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Pterodactyloidea

Reptilia - Pterosauria

Taxonomy
Pterodactyloidea was named by Plieninger (1901). It was considered monophyletic by Unwin (2003) and Pereda-Suberbiola et al. (2012) and Martill et al. (2013).

It was synonymized subjectively with Ornithocephaliformes by Hay (1930); it was reranked as the unranked clade Pterodactyloidea by Unwin (2003), Osi et al. (2005), Naish et al. (2013), Andres and Myers (2013), Rodrigues and Kellner (2013), Vidovic and Martill (2014), Bantim et al. (2014), Pegas et al. (2016), Zhou et al. (2017), Jacobs et al. (2019), Zhou et al. (2019), Kellner et al. (2019), Holgado et al. (2019), Holgado and Pegas (2020), Pegas et al. (2020), Richards et al. (2023), Martill et al. (2023) and Hone et al. (2023).

It was assigned to Breviquartossa by Unwin (2003); to Caelidracones by Andres et al. (2010) and Andres et al. (2014); to Monofenestrata by Martill et al. (2013), Vidovic and Martill (2014) and Martill et al. (2023); to Pterosauria by Roxo (1937), Kuhn (1946), Huene (1954), Delair (1958), Young (1964), Kuhn (1966), Charig (1967), Haubold (1971), Gasparini et al. (1987), Carroll (1988), Sereno (1991), Kellner and Campos (2002), Chure et al. (2006), Wang and Zhou (2006), Andres and Ji (2006), Pereda-Suberbiola et al. (2012), Jiang et al. (2014) and Agnolín et al. (2023); and to Pterosauria by Osi et al. (2005), Naish et al. (2013), Andres and Myers (2013), Rodrigues and Kellner (2013), Headden and Campos (2014), Bantim et al. (2014), Pegas et al. (2016), Zhou et al. (2017), O'Sullivan and Martill (2018), Jacobs et al. (2019), Zhou et al. (2019), Kellner et al. (2019), Holgado et al. (2019), Holgado and Pegas (2020), Pegas et al. (2020), Richards et al. (2023), Hone et al. (2023) and Pentland et al. (2024).

Subtaxa
Agadirichnidae, Araripedactylus, Araripedactylus dehmi, Araripesaurus castilhoi, Archaeopterodactyloidea, Ardeadactylus, Aurorazhdarchidae, Azhdarchoidea, Belonochasmatidae, Cearadactylidae, Dermodactylus montanus, Doratorhynchus validus, Dsungaripteroidea, Eupterodactyloidea, Haenamichnus, Herbstosaurus, Kryptodrakon, Lonchodraconidae, Lophocratia, Mesadactylus, Ningchengopterus, Nyctosaurus (syn. Nyctodactylus), Ornithocheiridae (syn. Criorhynchidae), Ornithocheirus brachyrhinus, Ornithocheirus bunzeli, Ornithocheirus carteri, Ornithocheirus crassidens, Ornithocheirus dentatus, Ornithocheirus enchorhynchus, Ornithocheirus eurygnathus, Ornithocheirus oxyrhinus, Ornithocheirus scaphorhynchus, Ornithocheirus tenuirostris, Ornithocheirus xyphorhynchus, Ornithodesmidae, Osteornis, Pteranodontia, Pteranodontoidea, Pterodactylus grandipelvis, Pterodactylus grandis, Pterodactylus longipes, Pterodactylus woodwardi, Santanadactylus pricei, Saraikisauridae, Tendaguripteridae, Titanopterygiidae, Wenupteryx
Sister taxa
Archaeopterodactyloidea, Azhdarchoidea, Dsungaripteroidea, Pteranodontoidea

Synonyms
Synonymy list
YearName and author
1824Pterodactylus grandis Cuvier
1829Pterodactylus grandis Holl p. 82
1842Pterodactylus grandis Owen p. 156
1858Pterodactylus grandis Buckland p. 218
1859Pterodactylus longipes Münster
