Alberti 1983
G. †Spinicryphops Crônier and Feist 2000
+
†Spinicryphops wocklumeriae Richter and Richter 1926
+
Invalid names: Spinicryphops lemkei Crônier and Feist 2000 [synonym]
G. †Trimerocephaloides Feist et al. 2009
+
†Trimerocephaloides sinevisus Feist et al. 2009
G. †Viaphacops Maximova 1972
+
†Viaphacops canadensis Stumm 1954
†Viaphacops cristatus Hall 1861
†Viaphacops hanusi Chlupáč 1977
†Viaphacops nasutus Stumm 1954
†Viaphacops pipa Hall and Clarke 1888
†Viaphacops salteri Kozlowski 1923
†Viaphacops venezuelensis Carvalho and Moody 2000
G. †Weyerites Crônier and Feist 2000
+
†Weyerites ensae Richter and Richter 1926
G. †Phacops Emmrich 1839
+
Subg. †Phacops (Boeckops) Chlupac 1972
+
Subg. †Phacops (Chasmops) M'Coy 1849
Subg. †Phacops (Chotecops) Chlupac 1971
Subg. †Phacops (Dalmania) Emmrich 1844
+
†Dalmania caudatus Brünnich 1781
Subg. †Phacops (Nephranops) Richter 1926
+
Subg. †Nephranops (Maternia) Feist 2019
Subg. †Nephranops (Nephranops) Richter and Richter 1926
†Nephranops dillanus Richter and Richter 1926
Subg. †Phacops (Omegops) Struve 1976
+
Subg. †Phacops (Phacops) Emmrich 1839
+
†Phacops (Phacops) brocki Feist and Talent 2000
†Phacops (Phacops) nyter Struve 1970
†Phacops (Phacops) oehlerti Morzadec 1969
†Phacops (Phacops) papulatus Richter and Richter 1943
Subg. †Phacops (Portlockia) M'Coy 1849
Subg. †Phacops (Prokops) Chlupac 1971
+
Subg. †Phacops (Pterygometopus) Schmidt 1881
†Phacops alifrons McCoy 1851
†Phacops araw McKellar and Chatterton 2009
†Phacops breviceps Angelin 1854
†Phacops circumspectans Paeckelmann 1913
†Phacops fragosus Struve 1970
†Phacops granulatus Munster 1840
+
Invalid names: Phacops posidoniae Gürich 1896 [synonym]
†Phacops guranensis Maksimova 1969
†Phacops iowensis Delo 1935
†Phacops latifrons Bronn 1825
†Phacops maurusius Alberti 1983
†Phacops maximus Törnquist 1884
†Phacops natlensis Perry and Chatterton 1976
†Phacops ouarouroutensis Crônier et al. 2018
†Phacops papulatus Richter and Richter 1943
†Phacops postaltaicus Maksimova 1960
†Phacops pronini Maksimova 1955
†Phacops spedeni Chatterton 1971
†Phacops tafilaltensis Crônier and Clarkson 2001
†Phacops tamtertensis Cronier et al. 2013
†Phacops transversalis Yi and Xiang 1975
†Phacops turco Richter and Richter 1939
†Phacops zeregensis Cronier et al. 2013
G. †Plagiolaria Kegel 1952
+
†Plagiolaria kockeli Alberti 1968
G. †Rabienops Struve 1989
+
†Rabienops evae Struve 1989
†Rabienops horni Struve 1989
†Rabienops wedekindi Richter and Richter 1926
G. †Rhinophacops Kaneko 1990
G. †Rhinoreedops Maksimova 1978
G. †Sambremeusaspis Lespérance and Sheehan 1987
+
†Sambremeusaspis fossesensis Lespérance and Sheehan 1987
G. †Signatops Pribyl and Vanek 1971
G. †Teichertops Struve 1992
G. †Trifoliops Crônier 2003
+
†Trifoliops cronierae Feist et al. 2016
†Trifoliops trifolius Osmólska 1958
G. †Trimerocephalus M'Coy 1849
+
†Trimerocephalus brevissimus Drevermann 1901
†Trimerocephalus caecus Gürich 1896
†Trimerocephalus chopini Kin and Blazejowski 2013
†Trimerocephalus lelievrei Crônier and Feist 1997
†Trimerocephalus lotzi Drevermann 1901
†Trimerocephalus mimbi Feist et al. 2009
†Trimerocephalus nigritus Crônier 2003
†Trimerocephalus shotoriensis Feist et al. 2003
†Trimerocephalus sulcatus Drevermann 1901
†Trimerocephalus tardispinosus Feist and Becker 1997
Invalid names: Eutrimerocephalus Gortani 1907 [objective synonym]
G. †Zaplaops Baldis and Blasco 1976
Fm. †Pterygometopidae Reed 1905
+
G. †Calliops Delo 1935
+
†Calliops annulatus Raymond 1905
†Calliops armatus Ulrich and Delo 1940
†Calliops brevipostica Cooper 1953
†Calliops brevis Roy 1941
+
Invalid names: Eomonorachus deloi Roy 1941 [synonym]
†Calliops mesleri Cooper 1953
†Calliops troosti Safford and Vogdes 1889
Subfm. †Chasmopinae Pillet 1954
+
G. †Bolbochasmops McNamara 1980
G. †Chasmops M'Coy 1849
+
†Chasmops extensa Boeck 1838
†Chasmops inge Roomusoks 1953
†Chasmops marginatus Schmidt 1881
†Chasmops odini Eichwald 1860
†Chasmops wesenbergensis Schmidt 1881
G. †Oculichasmops Rõõmusoks 1986
G. †Rollmops McNamara 1980
G. †Scopelochasmops McNamara 1980
+
†Scopelochasmops cambrensis Whittington 1962
G. †Toxochasmops McNamara 1979
+
Subg. †Toxochasmops (Schmidtops) Roomusoks 1998
Subg. †Toxochasmops (Toxochasmops) McNamara 1979
G. †Uralops Antsygin 1970
G. †Valdariops Rõõmusoks 2000
+
†Valdariops aliquantulus Rõõmusoks 2000
†Valdariops angustus Rõõmusoks 2000
†Valdariops eichwaldi Schmidt 1881
†Valdariops kernuensis Rõõmusoks 2000
Subfm. †Eomonorachinae Pillet 1954
