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Archaeorthoptera

Insecta

Synonymy list
YearName and author
2002Archaeorthoptera Béthoux and Nel p. 14
2005Archaeorthoptera Béthoux
2007Archaeorthoptera Béthoux
2009Archaeorthoptera Zhang et al.

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Protostomia
Ecdysozoa
Panarthropoda
phylumArthropodaLatreille 1829
superclassHexapodaLatreille 1825
classInsecta
RankNameAuthor
Dicondylia
Paranotalia
subclassPterygota()
NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
PolyneopteraMartynov 1938
superorderArchaeorthoptera
superorderArchaeorthoptera

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Superor. Archaeorthoptera Béthoux and Nel 2002
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G. †Aviologus Coty et al. 2014
+
Aviologus duquesnei Coty et al. 2014
G. †Bethouxia Prokop et al. 2015
+
Bethouxia ariegensis Prokop et al. 2015
Fm. †Chresmodidae Haase 1890
+
G. †Chresmoda Germar 1839
+
Chresmoda aquatica Martínez-Delclòs 1989
Chresmoda chikuni Zhang and Ge 2017
Chresmoda libanica Nel et al. 2004
Chresmoda mongolica Ponomarenko 1986
Chresmoda multinervis Zhang et al. 2009
Chresmoda neotropica Engel and Heads 2008
Chresmoda obscura Germar 1839
+
Invalid names: Halometra gigantea Germar 1839 [synonym], Halometra minor Oppenheim 1888 [synonym], Propygolampis bronni Weyenbergh 1874 [synonym], Pygolampis gigantea Germar 1839 [synonym], Sternarthron zitteli Haase 1890 [synonym]
Chresmoda orientalis Esaki 1949
Chresmoda oweni Westwood 1854
Chresmoda shihi Zhang et al. 2009
Invalid names: Gryllidium Westwood 1854 [synonym], Halometra Oppenheim 1888 [synonym], Propygolampis Weyenbergh 1874 [synonym], Saurophthiroides Ponomarenko 1986 [synonym], Sternarthron Haase 1890 [synonym]
G. †Jurachresmoda Zhang et al. 2008
+
Jurachresmoda gaskelli Zhang et al. 2008
Jurachresmoda sanyica Zhang et al. 2009
G. †Sinochresmoda Zhang et al. 2008
+
Sinochresmoda magnicornia Zhang et al. 2008
Or. †Cnemidolestodea Handlirsch 1937
+
G. †Aviocladus Prokop et al. 2014
+
Aviocladus pectinatus Prokop et al. 2014
G. †Aviohapaloptera Prokop et al. 2014
+
Aviohapaloptera bethouxi Prokop et al. 2014
Subor. †Cnemidolestina Handlirsch 1937
+
Fm. †Cnemidolestidae Handlirsch 1906
+
G. †Aetophlebia Scudder 1885
+
Aethophlebia singularis Scudder 1885
G. †Amphiboliacridites Langiaux and Parriat 1974
+
Amphiboliacridites joanesi Langiaux and Parriat 1974
G. †Anarkemina Aristov 2014
+
G. †Argentinonarkemina Martins-Neto et al. 2007
+
G. †Bouleites Lameere 1917
+
Bouleites latipennis Lameere 1917
G. †Carbonokata Aristov 2013
+
Carbonokata storozhenkoi Aristov 2013
G. †Cnemidolestes Handlirsch 1906
+
Cnemidolestes woodwardi Brongniart 1883
G. †Evenkiophlebia Aristov 2013
+
Evenkiophlebia collucata Aristov 2013
G. †Irajanarkemina Martins-Neto et al. 2007
+
Irajanarkemina rodendorfi Pinto and de Ornellas 1978
G. †Narkema Handlirsch 1911
+
Narkema alternatum Cockerell 1924
Narkema taeniatum Handlirsch 1911
G. †Narkemina Martynov 1930
+
Narkemina angustata Martynov 1930
Narkemina angustiformis Sharov 1961
Narkemina genuina Sharov 1961
Narkemina kata Aristov 2013
G. †Narkeminopsis Whalley 1979
+
Narkeminopsis eddi Whalley 1979
Narkeminopsis inversa Aristov 2013
Narkeminopsis sowiaki Brauckmann and Herd 2005
G. †Narkeminuta Aristov 2013
+
Narkeminuta permiana Aristov 2013
G. †Narkemulla Aristov 2013
+
Narkemulla sibirica Aristov 2013
G. †Palatinarkema Nel and Poschmann 2021
+
Palatinarkema prokopi Nel and Poschmann 2021
G. †Paranarkemina Pinto and De Ornellas 1980
+
Paranarkemina kurtzi Pinto and De Ornellas 1980
+
Invalid names: Paranarkemina velizensis Pinto and De Ornellas 1981 [synonym]
G. †Piesbergopterum Dvořák et al. 2021
+
Piesbergopterum avionensis Nel and Roques 2021
Piesbergopterum punctatum Dvořák et al. 2021
G. †Protodiamphipnoa Brongniart 1894
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Protodiamphipnoa gaudryi Brongniart 1885
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Invalid names: Protodiamphipnoa tertrini Brongniart 1894 [synonym]
G. †Tshunoptera Aristov 2013
+
Tshunoptera ampla Aristov 2013
G. †Velizphlebia Martins-Neto et al. 2007
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Velizphlebia cruzi Martins-Neto et al. 2007
Invalid names: Aetophlebiidae Handlirsch 1906 [synonym], Narkemidae Handlirsch 1911 [synonym], Narkeminidae Storozhenko 1996 [synonym]
Fm. †Ctenoptilidae Aristov 2014
+
G. †Ctenoptilus Lameere 1917
+
Ctenoptilus elongatus Brongniart 1894
+
Invalid names: Eoblattina complexa Bolton 1925 [synonym], Homalophlebia trouessarti Meunier 1911 [synonym]
Ctenoptilus frequens Chen et al. 2021
Invalid names: Eoblattina Bolton 1925 [synonym], Ischnoneurilla Handlirsch 1919 [synonym], Ischnoneurula Handlirsch 1919 [objective synonym]