1859Pterodactylus grandis Owen p. 151–152
1860Pterodactylus grandipelvis Meyer
1861Pterodactylus woodwardi Owen p. 4
1861Pterodactylus grandis Owen p. 274
1865Pterodactylus carteri Seeley
1869Ornithocheirus carteri Seeley
1869Ptenodactylus woodwardi Seeley
1870Pterodactylus validus Owen
1870Ornithocheirus tenuirostris Seeley
1870Ornithocheirus woodwardi Seeley
1870Pterodactylus carteri Seeley
1870Ornithocheirus oxyrhinus Seeley p. 117
1870Ornithocheirus xyphorhynchus Seeley p. 117
1870Ornithocheirus dentatus Seeley p. 119
1870Ornithocheirus scaphorhynchus Seeley p. 119
1870Ornithocheirus crassidens Seeley p. 122
1870Ornithocheirus brachyrhinus Seeley p. 123
1870Ornithocheirus enchorhynchus Seeley p. 123
1870Ornithocheirus eurygnathus Seeley p. 123
1875Doratorhynchus validus Seeley
1878Pterodactylus montanus Marsh p. 233
1878Pterodactylus montanus Williston p. 44
1881Dermodactylus montanus Marsh p. 342
1881Ornithocheirus bunzeli Seeley p. 701
1886Pterodactylus grandipelvis Winkler p. 200
1890Pterodactylus grandipelvis Zittel p. 792
1890Pterodactylus longipes Zittel p. 792
1890Dermodactylus montanus Zittel p. 798
1890Doratorhynchus validus Zittel p. 798
1892Pterodactylus grandis Lortet p. 132
1897Dermodactylus montanus Marsh p. 508
1900Doratorhynchus validum Van Den Broeck p. 106
1901Pterodactyloidea Plieninger
1902Dermodactylus montanus Hay p. 507
1930Dermodactylus montanus Hay p. 161
1937Pterodactyloidea Roxo p. 44
1946Pterodactyloidea Kuhn p. 67
1954Pterodactyloidea Huene p. 56
1958Pterodactyloidea Delair p. 69
1958Doratorhynchus validus Delair p. 71
1964Pterodactyloidea Young p. 251
1966Pterodactyloidea Kuhn p. 109
1967Pterodactyloidea Charig p. 717
1970Doratorhynchus validus Macfadyen p. 143
1971Pterodactyloidea Haubold p. 89
1971Araripesaurus castilhoi Price p. 454
1977Araripedactylus Wellnhofer
1977Araripedactylus dehmi Wellnhofer p. 160
1981Dermodactylus montanus Galton p. 1120
1985Santanadactylus pricei Wellnhofer p. 132 figs. 19-24
1987Pterodactyloidea Gasparini et al. p. 141
1988Araripedactylus Carroll
1988Pterodactyloidea Carroll
1991Pterodactyloidea Sereno p. 19
2002Pterodactyloidea Kellner and Campos p. 389
2003Pterodactyloidea Unwin p. 158
2005Pterodactyloidea Osi et al.
2006Pterodactyloidea Andres and Ji p. 70
2006Dermodactylus montanus Chure et al. p. 236
2006Pterodactyloidea Chure et al. p. 236
2006Pterodactyloidea Wang and Zhou p. 406
2010Pterodactyloidea Andres et al. fig. 7
2012Pterodactyloidea Pereda-Suberbiola et al.
2013Pterodactyloidea Andres and Myers
2013Pterodactyloidea Martill et al.
2013Pterodactyloidea Naish et al.
2013Osteornis Rodrigues and Kellner
2013Pterodactyloidea Rodrigues and Kellner p. 3
2014Pterodactyloidea Andres et al.
2014Pterodactyloidea Bantim et al.
2014Pterodactyloidea Headden and Campos
2014Pterodactyloidea Jiang et al. p. 823
2014Pterodactyloidea Vidovic and Martill
2016Pterodactyloidea Pegas et al.
2017Pterodactyloidea Zhou et al.
2018Pterodactyloidea O'Sullivan and Martill
2019Pterodactyloidea Holgado et al.
2019Pterodactyloidea Jacobs et al.
2019Pterodactyloidea Kellner et al.
2019Pterodactyloidea Zhou et al.
2020Pterodactyloidea Holgado and Pegas
2020Pterodactyloidea Pegas et al.
