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Tracheophyta

Taxonomy
Tracheophyta was named by Sinnott (1935) [Kenrick and Crane 1997 rank this as a division.]. It is extant. It was considered monophyletic by Jud (2011).

It was reranked as the unranked clade Tracheophyta by Doweld (2001); it was reranked as the phylum Tracheophyta by Jud (2011).

It was assigned to Polysporangiomorpha by Kenrick and Crane (1997); and to Plantae by Doweld (2001) and Jud (2011).

Synonymy list
YearName and author
1935Tracheophyta Sinnott
1997Tracheophyta Kenrick and Crane
2001Tracheophyta Doweld
2011Tracheophyta Jud

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RankNameAuthor
kingdomPlantae
phylumTracheophyta(Sinnott 1935)
RankNameAuthor
phylumTracheophyta(Sinnott 1935)

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Ph. Tracheophyta Sinnott 1935
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Unr. †Bryobiotina Doweld 2001
+
Ph. †Langiophytophyta Doweld 2001
+
Cl. †Langiophytopsida Doweld 2001
+
Or. †Langiophytales Doweld 2001
+
Invalid names: Langiophytaceae Doweld 2001 [empty]
Or. †Lyonophytales Doweld 2001
+
Invalid names: Lyonophytaceae Doweld 2001 [empty]
G. †Catenalis Hao and Xue 2013
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†Catenalis digitata Hao and Xue 2013
Subph. †Euphyllophytina
+
+
Cl. †Cladoxylopsida Novák 1930
+
G. †Pietzschia Gothan 1921
+
†Pietzschia schulleri Gothan 1927
+
G. †Calamophyton Krausel and Weyland
G. †Cladoxylon Unger 1856
+
Invalid names: Hierogramma Unger 1856 [synonym], Schizoxylon Unger 1856 [synonym]
G. †Eospermatopteris Goldring 1924
G. †Hyenia Nathorst 1915
G. †Lorophyton Fairon-Demaret and Li 1993
G. †Rhymokalon Scheckler 1975
Or. †Steloxylales Doweld 2001
+
Invalid names: Steloxylaceae Doweld 2001 [empty]
G. †Syncardia Unger 1856
+
†Syncardia pusilla Unger 1856
Unr. †Zygopteridae
+
G. †Austroclepsis Sahni 1932
G. †Corynepteris Bailey 1860
Invalid names: Eoclepsydropsis Bertrand 1909 [nomen nudum]
Invalid names: Cladoxyliidae [empty], Cladoxylophyceae [replaced], Stauropteridae [empty]
Unr. †Radiatopses
+
Unr. †Lignophytia
+
+
G. †Cardioneura Zalessky 1934
Fm. †Medullosaceae Göppert 1864
Invalid names: Hydraspermaceae [empty]
Invalid names: Protopityales Walton 1957 [empty]
G. †Pertica Kasper and Andrews 1972
+
†Pertica quadrifaria Kasper and Andrews 1972
+
Or. †Corystospermales Thomas 1933
+
Fm. †Corystospermaceae Thomas 1933
+
G. †Cuneumxylon Artabe and Brea 2003
+
†Cuneumxylon spallettii Artabe and Brea 2003
G. †Cycadopteris Zigno 1853
+
†Cycadopteris auriculata Bose and Sukh-Dev 1959
†Cycadopteris brauniana Zigno 1861
†Cycadopteris indica Bose and Sukh-Dev 1959
†Cycadopteris majus Bose and Sukh-Dev 1959
†Cycadopteris pulcherrima Bose and Sukh-Dev 1959
†Cycadopteris ungeri Zigno 1853
G. †Dicroidiopsis Frenguelli 1943
+
†Dicroidiopsis crassa Menéndez 1951
†Dicroidiopsis incisa DuToit 1943
G. †Dicroidium Gothan 1912
+
†Dicroidium coriaceum Johnston 1886
†Dicroidium crassinervis Geinitz 1876
†Dicroidium dubium Feistmantel 1878
+