Fm. †Cymenophlebiidae Pruvost 1919
+
G. †Cymenophlebia Pruvost 1919
+
Cymenophlebia carpentieri Pruvost 1919
Fm. †Gerapompidae Handlirsch 1906
+
G. †Aenigmatella Sharov 1961
+
G. †Gerapompus Scudder 1885
+
Gerapompus blattinoides Scudder 1885
Gerapompus extensus Scudder 1885
Gerapompus schucherti Handlirsch 1906
Fm. †Issatermidae Aristov 2018
+
G. †Issatermes Aristov 2018
+
Issatermes parvus Aristov 2018
Fm. †Neraphidiidae Aristov 2014
+
G. †Fodinopenna Aristov 2014
+
Fodinopenna abakanica Aristov 2014
G. †Neraphidia Novokshonov and Novokshonova 1997
+
Neraphidia mitis Novokshonov and Novokshonova 1997
Neraphidia rigida Aristov 2004
G. †Yaria Aristov 2014
+
Yaria arenaria Aristov 2014
Fm. †Prygidae Aristov and Rasnitsyn 2014
+
G. †Pryg Aristov and Rasnitsyn 2014
+
Pryg absurdus Aristov and Rasnitsyn 2014
Fm. †Psoropteridae Carpenter 1976
+
G. †Hammelburgia Aristov 2014
+
Hammelburgia selli Aristov 2014
G. †Psoroptera Carpenter 1976
+
Psoroptera cubitalia Carpenter 1976
Fm. †Spanioderidae Handlirsch 1906
+
G. †Carpenteroptera Pinto 1990
+
Carpenteroptera onzii Pinto 1990
Carpenteroptera rochacamposi Pinto and de Ornellas 1978
G. †Chenxiella Liu et al. 2009
+
Chenxiella liuae Liu et al. 2009
G. †Dieconeura Scudder 1885
+
Dieconeura arcuata Scudder 1885
Dieconeura mazona Handlirsch 1911
G. †Dieconeurites Handlirsch 1906
+
Dieconeurites rigidus Scudder 1885
G. †Etotabla Béthoux and Jarzembowski 2010
+
Etotabla gillae Béthoux and Jarzembowski 2010
G. †Izykhia Aristov 2013
+
Izykhia tridentis Aristov 2013
G. †Longzhua Gu et al. 2011
+
Longzhua loculata Gu et al. 2011
G. †Miamia Dana 1864
+
Miamia bronsoni Dana 1864
+
Invalid names: Cheliphlebia extensa Melander 1903 [synonym], Dictyoneura clarinervis Melander 1903 [synonym], Metryia analis Handlirsch 1906 [synonym], Petromartus indistinctus Melander 1903 [synonym], Propteticus infernus Scudder 1885 [synonym], Spaniodera acutipennis Handlirsch 1911 [synonym], Spaniodera ambulans Handlirsch 1906 [synonym], Spaniodera angusta Handlirsch 1911 [synonym], Spaniodera elatior Handlirsch 1911 [synonym], Spaniodera lata Handlirsch 1911 [synonym], Spaniodera longicollis Handlirsch 1911 [synonym], Spaniodera parvula Handlirsch 1911 [synonym], Spaniodera schucherti Handlirsch 1911 [synonym], Termes contusus Scudder 1878 [synonym]
Miamia maimai Béthoux et al. 2012
Miamia rossorum Béthoux and Jarzembowski 2010
Miamia silvatica Laurentiaux and Laurentiaux-Vieira 1980
Invalid names: Anthraconeura Laurentiaux and Laurentiaux-Vieira 1980 [synonym], Camptophlebia Handlirsch 1906 [synonym], Didymophleps Scudder 1885 [synonym], Metryia Handlirsch 1906 [synonym], Paracheliphlebia Handlirsch 1906 [synonym], Petromartus Melander 1903 [synonym], Propteticus Scudder 1885 [synonym], Spaniodera Handlirsch 1906 [synonym]
G. †Sinopteron Prokop and Ren 2007
+
Sinopteron huangheense Prokop and Ren 2007
Invalid names: Anthraconeuridae Laurentiaux and Laurentiaux-Vieira 1980 [synonym], Carpenteropteridae Pinto and Ornellas 1991 [synonym]
Fm. †Sylvabestiidae Aristov 2000
+
G. †Aiban Aristov 2018
+
Aiban kichineis Aristov 2018
G. †Cerasopterum Kukalová 1964
+
Cerasopterum gracile Kukalová 1964
G. †Elmopterum Béthoux and Beckemeyer 2007
+
Elmopterum rotundum Béthoux and Beckemeyer 2007
G. †Ivkinus Aristov 2014
+
Ivkinus conjunctus Aristov and Storozhenko 2011
G. †Kazanalicula Aristov and Storozhenko 2011
+
Kazanalicula reducta Aristov and Storozhenko 2011
G. †Kityakia Aristov 2014
+
Kityakia attrita Aristov 2014
G. †Micropermula Storozhenko 1992
+
Micropermula ceceya Aristov 2018
Micropermula mongolica Storozhenko 1992
G. †Neprotembia Aristov 2004
+
Neprotembia complicata Aristov 2005
Neprotembia truncata Aristov 2004
G. †Sharovala Aristov 2020
+
Sharovala triassica Aristov 2020
G. †Sibestia Aristov 2018
+
Sibestia nana Aristov 2018
G. †Sojanopermula Storozhenko 1992
+
Sojanopermula edemskii Aristov and Storozhenko 2011
Sojanopermula lucida Storozhenko 1992
Sojanopermula minor Aristov 2004
Sojanopermula rasnitsyni Aristov and Storozhenko 2011
G. †Sylvabestia Aristov 2000
+
Sylvabestia tenuis Aristov 2000
G. †Tshepanichoptera Aristov 2008
+
Tshepanichoptera lacera Aristov 2008
G. †Vokhmia Aristov 2018
+
Vokhmia laxa Aristov 2018
Vokhmia pyriformis Aristov 2018
G. †Votyak Aristov 2018
+
Votyak pictus Aristov 2018
Fm. †Tillyardembiidae Zalessky 1938
+
G. †Heteroptilon Carpenter 1976
+
Heteroptilon costale Carpenter 1976
G. †Kamamica Aristov and Rasnitsyn 2014
+
Kamamica promota Aristov and Rasnitsyn 2014
G. †Kungurembia Aristov 2004
+
Kungurembia biarmica Rasnitsyn and Aristov 2010
Kungurembia brevicervix Aristov 2004
Kungurembia pallida Aristov 2004
G. †Paralongzhua Prokop et al. 2015
+