2023Pterodactyloidea Agnolín et al. p. 310
2023Pterodactyloidea Hone et al. p. 3
2023Pterodactyloidea Martill et al. p. 393
2023Pterodactyloidea Richards et al. p. 5
2024Pterodactyloidea Pentland et al. p. 2

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
—Osteichthyes()
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
—Tetrapoda
—Reptiliomorpha
—Anthracosauria
subclassAmphibiosauriaKuhn 1967
—Cotylosauria()
—Amniota
RankNameAuthor
—Sauropsida
classReptilia
subclassEureptilia()
—Romeriida
—Diapsida()
—Archosauromorpha(Huene 1946)
—Crocopoda
—ArchosauriformesGauthier 1986
—Eucrocopoda
—Archosauria()
informalAvemetatarsalia
—Ornithodira
—PterosauromorphaPadian 1997
orderPterosauriaKaup 1834
suborderPterodactyloideaPlieninger 1901
suborderPterodactyloideaPlieninger 1901

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Subor. †Pterodactyloidea Plieninger 1901
show all | hide all
Fm. †Agadirichnidae Masrour et al. 2018
+
G. †Agadirichnus Ambroggi and Lapparent 1954
+
†Agadirichnus elegans Ambroggi and Lapparent 1954
Superfm. †Archaeopterodactyloidea Kellner 1996
+
Unr. †Boreopteridae Lü et al. 2006
+
G. †Boreopterus Lü and Ji 2005
+
†Boreopterus cuiae Lü and Ji 2005
†Boreopterus giganticus Jiang et al. 2014
Subfm. †Moganopterinae Lü et al. 2012
+
G. †Moganopterus Lü et al. 2012
+
†Moganopterus zhuiana Lü et al. 2012
G. †Zhenyuanopterus Lü 2010
+
Invalid names: Boreopterinae Lü et al. 2012 [empty]
G. †Ardeadactylus Bennett 2013
+
+
Invalid names: Pterodactylus vulturinus Wagner 1858 [synonym]
Fm. †Aurorazhdarchidae Vidovic and Martill 2014
+
G. †Aerodactylus Vidovic and Martill 2014
Superfm. †Azhdarchoidea Unwin 1992
+
Fm. †Azhdarchidae Nessov 1984
+
G. †Aerotitan Novas et al. 2012
+
†Aerotitan sudamericanus Novas et al. 2012
G. †Alanqa Ibrahim et al. 2010
+
†Alanqa saharica Ibrahim et al. 2010
G. †Albadraco Solomon et al. 2020
+
†Albadraco tharmisensis Solomon et al. 2020
G. †Aralazhdarcho Averianov 2007
+
†Aralazhdarcho bostobensis Averianov 2007
G. †Arambourgiania Nessov et al. 1987
+
†Arambourgiania philadelphiae Arambourg 1959
G. †Azhdarcho Nessov 1981
+
†Azhdarcho lancicollis Nessov 1984
+
Invalid names: Azhdarcho imparidens Nessov 1981 [replaced]
G. †Bakonydraco Osi et al. 2005
+
†Bakonydraco galaczi Osi et al. 2005
G. †Bennettazhia Nessov 1991
+
†Bennettazhia oregonensis Gilmore 1928
G. †Bogolubovia Nessov and Yarkov 1989
+
†Bogolubovia orientalis Bogoliubow 1914
G. †Chaoyangopterus Wang and Zhou 2003
+
†Chaoyangopterus zhangi Wang and Zhou 2003
G. †Cryodrakon Hone et al. 2019
+
†Cryodrakon boreas Hone et al. 2019
G. †Doratorhynchus Seeley 1875
G. †Eoazhdarcho Lü and Ji 2005
+
†Eoazhdarcho liaoxiensis Lü and Ji 2005
G. †Eurazhdarcho Vremir et al. 2013
+
†Eurazhdarcho langendorfensis Vremir et al. 2013
G. †Hatzegopteryx Buffetaut et al. 2002
+
†Hatzegopteryx thambema Buffetaut et al. 2002
G. †Mistralazhdarcho Vullo et al. 2018
+
†Mistralazhdarcho maggii Vullo et al. 2018