+
G. †Calyptaulax Cooper 1930
+
†Calyptaulax angusta Cooper 1953
†Calyptaulax annulata Raymond 1905
+
Invalid names: Pterygometopus gracilens Raymond 1925 [synonym]
†Calyptaulax brongniartii Portlock 1843
†Calyptaulax callirachis Cooper 1953
+
Invalid names: Calliops loxorachis Cooper 1953 [synonym]
†Calyptaulax cornwalli Ross and Barnes 1967
†Calyptaulax glabella Cooper 1930
†Calyptaulax holstonensis Raymond 1925
†Calyptaulax hunteri Reed 1914
†Calyptaulax incepta Whittington 1965
†Calyptaulax norvegicus Stormer 1945
†Calyptaulax strasburgensis Ulrich and Delo 1940
G. †Denella Ludvigsen and Chatterton 1982
+
†Denella cumera Ludvigsen and Chatterton 1982
†Denella goodridgii Schuchert 1900
G. †Eomonorachus Delo 1935
+
†Eomonorachus buttsi Cooper 1953
†Eomonorachus divaricatus Frederickson 1964
†Eomonorachus goodridgii Schuchert 1900
†Eomonorachus intermedius Walcott 1879
G. †Liocnemis Kielan 1960
G. †Podowrinella Clarkson et al. 1977
+
G. †Sceptaspis Ludvigsen and Chatterton 1982
+
†Sceptaspis lincolnensis Branson 1909
+
Invalid names: Calliops huronensis Okulitch 1942 [synonym], Calyptaulax calderi Wilson 1947 [synonym], Calyptaulax conjuens Foerste 1919 [synonym], Calyptaulax ottawanus Okulitch 1942 [synonym]
G. †Tricopelta Ludvigsen and Chatterton 1982
+
†Tricopelta breviceps Hall 1872
†Tricopelta mackenziensis Ludvigsen and Chatterton 1982
Subfm. †Monorakinae Kramarenko 1952
+
G. †Carinopyge Maksimova 1955
+
†Carinopyge abscisa Maksimova 1955
†Carinopyge fracta Masimova 1955
G. †Ceratevenkaspis Kramarenko 1952
+
†Ceratevenkaspis armata Kramaranko 1952
†Ceratevenkaspis parnaica Balashova 1960
†Ceratevenkaspis pentagona Balashova 1960
†Ceratevenkaspis taimyrica Balashova 1960
G. †Elasmaspis Kramarenko 1956
+
†Elasmaspis speciosa Kramarenko 1956
G. †Evenkaspis Kramarenko 1952
+
Subg. †Evenkaspis (Evenkaspis) Kramarenko 1952
Subg. †Evenkaspis (Parevenkaspis) Kramarenko 1952
G. †Monorakos Schmidt 1886
+
†Monorakos consimilis Semenova 1978
†Monorakos kledos Ormiston 1978
†Monorakos lopatini Schmidt 1886
†Monorakos magnus Kramarenko 1952
†Monorakos morkokensis Maksimova 1962
†Monorakos mutabilis Kramarenko 1952
†Monorakos ossacrum Kramarenko 1962
†Monorakos planiusculus Kramarenko 1957
†Monorakos planus Kramarenko 1957
†Monorakos tunguskaensis Kramarenko 1952
G. †Parevenkaspis Kramarenko 1952
Subfm. †Pterygometopinae Reed 1905
+
G. †Achatella Delo 1935
+
Subg. †Achatella (Achatella) Delo 1935
†Achatella achates Billings 1860
+
Invalid names: Pterygometopus billingsi Wilson 1947 [synonym]
†Achatella katharina Bradley 1930
†Achatella kuckersianus Schmidt 1881
†Achatella schmidti Warburg 1925
†Achatella truncatocaudata Portlock 1843
Invalid names: Vironiaspis Jaanusson and Ramsköld 1993 [synonym]
G. †Estoniops Mannil 1958
+
†Estoniops alifrons McCoy 1851
†Estoniops bekkeri Mannil 1958
†Estoniops exilis Eichwald 1858
†Estoniops fjaeckensis Jaanusson and Ramsköld 1993
†Estoniops maennili Jaanusson and Ramsköld 1993
†Estoniops oculeus Antsygin 1970
†Estoniops panderi Schmidt 1881
†Estoniops sandbyensis Olin 1906
G. †Ingriops Jaanusson and Ramskold 1993
+
†Ingriops intermedius Schmidt 1881
†Ingriops trigonocephala Schmidt 1881
G. †Isalaux Frederickson and Pollack 1952
+
†Isalaux canyonensis Frederickson and Pollack 1952
G. †Keilapyge Jaanusson and Ramsköld 1993
+
†Keilapyge laevigatus Schmidt 1881
†Keilapyge latus Antsygin 1970
G. †Oelandiops Jaanusson and Ramsköld 1993
+
†Oelandiops mirificus Jaanusson and Ramsköld 1993
G. †Pterygometopus Schmidt 1881
+
Subg. †Pterygometopus (Pterygometopus) Schmidt 1881
†Pterygometopus recurvus Brogger 1887
†Pterygometopus schmidti Warburg 1925
†Pterygometopus sclerops Dalman 1827
G. †Upplandiops Jaanusson and Ramsköld 1993
+
†Upplandiops calvus Jaanusson and Ramsköld 1993
G. †Yanhaoia Zhou et al. 1998
+
G. †Truncatometopus Pillet 1960
G. †Volkops Struve 1959
+
†Volkops volki Struve 1959
G. †Yankaoia Zhou et al. 1998
Invalid names: Creatarginae Tripp 1957 [empty]
Fm. †Prosopiscidae Fortey and Shergold 1984
hide
G. †Prosopiscus Salter 1865
+
†Prosopiscus praecox Fortey and Shergold 1984
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: low Mg calcitec
Composition 2: chitinc
Ontogeny: molting, addition of partsc
Environment: marinec
Locomotion: fast-movingc
Life habit: low-level epifaunalc
Diet: carnivoresubo
Created: 2012-08-27 22:18:01
Modified: 2012-08-27 07:18:01
Source: subo = suborder, c = class
References: Fortey and Owens 1999, Aberhan et al. 2004