Paralongzhua elongata Prokop et al. 2015
G. †Permedax Aristov 2004
+
Permedax effertus Aristov 2004
G. †Tillyardembia Zalessky 1937
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Tillyardembia antennaeplana Zalessky 1937
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Invalid names: Permocapnia brevipes Martynov 1940 [synonym], Tillyardembia biarmica Zalessky 1937 [synonym], Tillyardembia minuta Zalessky 1950 [synonym]
Tillyardembia ravisedorum Vilesov and Novokshonov 1993
Tillyardembia zalazna Aristov 2018
Invalid names: Permocapnia Martynov 1940 [synonym]
G. †Udembia Aristov 2018
+
Udembia udmurtica Aristov 2018
Invalid names: Heteroptilidae Carpenter 1976 [synonym], Permocapniidae Martynov 1940 [synonym]
Subor. †Parmapterina Aristov 2016
+
Fm. †Necrophasmidae Martynov 1925
+
G. †Ferganamadygenia Storozhenko and Vršanský 1995
+
Ferganamadygenia plicata Storozhenko and Vršanský 1995
G. †Juraperla Huang and Nel 2007
+
Juraperla daohugouensis Huang and Nel 2007
Juraperla grandis Cui et al. 2010
G. †Necrophasma Martynov 1925
+
Necrophasma shabarovi Martynov 1925
Invalid names: Juraperlidae Huang and Nel 2007 [synonym]
Fm. †Parmapteridae Aristov and Rasnitsyn 2015
+
G. †Heterologus Carpenter 1943
+
Heterologus duyiwuer Béthoux et al. 2012
Heterologus langfordorum Carpenter 1943
G. †Opisthocladus Carpenter 1976
+
Opisthocladus arcuatus Carpenter 1976
G. †Parmaptera Aristov and Rasnitsyn 2015
+
Parmaptera permiana Aristov and Rasnitsyn 2015
Fm. †Pinideliidae Storozhenko 1997
+
G. †Idelopterum Aristov 2015
+
Idelopterum samoedum Aristov 2015
G. †Kishertia Aristov 2004
+
Kishertia tricubitalis Aristov 2004
G. †Pinidelia Storozhenko 1994
+
Pinidelia sukatshevae Storozhenko 1994
Fm. †Protembiidae Tillyard 1937
+
G. †Aibolitus Novokshonov and Storozhenko 1996
+
Aibolitus medicinus Novokshonov and Storozhenko 1996
Aibolitus minutus Béthoux and Beckemeyer 2007
G. †Barmaleus Novokshonov 1997
+
Barmaleus dentatus Novokshonov 1997
G. †Kirovopteron Aristov 2015
+
Kirovopteron luteus Aristov 2015
G. †Kuplya Aristov 2013
+
Kuplya minutissima Aristov 2013
G. †Paratillyardembia Aristov 2000
+
G. †Parbarmaleus Novokshonov 1997
+
Parbarmaleus sojanensis Novokshonov 1997
G. †Protembia Tillyard 1937
+
Protembia permiana Tillyard 1937
+
Invalid names: Telactinopteryx striatipennis Tillyard 1937 [synonym]
Invalid names: Telactinopteryx Tillyard 1937 [synonym]
G. †Repka Aristov and Rasnitsyn 2011
+
Repka curta Aristov and Rasnitsyn 2011
Repka repens Aristov and Rasnitsyn 2011
Repka stramenis Aristov and Rasnitsyn 2011
G. †Sigmophlebia Béthoux and Beckemeyer 2007
+
Sigmophlebia engeli Béthoux and Beckemeyer 2007
Sigmophlebia rugulosa Aristov 2013
Sigmophlebia udmurtica Aristov 2008
G. †Sojanoraphidia Martynova 1952
+
Sojanoraphidia martynovae Storozhenko and Novokshonov 1995
Sojanoraphidia rossica Martynova 1952
G. †Soyanocadaver Aristov 2015
+
Soyanocadaver crypticus Aristov 2015
G. †Sylvardembia Novokshonov 1997
+
Sylvardembia matura Aristov 2000
Sylvardembia tamaena Novokshonov 1997
G. †Tshekardembia Novokshonov 1995
+
Tshekardembia sharovi Novokshonov 1995
G. †Tshekardomina Novokshonov and Aristov 2002
+
Tshekardomina imbecilla Aristov 2004
Tshekardomina imbicilla Aristov 2004
Tshekardomina maculosa Novokshonov and Aristov 2002
Tshekardomina mongolica Aristov 2009
G. †Ventopterum Kukalová 1964
+
Ventopterum rapidum Kukalová 1964
Invalid names: Sojanoraphidiidae Martynova 1952 [synonym], Sylvardembiidae Novokshonov 2000 [synonym], Telactinopterygidae Carpenter 1943 [synonym], Tshekardominidae Novokshonov and Aristov 2002 [synonym]
Fm. †Roqueiidae Nel et al. 2022
+
G. †Roqueia Nel et al. 2022
+
Roqueia alpina Nel et al. 2022
Fm. †Tococladidae Carpenter 1966
+
G. †Elmocladus Rasnitsyn and Aristov 2021
+
Elmocladus strictus Carpenter 1976
G. †Tococladus Carpenter 1966
+
Tococladus garrici Béthoux et al. 2003
Tococladus kargalensis Rasnitsyn and Aristov 2013
Tococladus rallus Carpenter 1966
Tococladus similis Novokshonov 1999
G. †Tshekardocladus Aristov and Rasnitsyn 2015
+
Tshekardocladus sparsus Aristov and Rasnitsyn 2015
G. †Xixia Gu et al. 2014
+
Xixia huban Gu et al. 2014
G. †Contracladus Dvořák et al. 2021
+
Contracladus impar Dvořák et al. 2021
G. †Coselia Bolton 1922
+
Coselia palmiformis Bolton 1922
Fm. †Emphylopteridae Handlirsch 1922
+
G. †Emphyloptera Pruvost 1919
+
Emphyloptera lecrivaini Pruvost 1919
Fm. †Eoblattidae Handlirsch 1906
+
G. †Anegertus Handlirsch 1911
+
Anegertus cubitalis Handlirsch 1911
G. †Beloatta Nel et al. 2020
+
Beloatta duquesnei Nel et al. 2020
G. †Eoblatta Handlirsch 1906
+
Eoblatta robusta Brongniart 1894
Eoblatta zeilleri Langiaux and Parriat 1974
G. †Lobeatta Béthoux 2005
+
Lobeatta schneideri Béthoux 2005
G. †Lomovatka Aristov 2015