G. †Montanazhdarcho Padian et al. 1995
+
†Montanazhdarcho minor Padian et al. 1995
G. †Navajodactylus Sullivan and Fowler 2011
+
†Navajodactylus boerei Sullivan and Fowler 2011
G. †Phosphatodraco Pereda Suberbiola et al. 2003
+
†Phosphatodraco mauritanicus Pereda Suberbiola et al. 2003
Subfm. †Quetzalcoatlinae Andres et al. 2014
+
G. †Quetzalcoatlus Lawson 1975
+
†Quetzalcoatlus lawsoni Andres and Langston Jr 2021
†Quetzalcoatlus northropi Lawson 1975
G. †Thanatosdrakon Ortiz David et al. 2022
+
†Thanatosdrakon amaru Ortiz David et al. 2022
G. †Wellnhopterus Andres and Langston Jr 2021
+
†Wellnhopterus brevirostris Andres and Langston Jr 2021
G. †Titanopteryx Arambourg 1959
G. †Zhejiangopterus Cai and Wei 1994
+
†Zhejiangopterus linhaiensis Cai and Wei 1994
G. †Cretornis Fritsch 1881
+
†Cretornis hlavaci Fritsch 1881
Fm. †Dsungaripteridae Young 1964
+
G. †Banguela Headden and Campos 2014
+
†Banguela oberlii Headden and Campos 2014
G. †Domeykodactylus Martill et al. 2000
+
†Domeykodactylus ceciliae Martill et al. 2000
G. †Dsungaripterus Young 1964
+
†Dsungaripterus weii Young 1964
G. †Lonchognathosaurus Maisch et al. 2004
+
†Lonchognathosaurus acutirostris Maisch et al. 2004
G. †Noripterus Young 1973
+
†Noripterus parvus Bakhurina 1982
Invalid names: Phobetor Bakhurina 1986 [synonym]
G. †Eopteranodon Lü and Zhang 2005
+
†Eopteranodon lii Lü and Zhang 2005
G. †Europejara Vullo et al. 2012
+
†Europejara olcadesorum Vullo et al. 2012
G. †Leptostomia Smith et al. 2020
+
†Leptostomia begaaensis Smith et al. 2020
G. †Microtuban Elgin and Frey 2011
+
†Microtuban altivolans Elgin and Frey 2011
G. †Nemicolopterus Wang et al. 2008 [forest dweller wing]
+
†Nemicolopterus crypticus Wang et al. 2008
Unr. †Neoazhdarchia Unwin 2003
+
Fm. †Chaoyangopteridae Lü et al. 2008
+
G. †Apatorhamphus McPhee et al. 2020
+
†Apatorhamphus gyrostega McPhee et al. 2020
G. †Lacusovagus Witton 2008
+
†Lacusovagus magnificens Witton 2008
G. †Shenzhoupterus Lü et al. 2008
+
†Shenzhoupterus chaoyangensis Lü et al. 2008
G. †Radiodactylus Andres and Myers 2013
+
†Radiodactylus langstoni Andres and Myers 2013
Superfm. †Tapejaroidea Kellner 1989
+
Fm. †Tapejaridae Kellner 1989
+
G. †Afrotapejara Martill et al. 2020
Unr. †Chaoyangopterinae Pinheiro et al. 2011
G. †Huaxiapterus Lü and Yuan 2005
G. †Ingridia Unwin and Martill 2007
G. †Sinopterus Wang and Zhou 2002
G. †Tapejara Kellner 1989
Subfm. †Tapejarinae Kellner 1989
Subfm. †Thalassodrominae Kellner and Campos 2007
G. †Tupandactylus Kellner and Campos 2007
G. †Wightia Martill et al. 2020
Fm. †Thalassodromidae Kellner and Campos 2007
+
G. †Thalassodromeus Kellner and Campos 2002
G. †Tupuxuara Kellner and Campos 1988
G. †Xericeps Martill et al. 2018
+
†Xericeps curvirostris Martill et al. 2018
G. †Ornithostoma Seeley 1871
+
†Ornithostoma sedgwicki Seeley 1871
+
Invalid names: Ornithostoma seeleyi Lydekker 1904 [synonym]
Fm. †Protazhdarchidae Frey et al. 2011