Invalid names: Sphenopteris bergina Jones and de Jersey 1947 [synonym]
†Dicroidium elongatum Carruthers 1872
†Dicroidium eskense Walkom 1928
†Dicroidium feistmantelii Johnston 1847
†Dicroidium gopadensis Bose and Srivastava 1971
†Dicroidium irnensis Abu Hamad et al. 2008
†Dicroidium jordanensis Abu Hamad et al. 2008
†Dicroidium natalense Frenguelli 1943
†Dicroidium nidpurensis Bose and Srivastava 1971
+
Invalid names: Thinnfeldia subtrigona Feistmantel 1879 [synonym]
†Dicroidium papillosum Bose and Srivastava 1971
†Dicroidium prolungatum Menéndez 1951
†Dicroidium robustum Abu Hamad et al. 2008
†Dicroidium sahnii Seward 1932
†Dicroidium shirleyi Holmes 1982
†Dicroidium spinifolium Tenison-Woods 1883
†Dicroidium stelznerianum Geinitz 1876
†Dicroidium superbum Shirley 1898
†Dicroidium voiseyi Holmes and Ash 1979
†Dicroidium zuberi Szajnocha 1888
G. †Elchaxylon Artabe and Zamuner 2007
+
†Elchaxylon zavattieriae Artabe and Zamuner 2007
G. †Fanerotheca Frenguelli 1944
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†Fanerotheca extans Frenguelli 1944
G. †Feruglioa Frenguelli 1944
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†Feruglioa samaroides Frenguelli 1944
G. †Komlopteris Barbacka 1994
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†Komlopteris cenozoicus McLoughlin et al. 2008
G. †Pteruchus Thomas 1933
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†Pteruchus dubius Thomas 1933
†Pteruchus gopadensis Pant and Basu 1979
†Pteruchus indicus Pant and Basu 1973
†Pteruchus johnstonii Feistmantel 1890
+
Invalid names: Pteruchus africanus Thomas 1933 [synonym], Stachyopitys annularioides Shirley 1898 [synonym]
†Pteruchus nidpurensis Srivastava 1975
†Pteruchus rhaetica Geinitz 1876
†Pteruchus thomasii Pant and Basu 1979
G. †Trichomanides Tenison-Woods 1883
+
†Trichomanides laxum Tenison-Woods 1883
G. †Umkomasia Thomas 1933
+
†Umkomasia cacheutensis Frenguelli 1942
†Umkomasia distans Holmes 1987
†Umkomasia macleani Thomas 1933
†Umkomasia polycarpa Holmes 1987
†Umkomasia sessilis Holmes 1987
Invalid names: Beania geminata Shirley 1898 [nomen dubium], Karibacarpon Lacey 1974 [synonym], Pilophorosperma Thomas 1933 [synonym]
G. †Xylopteris Frenguelli 1943
+
†Xylopteris elongata Carruthers 1872
†Xylopteris rigida Dun 1909
+
Invalid names: Stenopteris densifolia Thomas 1933 [synonym]
Invalid names: Umkomastiaceae [replaced]
G. †Doylea Stockey and Rothwell 2009
G. †Laevigatasporites Potonié and Gelletich 1933
G. †Laricoidites Potonie et al. 1958
+
†Laricoidites communis Sah and Jain 1965
†Laricoidites desquamatus Goubin 1965
†Laricoidites gigantus Brenner 1963
†Laricoidites indicus Singh et al. 1964
†Laricoidites kutchensis Venkatachala et al. 1968
G. †Leioaletes Staplin
G. †Ovalipollis Krutzsch 1955
+
†Ovalipollis breviformis Krutzsch 1955
†Ovalipollis lunzenzis Klaus 1960
†Ovalipollis pseudoalutus Krutzsch 1955
G. †Singhia Srivastava 1967
+
†Singhia acicularis Lima 1980
†Singhia crenulata de Lima 1980
†Singhia elongata de Lima 1980
†Singhia minima Lima 1980
†Singhia montanaensis Lima 1980