+
Lomovatka udovichenkoi Aristov 2015
G. †Forfexala Béthoux and Herd 2009
+
Forfexala kiarae Béthoux and Herd 2009
Fm. †Ischnoneuridae Handlirsch 1906
+
G. †Ischnoneura Brongniart 1894
+
Ischnoneura brongniarti Lameere 1917
Ischnoneura delicatula Brongniart 1894
Ischnoneura oustaleti Brongniart 1885
Invalid names: Ischnoneurona Handlirsch 1919 [synonym], Leptoneura Brongniart 1885 [nomen nudum]
G. †Permarrhaphus Martynov 1931
+
Permarrhaphus venosus Cockerell 1927
Invalid names: Adiphlebiidae Handlirsch 1906 [synonym], Stenoneurellidae Handlirsch 1919 [synonym]
G. †Lodevolongzhua Prokop et al. 2015
+
Lodevolongzhua incompleta Prokop et al. 2015
Fm. †Nugonioneuridae Carpenter 1976
+
G. †Avionugonioneura Nel and Roques 2021
+
Avionugonioneura jouaulti Nel and Roques 2021
G. †Duquesnia Nel and Roques 2021
+
Duquesnia gallica Nel and Roques 2021
G. †Nugonioneura Tillyard 1937
+
Nugonioneura problematica Tillyard 1937
Fm. †Omaliaidae Handlirsch 1906
+
G. †Anthrakoris Richardson 1956
+
Anthrakoris aetherius Richardson 1956
G. †Omalia Van Beneden and Coemans 1867
+
Omalia macroptera Van Beneden and Coemans 1867
Invalid names: Coseliidae Bolton 1922 [synonym]
G. †Omaliella Béthoux and Nel 2005
+
Omaliella polonica Dvořák et al. 2019
Omaliella ramosa Béthoux and Nel 2005
G. †Palaeomastax Handlirsch 1904
+
Palaeomastax carbonis Handlirsch 1904
G. †Paleoischnoptera Schubnel et al. 2019
+
Paleoischnoptera diaphanes Béthoux and Nel 2005
Invalid names: Ischnoptera Béthoux and Nel 2005 [replaced]
Unr. Panorthoptera Crampton 1928
+
G. †Bruaylogus Coty et al. 2014
+
Bruaylogus magnificus Coty et al. 2014
Fm. †Cacurgidae Handlirsch 1911
+
G. †Cacurgus Handlirsch 1911
+
Cacurgus avionensis Schubnel et al. 2019
Cacurgus spilopterus Handlirsch 1911
G. †Heterologopsis Brauckmann and Koch 1982
+
Heterologopsis ruhrensis Brauckmann and Koch 1982
G. †Spilomastax Handlirsch 1911
+
Spilomastax oligoneurus Handlirsch 1911
Or. †Caloneurodea Martynov 1938
+
Fm. †Apsidoneuridae Carpenter 1961
+
G. †Apsidoneura Carpenter 1943
+
Apsidoneura flexa Carpenter 1943
G. †Homaloptila Handlirsch 1919
+
Homaloptila similis Meunier 1911
G. †Sinaspidoneura Huang et al. 2020
+
Sinaspidoneura magnifica Huang et al. 2020
G. †Aviogramma Prokop et al. 2014
+
Aviogramma gracilis Prokop et al. 2014
Fm. †Caloneuridae Handlirsch 1906
+
G. †Caloneura Brongniart 1885
+
Caloneura dawsoni Brongniart 1885
+
Invalid names: Caloneura longicornis Handlirsch 1919 [synonym], Caloneura major Handlirsch 1919 [synonym], Caloneura picta Handlirsch 1919 [synonym], Caloneura robusta Handlirsch 1919 [synonym], Caloneura subtilis Bolton 1925 [synonym], Homaloneura royeri Meunier 1911 [synonym]
Caloneura dawsonii Brongniart 1885
Invalid names: Boltonaloneura Rasnitsyn 2002 [synonym], Confusio Handlirsch 1919 [synonym]
G. †Caloneurella Carpenter 1934
+
Caloneurella carbonaria Carpenter 1934
G. †Eohymen Martynov 1937
+
Eohymen maculipennis Martynov 1937
G. †Isadistica Rasnitsyn and Aristov 2013
+
Isadistica issada Rasnitsyn and Aristov 2013
Isadistica longa Rasnitsyn and Aristov 2013
G. †Ligogramma Béthoux et al. 2004
+
Ligogramma sinuosa Béthoux et al. 2004
Ligogramma wichita Beckemeyer 2009
G. †Pleisiogramma Carpenter 1943
+
Pleisiogramma maueri Novokshonov 1998
Pleisiogramma medialis Carpenter 1943
+
Invalid names: Pleisiogramma reducta Carpenter 1943 [synonym]
G. †Vilviopsis Martynov 1938
+
Vilviopsis extensa Martynov 1938
Vilviopsis picta Rasnitsyn 2004
Invalid names: Amboneuridae Carpenter 1980 [synonym], Anomalogrammatidae Carpenter 1943 [synonym], Eohymenidae Martynov 1937 [synonym], Euthygrammatidae Martynov 1928 [synonym], Paleuthygrammatidae Carpenter 1943 [synonym], Plesiogrammatidae Carpenter 1943 [synonym], Sthenaroceridae Handlirsch 1906 [synonym]
G. †Gallogramma Garrouste et al. 2017
+
Gallogramma galadrieli Garrouste et al. 2017
G. †Gigagramma Béthoux et al. 2004
+
Gigagramma carpenteri Béthoux et al. 2004
Fm. †Hapalopteridae Handlirsch 1906
+
G. †Aenigmatodes Handlirsch 1906
+
Aenigmatodes danielsi Handlirsch 1906
G. †Arroyopteron Rasnitsyn 2004
+
Arroyopteron indolatum Rasnitsyn 2004
G. †Carrizarroyo Rasnitsyn 2004
+
Carrizarroyo calopterus Rasnitsyn 2004
G. †Carrizopteron Rasnitsyn 2004
+
Carrizopteron longum Rasnitsyn 2004
G. †Endoiasmus Handlirsch 1906
+
Endoiasmus reticulatus Handlirsch 1906
G. †Geroneura Matthew 1889
+
Geroneura ovata Bolton 1921
Geroneura wilsoni Matthew 1889
G. †Hapaloptera Handlirsch 1906
+
Hapaloptera gracilis Handlirsch 1906
Hapaloptera obovata Bolton 1934
Hapaloptera parallela Bolton 1934
G. †Protokollaria Brongniart 1885
+
Protokollaria ingens Brongniart 1885
G. †Spiculum Brauckmann and Herd 2005
+
Spiculum mendicum Brauckmann and Herd 2005
G. †Tshecalculus Novokshonov 2000