+
G. †Aurorazhdarcho Frey et al. 2011
+
†Aurorazhdarcho primordius Frey et al. 2011
Unr. †Tapejaromorpha Andres et al. 2014
+
G. †Keresdrakon Kellner et al. 2019
+
†Keresdrakon vilsoni Kellner et al. 2019
G. †Vectidraco Naish et al. 2013
+
†Vectidraco daisymorrisae Naish et al. 2013
Invalid names: Palaeornis cliftii Mantell 1844 [nomen dubium]
Superfm. †Dsungaripteroidea Young 1964
+
G. †Puntanipterus Bonaparte and Sánchez 1975
Invalid names: Germanodactylidae Young 1964 [invalid subgroup], Nyctosauridae Hay 1902 [empty]
G. †Haenamichnus Hwang et al. 2002
+
†Haenamichnus uhangriensis Hwang et al. 2002
G. †Herbstosaurus Casamiquela 1975 [Herbst lizard]
+
†Herbstosaurus pigmaeus Casamiquela 1975
G. †Kryptodrakon Andres et al. 2014
+
†Kryptodrakon progenitor Andres et al. 2014
Fm. †Lonchodraconidae Rodrigues and Kellner 2013
+
G. †Lonchodraco Rodrigues and Kellner 2013
+
†Lonchodraco giganteus Bowerbank 1846
†Lonchodraco microdon Seeley 1869
Unr. †Lophocratia Unwin 2003
+
G. †Beipiaopterus Lü 2003
+
G. †Ctenochasma Meyer 1851
+
†Ctenochasma elegans Wagner 1861
+
Invalid names: Ctenochasma gracile Oppel 1862 [synonym], Pterodactylus pulchellus Meyer 1859 [synonym]
†Ctenochasma porocristata de Buisonje 1981
†Ctenochasma roemeri Meyer 1851
†Ctenochasma taqueti Bennett 2007
Superfm. †Ctenochasmatoidea Unwin 1995
+
Unr. †Euctenochasmatia Unwin 2003
+
Fm. †Ctenochasmatidae Nopcsa 1928 [ctenochasmatid]
+
G. †Balaenognathus Martill et al. 2023
G. †Cathayopterus Wang and Zhou 2006
G. †Forfexopterus Jiang et al. 2016
Unr. †Gnathosaurinae Unwin 1992
G. †Liaodactylus Zhou et al. 2017
G. †Pterofiltrus Jiang and Wang 2011
Invalid names: Ctenochasmatinae Nopcsa 1928 [empty], Pterodaustridae Bonaparte 1971 [synonym]
Fm. †Gallodactylidae Fabre 1974
+
G. †Gladocephaloideus Lü et al. 2012
+
G. †Petrodactyle Hone et al. 2023
+
†Petrodactyle wellnhoferi Hone et al. 2023
Fm. †Lonchodectidae Hooley 1914
+
G. †Serradraco Rigal et al. 2017
+
G. †Yixianopterus Lü et al. 2006
+
†Yixianopterus jingangshanensis Lü et al. 2006
G. †Unwindia Martill 2011
+
†Unwindia trigonus Martill 2011
Invalid names: Pterodactylidae Bonaparte 1838 [invalid subgroup]
G. †Cycnorhamphus Seeley 1870
+
†Cycnorhamphus suevicus Quenstadt 1855
+
Invalid names: Pterodactylus eurychirus Wagner 1858 [synonym]
G. †Elanodactylus Andres and Ji 2008
+
†Elanodactylus prolatus Andres and Ji 2008
G. †Eosipterus Ji and Ji 1997
+
†Eosipterus yangi Ji and Ji 1997
G. †Feilongus Wang et al. 2005
+
†Feilongus youngi Wang et al. 2005
G. †Gallodactylus Fabre 1976
+
G. †Gegepterus Wang et al. 2007
+
†Gegepterus changi Wang et al. 2007
G. †Germanodactylus Young 1964
+
G. †Gnathosaurus Meyer 1833
+
†Gnathosaurus macrurus Seeley 1869
†Gnathosaurus subulatus Meyer 1833
G. †Huanhepterus Dong 1982
+
G. †Kepodactylus Harris and Carpenter 1996
+
†Kepodactylus insperatus Harris and Carpenter 1996
G. †Normannognathus Buffetaut et al. 1998
+