†Singhia multicostata Lima 1980
†Singhia punctata de Lima 1980
†Singhia reyrei de Lima 1980
†Singhia urwashii Srivastava 1967
Subph. †Lycophytina
+
G. †Zosterophyllum Penhallow 1892
+
†Zosterophyllum myretonianum Penhallow 1892
Unr. †Lycopodiobiotina Doweld 2001
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Ph. Isoetophyta Doweld 2001
+
Subph. Isoetophytina Doweld 2001
+
Cl. †Lepidocarpopsida Doweld 2001
+
Or. †Jurinodendrales Doweld 2001
Subph. †Leclercqiophytina Doweld 2001
+
Cl. †Leclercqiopsida Doweld 2001
+
Or. †Leclercqiales Doweld 2001
Subph. Selaginellophytina Doweld 2001
+
Cl. Selaginellopsida Frank 1874
+
Invalid names: Miadesmiales Doweld 2001 [empty]
Cl. †Progymnospermopsida Lang 1925
+
G. †Cairoa Matten 1973
+
†Cairoa lamanekii Matten 1973
G. †Praeramunculus Anderson and Anderson 1985
G. †Tetraxylopteris Beck 1957
+
Unr. †Pteridobiotina Doweld 2001
+
Unr. †Cycadophytanae Doweld 2001
+
Ph. †Aneurophytophyta Bold 1973
+
Subph. †Aneurophytophytina Doweld 2001
+
Cl. †Aneurophytopsida Takhtajan 1978
Subph. †Archaeopteridophytina Doweld 2001
+
Cl. †Archaeopteridopsida Takhtajan 1978
Invalid names: Archaeopteridophyta Snigirevskaya 2000 [synonym]
Ph. †Cycadeoideophyta Taylor 1981
+
Cl. †Cycadeoideopsida Scott 1923
+
Or. †Cycadeoideales Berry 1916
G. †Cycadeoidella Ogura 1930
Invalid names: Bennettitopsida Engler 1892 [synonym]
Ph. Gnetophyta Bessey 1907
+
Cl. Gnetopsida Kirpotenko 1884
+
G. †Gontriglossa Anderson and Anderson 1989
G. †Pityophyllum Nathorst 1899
Ph. Magnoliophyta Cronquist et al.
+
G. †Andreisporis Belsky et al. 1965
+
†Andreisporis mariae Belsky et al. 1965
+
G. †Aquilapollenites Rouse 1957
G. †Cranwellia Srivastava 1966
Unr. †Liliopsida Kenrick and Crane 1997
+
Invalid names: Liliophyceae Kenrick and Crane 1997 [replaced]
G. †Loranthacites Mtchedlishvili 1961
G. †Araripia Mohr and Eklund 2003
+
†Araripia florifera Mohr and Eklund 2003
G. †Asteropollis Hedlund and Norris 1968
+
†Asteropollis asteroides Hedlund and Norris 1968
†Asteropollis transitorius Vallati 2006
†Asteropollis vulgaris Singh 1983
G. †Auriculiidites Elsik 1964
+
†Auriculiidites hourcqui Boltenhagen 1967
G. †Auriculopollenites Legoux et al. 1970
+
†Auriculopollenites reticulatus Salard-Cheboldaeff 1978
G. †Bacumorphomonocolpites Sole de Porta 1971
+
†Bacumorphomonocolpites tausae Sole de Porta 1971
G. †Balanopollis Salard-Cheboldaeff 1978
+
†Balanopollis minutus Salard-Cheboldaeff 1978
G. Bignonia Linnaeus 1753
+
Bignonia caerulea Linnaeus 1753
Bignonia catalpa Linnaeus 1753
Bignonia crucigera Linnaeus 1753
Bignonia espreolata Linnaeus 1753
†Bignonia gigantifolia Engelhardt 1891
Bignonia indica Linnaeus 1753
Bignonia loucoxylon Linnaeus 1753
Bignonia paniculata Linnaeus 1753
Bignonia peruviana Linnaeus 1753
Bignonia radiata Linnaeus 1753
Bignonia radicans Linnaeus 1753
Bignonia sempervireus Linnaeus 1753
Bignonia unguiscati Linnaeus 1753
G. †Brenneripollis Juhász and Goczan 1985
+
†Brenneripollis pellitusreticulatus Juhász and Goczan 1985
G. †Brevicolporites Anderson 1960
+
†Brevicolporites guinetti Salard-Cheboldaeff 1978