+
Tshecalculus inaspectus Novokshonov 2000
Invalid names: Aenigmatodidae Handlirsch 1906 [synonym], Protokollariidae Handlirsch 1906 [synonym]
G. †Jarmiloptera Prokop et al. 2015
+
Jarmiloptera mouralensis Prokop et al. 2015
Fm. †Lodevogrammatidae Nel et al. 2023
+
G. †Lodevogramma Nel et al. 2023
+
Lodevogramma pumilia Nel et al. 2023
G. †Lusitaneura Loureiro et al. 2010
+
Lusitaneura covensis Loureiro et al. 2010
G. †Nanogramma Béthoux et al. 2004
+
Nanogramma gandi Béthoux et al. 2004
G. †Oudardgramma Nel and Roques 2021
+
Oudardgramma bruayensis Nel and Roques 2021
G. †Paleuthygramma Martynov 1930
+
Paleuthygramma acuta Carpenter 1943
+
Invalid names: Apsidoneura reducta Zimmerman 1962 [synonym]
Paleuthygramma sharovi Béthoux et al. 2004
Paleuthygramma sottyi Burnham 1984
Paleuthygramma tenuicorne Martynov 1930
Paleuthygramma ventricosa Rasnitsyn 2004
G. †Paraspiculum Dvořák et al. 2021
+
Paraspiculum brauckmanni Dvořák et al. 2021
Fm. †Permobiellidae Tillyard 1937
+
G. †Aviobiella Nel and Roques 2021
+
Aviobiella garroustei Nel and Roques 2021
G. †Permobiella Tillyard 1937
+
Permobiella perspicua Tillyard 1937
G. †Pseudobiella Carpenter 1970
+
Pseudobiella fasciata Carpenter 1970
G. †Pruvostiella Handlirsch 1922
+
Pruvostiella lecomtei Pruvost 1919
Invalid names: Euthyneura Pruvost 1919 [replaced]
Invalid names: Hapalopteroidea Handlirsch 1906 [synonym], Pleisiogramma kemerovoense Martynova 1952 [nomen dubium], Stenaroceridae [empty]
Fm. †Geraridae Scudder 1885
+
G. †Anepitedius Handlirsch 1911
+
Anepitedius giraffa Handlirsch 1911
G. †Axiologus Handlirsch 1906
+
Axiologus thoracicus Handlirsch 1906
G. †Cantabrala Kukalová-Peck and Brauckmann 1992
+
Cantabrala gandli Kukalová-Peck and Brauckmann 1992
G. †Genentomum Scudder 1885
+
Genentomum subacutum Bolton 1911
Genentomum validum Scudder 1885
+
Invalid names: Genentomum carri Cockerell 1917 [synonym]
G. †Gerarulus Handlirsch 1911
+
Gerarulus radialis Handlirsch 1911
G. †Gerarus Scudder 1885
+
Gerarus collaris Handlirsch 1911
+
Invalid names: Gerarus longicollis Handlirsch 1911 [synonym]
Gerarus constrictus Scudder 1885
+
Invalid names: Gerarus angustus Handlirsch 1906 [synonym], Gerarus latus Handlirsch 1911 [synonym], Gerarus longus Handlirsch 1906 [synonym], Gerarus reductus Handlirsch 1911 [synonym], Rossites inopinus Richardson 1956 [synonym]
Gerarus danielsi Handlirsch 1906
Gerarus fischeri Brongniart 1885
+
Invalid names: Archaeacridites bruesi Meunier 1909 [synonym], Archaeacridites elegantissima Meunier 1909 [synonym], Sthenaropoda agnusi Lameere 1917 [synonym], Sthenaropoda lerichei Lameere 1917 [synonym]
Gerarus mazonus Scudder 1885
Gerarus symmetricus Bolton 1934
Gerarus teutonicus Brauckmann and Herd 2005
Gerarus vetus Scudder 1885
Invalid names: Archaeacridites Meunier 1909 [synonym], Genopteryx Scudder 1885 [synonym], Rossites Richardson 1956 [synonym], Sthenaropoda Brongniart 1885 [synonym]
G. †Hirsutgerarus Zessin 2012
+
Hirsutgerarus saxonicus Zessin 2012
G. †Nacekomia Richardson 1956
+
Nacekomia rossae Richardson 1956
G. †Osnogerarus Kukalová-Peck and Brauckmann 1992
+
Osnogerarus trecwithiensis Kukalová-Peck and Brauckmann 1992
G. †Owadpteron Dvořák et al. 2019
+
Owadpteron dareki Dvořák et al. 2019
G. †Parapalaeomastax Dvořák et al. 2019
+
Parapalaeomastax dariuszi Dvořák et al. 2019
G. †Ploetzgerarus Zessin 2009
+
Ploetzgerarus krempieni Zessin 2009
G. †Progenentomum Handlirsch 1906
+
Progenentomum carbonis Handlirsch 1906
Invalid names: Genopterygidae Richardson 1956 [synonym], Sthenaropodidae Handlirsch 1906 [synonym]
Fm. †Permostridulidae Béthoux et al. 2003
+
G. †Permostridulus Béthoux et al. 2003
+
Permostridulus brongniarti Béthoux et al. 2003
G. †Sinogerarus Gu et al. 2017
+
Sinogerarus pectinatus Gu et al. 2017
G. †Phtanomiamia Chen et al. 2020
+
Phtanomiamia gui Chen et al. 2020
G. †Protodictyon Melander 1903
+
Protodictyon pulchripenne Melander 1903
G. †Protomiamia Du et al. 2016
+
Protomiamia yangi Du et al. 2016
G. †Sumixam Béthoux and Grossmann 2011
+
Sumixam maximus Béthoux and Grossmann 2011
Invalid names: Lobeattida Béthoux 2005 [empty]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Kiessling 2004, Bush and Bambach 2015

Age range

Maximum range based only on fossils: base of the Marsdenian to the top of the Middle Cenomanian or 321.30000 to 93.50000 Ma
Minimum age of oldest fossil (stem group age): 320.0 Ma

Collections (191 total)


Time interval Ma Country or state Original ID and collection number
Marsdenian321.3 - 320.0Germany (Ruhr) Heterologopsis ruhrensis (155261)
Westphalian A318.7 - 316.9France (Nord-Pas-de-Calais) Omalia macroptera (152785)
Westphalian A318.7 - 316.9Belgium Omalia macroptera (123821)