†Normannognathus wellnhoferi Buffetaut et al. 1998
G. †Plataleorhynchus Howse and Milner 1995
+
†Plataleorhynchus streptophorodon Howse and Milner 1995
G. †Pterodaustro Bonaparte 1970
+
†Pterodaustro guinazui Bonaparte 1970
+
Invalid names: Puntanipterus globosus Bonaparte and Sánchez 1975 [synonym]
G. †Mesadactylus Jensen and Padian 1989
+
†Mesadactylus ornithosphyos Jensen and Padian 1989
G. †Ningchengopterus Lü 2009
+
G. †Nyctosaurus Marsh 1876
+
†Nyctosaurus bonneri Miller 1971
†Nyctosaurus gracilis Marsh 1876
†Nyctosaurus lamegoi Price 1953
†Nyctosaurus nanus Marsh 1881
Invalid names: Nyctodactylus Marsh 1881 [synonym]
Fm. †Ornithocheiridae Seeley 1870
+
G. †Aetodactylus Myers 2010
+
†Aetodactylus halli Myers 2010
G. †Araripesaurus Price 1971
G. †Arthurdactylus Frey and Martill 1994
+
†Arthurdactylus conandoylei Frey and Martill 1994
G. †Barbosania Elgin and Frey 2011
+
†Barbosania gracilirostris Elgin and Frey 2011
G. †Caulkicephalus Steel et al. 2005
+
†Caulkicephalus trimicrodon Steel et al. 2005
Invalid names: Criorhynchidae Hooley 1914 [synonym], Ornithocheirinae Williston 1897 [empty], Pteranodontinae Williston 1892 [empty]
Unr. †Pteranodontia Marsh 1876
+
G. †Alamodactylus Andres and Myers 2013
+
†Alamodactylus byrdi Andres and Myers 2013
Unr. †Aponyctosauria Fernandes et al. 2022
+
G. †Alcione Longrich et al. 2018
+
†Alcione elainus Longrich et al. 2018
G. †Epapatelo Fernandes et al. 2022
+
†Epapatelo otyikokolo Fernandes et al. 2022
Unr. †Nyctosauridae Hay 1902
+
G. †Barbaridactylus Longrich et al. 2018
+
†Barbaridactylus grandis Longrich et al. 2018
G. †Muzquizopteryx Frey et al. 2006
+
†Muzquizopteryx coahuilensis Frey et al. 2006
G. †Simurghia Longrich et al. 2018
+
†Simurghia robusta Longrich et al. 2018
Superfm. †Ornithocheiroidea Seeley 1870
+
G. †Anhanguera Campos and Kellner 1985
+
†Anhanguera araripensis Wellnhofer 1985
†Anhanguera blittersdorffi Campos and Kellner 1985
†Anhanguera cuvieri Bowerbank 1851
+
Invalid names: Ornithocheirus colorhinus Seeley 1869 [synonym], Ornithocheirus denticulatus Seeley 1870 [synonym]
†Anhanguera ligabuei Dalla Vecchia 1993
†Anhanguera piscator Kellner and Tomida 2000
†Anhanguera robustus Wellnhofer 1987
†Anhanguera santanae Wellnhofer 1985
Fm. †Anhangueridae Campos and Kellner 1985
+
Subfm. †Coloborhynchinae Holgado et al. 2019
+
G. †Aerodraco Holgado and Pegas 2020
G. †Coloborhynchus Owen 1874
+
Invalid names: Amblydectes Hooley 1914 [synonym]
G. †Nicorhynchus Holgado and Pegas 2020
G. †Uktenadactylus Rodrigues and Kellner 2008
G. †Haliskia Pentland et al. 2024
+
†Haliskia peterseni Pentland et al. 2024
G. †Maaradactylus Bantim et al. 2014
+
†Maaradactylus kellneri Bantim et al. 2014
†Maaradactylus spielbergi Veldmeijer 2003
Subfm. †Tropeognathinae Holgado and Pegas 2020
+
G. †Ferrodraco Pentland et al. 2019
G. †Mythunga Molnar and Thulborn 2007
G. †Siroccopteryx Mader and Kellner 1999
G. †Thapunngaka Richards et al. 2021