†Brevicolporites molinae Salard-Cheboldaeff 1978
G. †Ceratocarpus Velenovsky and Viniklar 1931
+
†Ceratocarpus fendrychi Velenovsky and Viniklar 1931
G. †Constructipollenites van Hoeken-Klinkenberg 1964
+
†Constructipollenites ineffectus van Hoeken-Klinkenberg 1964
G. †Cornetipollis Pocock and Vasanthy 1988
+
†Cornetipollis herngreenii Deaf et al. 2014
G. †Crassitriapertites Herngreen 1972
+
†Crassitriapertites uesugii Herngreen 1972
G. †Crassitricolporites Herngreen 1972
+
G. †Cristaecolpites Schrank 1994
+
G. †Cristaeturites Schrank 1994
+
†Cristaeturites cristatus Schrank 1994
G. †Crusafontites Sole de Porta 1971
+
†Crusafontites megagemmatus Jaramillo and Dilcher 2001
G. †Cupuliferoidaepollenites Potonié et al. 1950
+
+
Invalid names: Psilatricolpites parvulus Norris 1967 [synonym]
G. †Dettmannaepollenites Kedves 1999
+
†Dettmannaepollenites clavatus Schrank and Mahmoud 2000
G. †Dicolpopollis Pflanzl 1956
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†Dicolpopollis malesianus Muller 1968
G. †Dryadopollis Srivastava 1977
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†Dryadopollis argus Srivastava 1975
†Dryadopollis minima Barreda 1997
G. †Echitricolpites Regali et al. 1974
G. †Foveomonocolpites Wijmstra 1971
+
G. †Foveotetradites Singh 1983
+
G. †Foveotricolpites Pierce 1961
+
†Foveotricolpites genuinus González-Guzmán 1967
†Foveotricolpites giganteus Du Chene 1978
†Foveotricolpites perforatus Van der Hammen and Garcia de Mutis 1966
†Foveotricolpites pomarius Van der Hammen and Garcia de Mutis 1966
†Foveotricolpites prolatus Frederiksen 1973
†Foveotricolpites santanderianus Van der Hammen and Garcia de Mutis 1966
G. †Fraxinoipollenites Potonié 1960
+
†Fraxinoipollenites variformis Norvick and Burger 1975
G. †Hammenia Ward 1986
G. †Hexacolpites Erdtman 1947
+
†Hexacolpites echinatus Salard-Cheboldaeff 1978
†Hexacolpites indicus Ramanujam 1966
G. †Holkopollenites Stover 1966
+
†Holkopollenites chemardensis Stover et al. 1966
†Holkopollenites forix Christopher and Prowell 2010
†Holkopollenites irregularis Christopher and Prowell 2002
†Holkopollenites propinquus Christopher and Prowell 2010
G. †Inaperturotetradites van Hoeken-Klinkenberg 1964
+
†Inaperturotetradites lacunosus van Hoeken-Klinkenberg 1964
†Inaperturotetradites reticulatus Salard-Cheboldaeff 1978
G. †Integritetradites Schrank and Mahmoud 2000
+
†Integritetradites porosus Schrank and Mahmoud 2000
G. †Intrareticulitis Kar 1985
+
G. †Jugella Mchedlishvili and Shakmundes 1973
+
†Jugella caichiguensis Schrank 2010
†Jugella semistriata Schrank 2010
†Jugella sibirica Mtchedlishvili and Shakmundes 1973
G. †Lethomasites Ward 1989
G. †Microfoveotriporites van Hoeken-Klinkenberg 1966
+
†Microfoveotriporites cretaceus van Hoeken-Klinkenberg 1966
G. †Monocolpopollenites Pflug and Thomson 1953
+
†Monocolpopollenites ovatus Jaramillo and Dilcher 2001
†Monocolpopollenites sphaeroidites Jardiné and Magloire 1965
†Monocolpopollenites spheroidites Jardiné and Magloire 1965
G. †Multiporopollenites Pflug 1953
+