Age range: base of the Tremadocian to the top of the Hastarian or 486.85000 to 353.70000 Ma

Collections (2728 total)

Oldest occurrences

Time interval Ma Country or state Original ID and collection number
Ibexian488.9 - 471.3USA (Nevada) Calyptaulax sp. (104439)
Tremadocian486.85 - 477.1Morocco (Anti-Atlas) Toletanaspis sp. (166727)
Tremadocian486.85 - 477.1Morocco Toletanaspis sp. (235487)
Floian477.1 - 471.3Morocco (Anti Atlas) Toletanaspis borni (166726 166730)
Floian477.1 - 471.3Morocco (Tarfaya) Toletanaspis borni (56583)
Floian477.1 - 471.3Morocco Dalmanitidae indet. (235492) Ormathops clariondi, Toletanaspis borni (56592) Toletanaspis sp. (235491)
Klabava477.1 - 469.4Czechia (Prague) Ormathops sp. (76457)
Klabava477.1 - 469.4Czechia Ormathops sp. (3841)
Arenig477.1 - 468.0China (GanSu) Dalmanitina sp. (8575)
Arenig477.1 - 468.0China (YunNan) Prosopiscus sp. (7734)

Youngest occurrences
Time interval Ma Country or state PBDB collection number
Late/Upper Famennian365.2 - 358.86China (Guizhou) Dianops guizhouensis (3954) Phacops granulatus (218957 218958) Phacops guilinensis (234696)
Late/Upper Famennian365.2 - 358.86Austria Phacops granulatus (38596 38608 42404 42406)
Late/Upper Famennian365.2 - 358.86Poland Dianops anophthalmus (134803)
Late/Upper Famennian365.2 - 358.86Iran (Alborz) Omegops tilabadensis (234970 234971 234973 234974)
Late/Upper Famennian365.2 - 358.86Germany (Thuringen) Dianops bartzschi, Cryphops ensae (235215)
Late/Upper Famennian365.2 - 358.86Belgium (Namur) Phacops accipitrinus (223663)
Late/Upper Famennian365.2 - 358.86Belgium Omegops maretiolensis, Omegops accipitrinus (223664) Phacops maretiolensis (234969) Phacops sp. (235220)
Late/Upper Famennian365.2 - 358.86China (Xinjiang) Omegops honggulelengensis (234672)
Late/Upper Famennian365.2 - 358.86France Cryphops acuticeps (234854) Phacops wedekindi, Phacops granulatus, Phacops sp. (38641)
Hastarian358.86 - 353.7France Phacops sp. (38642)