G. †Tropeognathus Wellnhofer 1987
Invalid names: Mythungini Richards et al. 2023 [empty]
G. †Brasileodactylus Kellner 1984
+
G. †Cearadactylus Leonardi and Borgomanero 1985
+
†Cearadactylus atrox Leonardi and Borgomanero 1985
G. †Guidraco Wang et al. 2012
+
†Guidraco venator Wang et al. 2012
G. †Istiodactylus Howse et al. 2001
+
†Istiodactylus latidens Seeley 1901
†Istiodactylus sinensis Andres and Ji 2006
Unr. †Lanceodontia Andres et al. 2014
+
Unr. †Istiodactyliformes Kellner et al. 2019
+
G. †Hongshanopterus Wang et al. 2008
Fm. †Istiodactylidae Howse et al. 2001
Unr. †Mimodactylidae Kellner et al. 2019
Unr. †Targaryendraconia Pegas et al. 2020
+
G. †Targaryendraco Pegas et al. 2020
G. †Liaoningopterus Wang and Zhou 2003
+
†Liaoningopterus gui Wang and Zhou 2003
G. †Lonchodectes Hooley 1914
+
†Lonchodectes microdon Seeley 1869
+
Invalid names: Ornithocheirus huxleyi Seeley 1870 [synonym], Ornithocheirus oweni Seeley 1870 [synonym]
†Lonchodectes platystomus Seeley 1870
+
Invalid names: Pterodactylus daviesii Owen 1874 [synonym]
G. †Longchengpterus Wang et al. 2006
+
†Longchengpterus zhaoi Wang et al. 2006
G. †Ludodactylus Frey et al. 2003
+
†Ludodactylus sibbicki Frey et al. 2003
G. †Nurhachius Wang et al. 2005
+
†Nurhachius ignaciobritoi Wang et al. 2005
†Nurhachius luei Zhou et al. 2019
G. †Ornithocheirus Seeley 1869
+
†Ornithocheirus simus Owen 1861
+
Invalid names: Ornithocheirus platyrhinus Seeley 1870 [synonym]
Invalid names: Cimoliornis Owen 1846 [synonym], Criorhynchus Owen 1874 [objective synonym], Palaeornis Mantell 1835 [synonym]
G. †Ornithodesmus Seeley 1887
+
†Ornithodesmus cluniculus Seeley 1887
G. †Piksi Varricchio 2002
+
†Piksi barbarulna Varricchio 2002
Invalid names: Euornithocheira Unwin 2003 [empty]
G. †Pteranodon Marsh 1876
+
Subg. †Pteranodon (Longicepia) Miller 1971
Subg. †Pteranodon (Occidentalia) Miller 1971
Subg. †Pteranodon (Sternbergia) Miller 1971
†Pteranodon longiceps Marsh 1876
+
Invalid names: Pteranodon (Longicepia) marshi Miller 1971 [synonym], Pteranodon (Sternbergia) walkeri Miller 1971 [synonym]
†Pteranodon sternbergi Harksen 1966
+
Invalid names: Pteranodon (Occidentalia) eatoni Miller 1971 [synonym]
Invalid names: Ornithocheirus harpyia Cope 1872 [nomen dubium], Ornithocheirus umbrosus Cope 1872 [nomen dubium], Pteranodon comptus Marsh 1876 [nomen dubium], Pterodactylus ingens Marsh 1872 [nomen dubium], Pterodactylus occidentalis Marsh 1872 [nomen dubium], Pterodactylus oweni Marsh 1871 [nomen dubium], Pterodactylus velox Marsh 1872 [nomen dubium]
Fm. †Pteranodontidae Marsh 1876
+
G. †Dawndraco Kellner 2010
+
†Dawndraco kanzai Kellner 2010
G. †Geosternbergia Miller 1978
+
†Geosternbergia maiseyi Kellner 2010
G. †Santanadactylus Buisonjé 1980
+
†Santanadactylus brasilensis Buisonjé 1980
†Santanadactylus spixi Wellnhofer 1985
G. †Tethydraco Longrich et al. 2018
+
†Tethydraco regalis Longrich et al. 2018
G. †Volgadraco Averianov et al. 2008
+