†Multiporopollenites pauciporatus Jaramillo and Dilcher 2001
G. †Nigeripollis Srivastava 1975
+
†Nigeripollis gemmatus Salami 1985
G. †Nyssapollenites Thiergart 1937
+
†Nyssapollenites barooahii Sah and Dutta 1968
†Nyssapollenites endobalteus Mildenhall and Pocknall 1989
†Nyssapollenites incertus Dutta and Sah 1970
†Nyssapollenites kutchensis Venkatachala and Kar 1969
†Nyssapollenites lanosus Dettmann 1973
†Nyssapollenites lehmanii Schrank 1994
†Nyssapollenites pachyexinus Baldoni and Askin 1993
†Nyssapollenites scabratus De Benedetti et al. 2023
†Nyssapollenites squamosus Dettmann 1973
G. †Pediculisporis Belsky et al. 1965
+
†Pediculisporis reticularis Belsky et al. 1965
†Pediculisporis rnicrogranulatus Belsky et al. 1965
G. †Penetetrapites Hedlund and Norris 1968
+
†Penetetrapites incipiens de Lima 1989
†Penetetrapites mollis Hedlund and Norris 1968
G. †Phimopollenites Dettmann 1973
+
†Phimopollenites pannosus Dettmann 1974
†Phimopollenites pseudocheros Srivastava 1975
G. †Pleurospermaepollenites Kulkova 1973
G. †Polycolpopollenites Nakoman 1967
G. †Polyporotetradites Salard-Cheboldaeff 1978
+
†Polyporotetradites laevigatus Salard-Cheboldaeff 1978
G. †Protrudodiporites van Hoeken-Klinkenberg 1966
+
†Protrudodiporites typicus van Hoeken-Klinkenberg 1966
G. †Psilabrevitricolpites van Hoeken-Klinkenberg 1966
+
†Psilabrevitricolpites flexibilis van Hoeken-Klinkenberg 1966
†Psilabrevitricolpites rotundus van Hoeken-Klinkenberg 1966
G. †Psilabrevitricolporites Van der Kaars 1983
+
†Psilabrevitricolporites triangularis Jaramillo and Dilcher 2001
G. †Psilamonocolpites Van der Hammen and Garcia de Mutis 1966
+
†Psilamonocolpites grandis Van der Hammen 1954
†Psilamonocolpites magnus Du Chene 1978
†Psilamonocolpites medius Van der Hammen 1954
G. †Psilastephanocolpites Leidelmeyer 1966
+
†Psilastephanocolpites adinos González-Guzmán 1967
†Psilastephanocolpites marginatus González-Guzmán 1967
†Psilastephanocolpites parvus Salard-Cheboldaeff 1978
†Psilastephanocolpites verrucosus González-Guzmán 1967
G. †Psilastephanocolporites Leidelmeyer 1966
+
†Psilastephanocolporites brevicolpatus Jaramillo and Dilcher 2001
†Psilastephanocolporites globulus van Hoeken-Klinkenberg 1966
†Psilastephanocolporites grandis Salard-Cheboldaeff 1978
†Psilastephanocolporites laevigatus Salard-Cheboldaeff 1978
†Psilastephanocolporites minor Salard-Cheboldaeff 1978
†Psilastephanocolporites perforatus Salard-Cheboldaeff 1978
†Psilastephanocolporites punctatus Salard-Cheboldaeff 1978
†Psilastephanocolporites scabratus Salard-Cheboldaeff 1978
G. †Psilastephanoporites Regali et al. 1974
G. †Psilatricolpites Van der Hammen and Wijmstra 1964
+
†Psilatricolpites acerbus González-Guzmán 1967
†Psilatricolpites brevis González-Guzmán 1967
†Psilatricolpites colpiconstrictus van Hoeken-Klinkenberg 1966
†Psilatricolpites inargutus Archangelsky 1973
+
Invalid names: Tricolpites inargutus McIntyre 1968 [replaced]
†Psilatricolpites minutus González-Guzmán 1967
†Psilatricolpites palaeocenica Van der Hammen and Garcia de Mutis 1966
†Psilatricolpites polaroides González-Guzmán 1967