Westphalian A318.7 - 316.9Poland Owadpteron dareki, Parapalaeomastax dariuszi (107805)
Westphalian A318.7 - 316.9United Kingdom (England) Genentomum subacutum (127291)
Westphalian A318.7 - 316.9France Oudardgramma bruayensis (122506)
Westphalian A318.7 - 316.9Canada (New Brunswick) Geroneura wilsoni (114468)
Westphalian A318.7 - 316.9Canada (Nova Scotia) Caloneurodea indet. (216802)
Westphalian B316.9 - 314.6United Kingdom (England) Coselia palmiformis (125238)
Westphalian B316.9 - 314.6Poland Omaliella polonica (206852)
Westphalian B316.9 - 314.6France Omalia palmiformis (151373)
Westphalian B316.9 - 314.6United Kingdom (Staffordshire) Geroneura ovata (34906)
Westphalian B316.9 - 314.6United Kingdom (Wales) Gerarus symmetricus, Hapaloptera obovata (151398) Hapaloptera parallela (151404)
Westphalian B - Westphalian C316.9 - 309.8China (Ningxia) Chenxiella liuae, Longzhua loculata, Sinopteron huangheense, Miamia maimai, Heterologus duyiwuer, Xixia huban, Protomiamia yangi, Sinogerarus pectinatus, Phtanomiamia gui, Ctenoptilus frequens (139576)
Moscovian315.2 - 307.0USA (Maryland) Narkema alternatum, Spaniodera simplex (124496)
Westphalian C314.6 - 309.8China (Ningxia) Longzhua loculata, Sinopteron huangheense, Miamia maimai (179078) Miamia maimai (179076 179077)
Westphalian C314.6 - 309.8Belgium Palaeomastax carbonis (123820)
Westphalian C314.6 - 309.8France (Nord Pas de Calais) Anthraconeura silvatica (128557)
Westphalian C - Westphalian D314.6 - 307.5France (Nord Pas de Calais) Archaeorthoptera indet., Bruaylogus magnificus (153462)
Westphalian C - Westphalian D314.6 - 307.5USA (Pennsylvania) Hapaloptera gracilis (114456)
Westphalian C - Westphalian D314.6 - 307.5France (Nord-Pas-de-Calais) Emphyloptera lecrivaini (151691)
Westphalian C - Westphalian D314.6 - 307.5Germany (Lower Saxony) Piesbergopterum punctatum (122514)
Westphalian C - Westphalian D314.6 - 307.5France Duquesnia gallica (224002) Tococladus sp., Aviogramma gracilis, Aviocladus pectinatus, Aviohapaloptera bethouxi, Aviologus duquesnei, Cacurgus avionensis, Avionugonioneura jouaulti, Beloatta duquesnei, Aviobiella garroustei, Piesbergopterum avionensis (122504)
Westphalian D309.8 - 307.5USA (Indiana) Sumixam maximus (110852)
Westphalian D309.8 - 307.5USA (Pennsylvania) Amboneura klosei (122502) Caloneurella carbonaria (124504) Caloneurodea indet. (114649) Dieconeura rigida (111045)
Westphalian D309.8 - 307.5USA (Illinois) Aenigmatodes danielsi, Gerarus danielsi, Progenentomum carbonis (146857) Cheliphlebia extensa (122521) Dictyoneura clarinervis (122522) Genentomum carri (122527) Gerarus collaris, Spaniodera ambulans, Metryia analis, Gerapompus blattinoides, Gerapompus extensus, Genopteryx constricta, Genentomum validum, Dieconeura arcuata, Aethophlebia singularis, Gerarus vetus, Gerarus mazonus, Gerarus longus, Gerarus angustus, Gerapompus schucherti, Axiologus thoracicus, Endoiasmus reticulatus (21981) Gerarus danielsi (122512) Gerarus vetus (151855) Gerarus vetus, Gerarus danielsi (151848 151849) Gerarus vetus, Gerarus danielsi, Nacekomia rossae, Anthrakoris aetherius (112638) Heterologus langfordorum (133734) Lobeatta schneideri (112637) Miamia bronsoni (151543) Narkema taeniatum, Anegertus cubitalis, Spaniodera longicollis, Spaniodera lata, Spaniodera elatior, Spaniodera schucherti, Spaniodera acutipennis, Spaniodera parvula, Spaniodera angusta, Dieconeura mazona, Gerarus latus, Gerarus collaris, Gerarus reductus, Gerarus longicollis, Gerarulus radialis, Anepitedius giraffa, Cacurgus spilopterus, Spilomastax oligoneurus (112630) Propteticus infernus (114476) Protodictyon pulchripenne (122523) Rossites inopinus (122530 122531) Termes contusus, Petromartus indistinctus (122524)
Westphalian D309.8 - 307.5France (Nord-Pas-de-Calais) Cymenophlebia carpentieri (107403) Euthyneura lecomtei (124902)
Westphalian D309.8 - 307.5United Kingdom (Somerset) Gerarus danielsi, Etotabla gillae (156594) Miamia rossorum (156595)
Westphalian D309.8 - 307.5United Kingdom (England) Narkeminopsis eddi (105765)
Westphalian D309.8 - 307.5USA (Missouri) Narkemina winsdoriensis (151845)
Westphalian D309.8 - 307.5Germany (Lower Saxony) Caloneurodea indet., Geraridae indet., Spiculum mendicum, Narkeminopsis sowiaki (122515) Forfexala kiarae (112605 112606) Gerarus teutonicus, Primakollaria amicorum, Contracladus impar (122518) Osnogerarus trecwithiensis (126589) Paraspiculum brauckmanni (224159)
Cantabrian307.5 - 305.1Spain Cantabrala gandli (126590)
Kasimovian307.0 - 303.7Ukraine (Voroshilovgrad) Lomovatka udovichenkoi (123534)
Kasimovian307.0 - 303.7USA (New Mexico) Pseudobiella fasciata (49663)
Kasimovian - Gzhelian307.0 - 298.9Russian Federation (Khaskassia) Narkemulla sibirica, Izykhia tridentis, Fodinopenna abakanica (145701)