†Volgadraco bogolubovi Averianov et al. 2008
Invalid names: Azhdarchinae Nessov 1984 [empty], Ornithostomatidae Williston 1893 [synonym], Ornithostomatinae Williston 1897 [synonym]
Superfm. †Pteranodontoidea Kellner 1996
+
Unr. †Anhangueria Rodrigues and Kellner 2013
+
G. †Camposipterus Rodrigues and Kellner 2013
+
†Camposipterus colorhinus Seeley 1869
†Camposipterus nasutus Seeley 1869
+
Invalid names: Pterodactylus fittoni Owen 1859 [synonym]
Fm. †Hamipteridae Holgado et al. 2019
+
G. †Iberodactylus Holgado et al. 2019
+
†Iberodactylus andreui Holgado et al. 2019
Invalid names: Ornithocheirae Seeley 1870 [empty]
G. †Aussiedraco Kellner et al. 2011
+
†Aussiedraco molnari Kellner et al. 2011
G. †Cimoliopterus Rodrigues and Kellner 2013
+
†Cimoliopterus cuvieri Bowerbank 1851
†Cimoliopterus dunni Myers 2015
G. †Hamipterus Wang et al. 2014
+
†Hamipterus tianshanensis Wang et al. 2014
G. †Ikrandraco Wang et al. 2014
+
†Ikrandraco avatar Wang et al. 2014
G. †Linlongopterus Rodrigues et al. 2015
+
†Linlongopterus jennyae Rodrigues et al. 2015
Fm. †Saraikisauridae Malkani 2014
+
Subfm. †Saraikisaurinae Malkani 2014
+
G. †Saraikisaurus Malkani 2014
+
†Saraikisaurus minhui Malkani 2014
Fm. †Tendaguripteridae Kellner et al. 2007
+
G. †Tendaguripterus Unwin and Heinrich 1999
+
†Tendaguripterus recki Unwin and Heinrich 1999
G. †Wenupteryx Codorniú and Gasparini 2013
+
†Wenupteryx uzi Codorniú and Gasparini 2013
Invalid names: Araripedactylus Wellnhofer 1977 [nomen dubium], Araripedactylus dehmi Wellnhofer 1977 [nomen dubium], Araripesaurus castilhoi Price 1971 [nomen dubium], Belonochasmatidae [empty], Cearadactylidae Wellnhofer 1991 [empty], Dermodactylus montanus Marsh 1878 [nomen dubium], Doratorhynchus validus Owen 1870 [nomen vanum], Eupterodactyloidea Bennett 1994 [empty], Ornithocheirus brachyrhinus Seeley 1870 [nomen dubium], Ornithocheirus bunzeli Seeley 1881 [nomen dubium], Ornithocheirus carteri Seeley 1870 [nomen dubium], Ornithocheirus crassidens Seeley 1870 [nomen dubium], Ornithocheirus dentatus Seeley 1870 [nomen dubium], Ornithocheirus enchorhynchus Seeley 1870 [nomen dubium], Ornithocheirus eurygnathus Seeley 1870 [nomen dubium], Ornithocheirus oxyrhinus Seeley 1870 [nomen dubium], Ornithocheirus scaphorhynchus Seeley 1870 [nomen dubium], Ornithocheirus tenuirostris Seeley 1870 [nomen dubium], Ornithocheirus xyphorhynchus Seeley 1870 [nomen dubium], Ornithodesmidae Hooley 1913 [empty], Osteornis Rodrigues and Kellner 2013 [nomen dubium], Pterodactylus grandipelvis Meyer 1860 [nomen dubium], Pterodactylus grandis Cuvier 1824 [nomen dubium], Pterodactylus longipes Münster 1859 [nomen dubium], Pterodactylus woodwardi Owen 1861 [nomen dubium], Santanadactylus pricei Wellnhofer 1985 [nomen dubium], Titanopterygiidae Padian 1984 [empty]
Diagnosis
ReferenceDiagnosis
D. M. Unwin 2003Pteranodon longiceps, Quetzalcoatlus northropi, their most recent common ancester and all descendants