Kasimovian - Gzhelian307.0 - 298.9Brazil (Santa Catarina) Carpenteroptera onzii (181409)
Kasimovian - Gzhelian307.0 - 298.9Russian Federation Narkemina angustiformis, Narkemina kata, Narkeminopsis inversa, Narkemulla sibirica, Evenkiophlebia collucata, Carbonokata storozhenkoi, Tshunoptera ampla (102503)
Kasimovian - Gzhelian307.0 - 298.9Argentina (San Luis) Paranarkemina velizensis, Paranarkemina kurtzi, Paranarkemina amosi (35057) Velizphlebia cruzi (127043)
Stephanian A305.1 - 301.5France Roqueia alpina (229588)
Gzhelian303.7 - 298.9France Caloneura dawsoni, Eoblatta robusta, Caloneura subtilis, Eoblattina complexa (124916) Gerarus fischeri, Sthenaropoda lerichei, Archaeacridites bruesi, Sthenarocera pachytyloides, Homaloneura similis, Homaloneura royeri, Ctenoptilus elongatus, Homalophlebia trouessarti, Ischnoptera diaphanes, Bouleites latipennis, Archaeacridites elegantissima, Sthenaropoda fischeri (121979) Sthenaropoda fischeri, Sthenaropoda lerichei, Archaeacridites bruesi, Leptoneura oustaleti, Protophasma gaudryi, Protophasma woodwardii, Protodiamphipnoa tertrini, Protokollaria ingens, Sthenarocera pachytyloides, Sthenarocera bureaui, Caloneura picta, Caloneura major, Caloneura longicornis, Caloneura robusta, Ischnoneura delicatula, Ischnoneura elongata, Stenoneura robusta, Ischnoneura brongniarti, Caloneura dawsonii (103635)
Gzhelian303.7 - 298.9Brazil (Rio Grande do Sul) Narkemina rochacamposi (105039)
Gzhelian303.7 - 298.9Brazil (São Paulo) Narkemina sp. A (117288) Narkemina sp. B, Narkemina rodendorfi (117285) Narkemina sp. C (117286)
Gzhelian303.7 - 298.9Russian Federation Aenigmatella comparabilis (124585) Narkemina angustata (124587) Narkemina angustiformis (124586) Narkemina genuina (124583)
Stephanian B301.5 - 301.0France Amphiboliacridites joanesi (172760) Apsidoneura sottyi (122490) Eoblatta zeilleri (172759)
Stephanian C301.0 - 300.4Germany (Saxony-Anhalt) Laspeyresia wettinensis (122578)
Stephanian C301.0 - 300.4Germany Hirsutgerarus saxonicus (163476) Ploetzgerarus krempieni (122505)
Stephanian C301.0 - 300.4Portugal Lusitaneura covensis (103240)
Asselian298.9 - 295.5USA (New Mexico) Paleuthygramma ventricosa, Carrizarroyo calopterus (124386) Vilviopsis picta (124385) Vilviopsis picta, Carrizopteron longum, Arroyopteron indolatum (124387)
Asselian298.9 - 295.5Czech Republic Cerasopterum sp., Cerasopterum gracile, Ventopterum rapidum (114186)
Asselian298.9 - 295.5Germany Palatinarkema prokopi (112604) Panorthoptera indet. (222678)
Artinskian290.1 - 279.3USA (Kansas) Aibolitus minutus, Elmopterum rotundum, Protembia permiana, Paleuthygramma acuta, Apsidoneura flexa, Plesiogramma medialis (111319) Anomalogramma parva (114151) Caloneurodea indet., Permobiella perspicua, Protembia permiana, Paleuthygramma acuta, Apsidoneura flexa, Tococladus rallus (111320) Caloneurodea indet., Pryg absurdus (114450) Nugonioneura problematica, Permobiella perspicua, Protembia permiana, Telactinopteryx striatipennis, Paleuthygramma acuta (111219) Nugonioneura problematica, Plesiogramma reducta, Opisthocladus arcuatus, Opisthocladus strictus, Psoroptera cubitalia, Heteroptilon costale (111217) Paleuthygramma acuta (114152)
Artinskian290.1 - 279.3France (Languedoc) Paleuthygramma acuta (192422) Panorthoptera indet. (174411) Tococladus sp., Nanogramma gandi, Jarmiloptera mouralensis (105044)
Artinskian290.1 - 279.3France Jarmiloptera mouralensis (174409) Lodevolongzhua incompleta (174414) Paralongzhua elongata (117228) Tococladus garrici (122935)
Artinskian290.1 - 279.3USA (Oklahoma) Apsidoneura reducta (104826) Ligogramma wichita (104821) Paleuthygramma acuta, Apsidoneura flexa, Anomalogramma parva, Gigagramma carpenteri, Ligogramma sinuosa (107405) Sigmophlebia engeli (104828)
Artinskian290.1 - 279.3Madagascar Mecopterida indet. (117289)
Artinskian290.1 - 279.3Russian Federation (Perm) Vilvia densinervosa (153669)
Artinskian - Wordian290.1 - 264.28France (Languedoc) Gallogramma galadrieli (192423)
Kungurian279.3 - 272.3France Tococladus garrici, Bethouxia ariegensis, Lodevogramma pumilia, Caloneurodea indet. (103637)
Kungurian279.3 - 272.3Russian Federation Aibolitus medicinus, Sylvabestia tenuis, Sojanoraphidia martynovae, Tshekardomina maculosa, Tillyardembia ravisedorum, Pleisiogramma maueri (107787) Aibolitus medicinus, Tshekardembia sharovi, Sylvardembia tamaena, Barmaleus dentatus, Sojanoraphidia martynovae (114207) Kishertia tricubitalis (122754) Tillyardembia antennaeplana, Tillyardembia biarmica (102504) Tillyardembia biarmica (155825) Tillyardembia biarmica, Tillyardembia ravisedorum (155819)
Kungurian279.3 - 272.3Russian Federation (Suksun) Aibolitus medicinus, Sylvabestia tenuis, Tillyardembia sp., Tillyardembia antennaeplana, Kungurembia brevicervix, Kungurembia pallida, Sylvardembia matura, Paratillyardembia sepicolorata, Sojanoraphidia martynovae, Tshekardomina imbecillissima, Tshekardomina subincurvata, Tshekardomina maculosa, Neprotembia truncata, Neraphidia mitis, Neraphidia rigida, Sojanopermula tshekardensis, Sojanopermula minor, Permedax effertus, Tillyardembia ravisedorum, Tococladus similis, Tshekardomina imbecilla, Tshecalculus inaspectus, Paleuthygramma tenuicorne, Parmaptera permiana, Tshekardocladus sparsus (108206) Paleuthygramma tenuicornis (157042) Permocapnia brevipes (113379 116641 116642) Tillyardembia sp., Tillyardembia antennaeplana, Kungurembia brevicervix, Kungurembia pallida, Tshekardomina imbecillissima, Neraphidia mitis, Tshekardomina imbecilla (122753)
Kungurian279.3 - 272.3France (Languedoc) Permostridulus brongniarti (104990 105053 122920)
Kungurian279.3 - 272.3Russian Federation (Kemerovo) Narkeminuta permiana (145702)
Kungurian279.3 - 272.3Russian Federation (Perm) Pinidelia sukatshevae, Kamamica promota (103639)
Kungurian279.3 - 272.3Russian Federation (Komi Republic) Neprotembia complicata (127047)
Roadian272.3 - 268.8Russian Federation (Kazan) Euthygramma curvatum, Permarrhaphus venosus (116803)
Roadian272.3 - 268.8Russian Federation Euthygramma parallellum (116935) Permula edemskii, Sojanopermula lucida (96860)
Roadian272.3 - 268.8Russian Federation (Kirov) Kityakia attrita, Kirovopteron luteus (105058)
Roadian272.3 - 268.8Russian Federation (Kemerovo) Pleisiogramma kemerovoense (124605) Sibestia nana (104704)
Roadian272.3 - 268.8Russian Federation (Arkangelsk) Euthygramma aberrans (104993) Euthygramma parallellum (116535 116933) Euthygramma parallellum, Sojanopermula rasnitsyni, Kazanalicula reducta, Sojanoraphidia rossica, Idelopterum samoedum, Tshekardomina novokshonovi (104716) Euthygramma parallellum, Vilviopsis extensa (116934) Kungurembia biarmica, Paleuthygramma sharovi, Euthygramma parallelum, Repka stramenis, Repka repens, Repka curta, Soyanocadaver crypticus (104992)
Wordian266.9 - 264.28Russian Federation (Khaskassia) Yaria arenaria (161991)
Wordian266.9 - 264.28Russian Federation (Udmurtia) Tshepanichoptera lacera, Sigmophlebia udmurtica (103502) Udembia udmurtica, Votyak pictus (104750)
Capitanian264.28 - 259.51China (Anhui) Sinaspidoneura magnifica (103239)
Capitanian264.28 - 259.51Russian Federation (Orenburg) Eohymen maculipennis (116607) Kuplya minutissima (104717) Opisthocladus kargalensis (104057)
Wuchiapingian259.9 - 254.17Mongolia Tshekardomina mongolica, Micropermula mongolica, Micropermula ceceya (97602)
Wuchiapingian259.9 - 254.17Russian Federation (Vologda) Mezenalicula conjuncta, Isadistica issada, Isadistica longa, Issatermes parvus (104754)
Changhsingian254.17 - 252.17Russian Federation (Vologda) Vokhmia pyriformis, Vokhmia laxa (105046)
Changhsingian254.17 - 252.17Kazakhstan (Zaisan) Sigmophlebia rugulosa (105062)
Changhsingian254.17 - 252.17Russian Federation (Kirov) Tillyardembia zalazna (140985)
Bithynian247.2 - 242.0Germany (Bavaria) Hammelburgia selli (115970)
Ladinian242.0 - 237.0Kyrgyzstan (Osh) Aiban kichineis (163986) Ferganamadygenia plicata (106088) Sharovala triassica (105007)
Callovian - Oxfordian166.1 - 157.3China (Nei Mongol) Jurachresmoda gaskelli (112664) Jurachresmoda sanyica (113362) Juraperla daohugouensis (113371) Juraperla grandis (113365)
Callovian - Oxfordian166.1 - 157.3Kazakhstan (Chimkent) Necrophasma shabarovi (31311)
Early/Lower Tithonian150.8 - 145.0Germany (Bayern) Chresmoda obscura (123144 133578) Chresmoda obscura, Pygolampis gigantea, Halometra minor, Sternarthron zitteli (123985) Pygolampis gigantea, Propygolampis bronni (123906)
Early/Lower Tithonian150.8 - 145.0Germany (Bavaria) Chresmoda obscura (122586 123994 139876)
Berriasian145.0 - 139.8United Kingdom (England) Gryllidium oweni (127481)
Late/Upper Hauterivian136.4 - 130.0China (Hebei) Chresmoda multinervis, Chresmoda shihi (113484)
Early/Lower Barremian130.0 - 125.45Spain Chresmoda aquatica (104052)
Late/Upper Barremian130.0 - 125.45China (Nei Mongol) Sinochresmoda magnicornia (125656)
Late/Upper Barremian130.0 - 125.45China (Liaoning) Chresmoda orientalis (140222)
Late/Upper Barremian130.0 - 125.45Spain (Castilla-La Mancha) Chresmoda aquatica (28006)
Late/Upper Barremian - Early/Lower Aptian130.0 - 122.46China (Liaoning) Chresmoda sp. (73687)
Aptian125.0 - 113.0Mongolia (Govi-Altai) Saurophthiroides mongolicus (136020)
Aptian125.0 - 113.0China (Jilin) Sinochresmoda magnicornia (196770)
Late/Upper Aptian122.46 - 113.0Brazil (Ceará) Chresmoda neotropica (122352 122557 122919 127034)
Early/Lower Cenomanian99.6 - 93.5Myanmar Chresmoda chikuni (178121)
Middle Cenomanian99.6 - 93.5Lebanon Chresmoda libanica (152969)