†Psilatricolpites psilascabratus Archangelsky 1973
+
Invalid names: Tricolpites psilascabratus Norton 1969 [replaced]
†Psilatricolpites psilatus Potonié 1966
†Psilatricolpites simplex Jaramillo and Dilcher 2001
†Psilatricolpites tetradus Brenner 1968
Invalid names: Psilatricolpites rubini Van der Hammen and Wymstra 1964 [nomen nudum]
G. †Ranunculacidites Sah 1967
+
†Ranunculacidites operculatus Jaramillo and Dilcher 2001
+
Invalid names: Psilatricolporites operculatus Van der Hammen and Wijmstra 1964 [synonym]
G. †Retibrevitricolpites van Hoeken-Klinkenberg 1966
+
†Retibrevitricolpites catatumbus González-Guzmán 1967
†Retibrevitricolpites distinctus van Hoeken-Klinkenberg 1966
†Retibrevitricolpites grandis Jaramillo and Dilcher 2001
†Retibrevitricolpites increatus González-Guzmán 1967
†Retibrevitricolpites retibolus Leidelmeyer 1966
†Retibrevitricolpites triangulatus van Hoeken-Klinkenberg 1966
G. †Retibrevitricolporites Legoux 1978
+
†Retibrevitricolporites grandis Jaramillo and Dilcher 2001
†Retibrevitricolporites speciosus Jaramillo and Dilcher 2001
G. †Retipollenites González-Guzmán 1967
+
†Retipollenites confusus González-Guzmán 1967
G. †Retistephanocolpites Leidelmeyer 1966
+
†Retistephanocolpites angeli Leidelmeyer 1966
†Retistephanocolpites finalis González-Guzmán 1967
†Retistephanocolpites fossulatus Jaramillo and Dilcher 2001
†Retistephanocolpites minutus González-Guzmán 1967
†Retistephanocolpites regularis van Hoeken-Klinkenberg 1966
†Retistephanocolpites williamsii Germeraad et al. 1968
G. †Retistephanoporites González-Guzmán 1967
+
†Retistephanoporites angelicus González-Guzmán 1967
†Retistephanoporites crassiexinatus Jaramillo and Dilcher 2001
†Retistephanoporites minutiporus Jaramillo and Dilcher 2001
G. †Retisyncolporites González-Guzmán 1967
+
†Retisyncolporites angularis González-Guzmán 1967
†Retisyncolporites aureus González-Guzmán 1967
G. †Retitrescolpites Sah 1967
+
†Retitrescolpites assamicus Dutta and Sah 1970
†Retitrescolpites baculatus Jaramillo and Dilcher 2001
†Retitrescolpites irregularis Jaramillo and Dilcher 2001
+
Invalid names: Retitricolporites irregularis Van der Hammen and Wijmstra 1964 [replaced]
†Retitrescolpites magnus Jaramillo and Dilcher 2001
+
Invalid names: Retitricolpites magnus González-Guzmán 1967 [replaced]
†Retitrescolpites minor Dutta and Sah 1970
†Retitrescolpites peculiaris Jaramillo and Dilcher 2001
†Retitrescolpites saturum Jaramillo and Dilcher 2001
+
Invalid names: Retitricolpites saturum González-Guzmán 1967 [replaced]
G. †Retitricolpites Van der Hammen 1956
+
†Retitricolpites absolutus Jaramillo and Dilcher 2001
†Retitricolpites adeptus González-Guzmán 1967
†Retitricolpites adultus González-Guzmán 1967
†Retitricolpites amapaensis Regali et al. 1974
†Retitricolpites americana Wymstra 1966
†Retitricolpites antonii González-Guzmán 1967
†Retitricolpites bonus González-Guzmán 1967
†Retitricolpites clarensis González-Guzmán 1967
†Retitricolpites conciliatus González-Guzmán 1967
†Retitricolpites constrictus González-Guzmán 1967
†Retitricolpites fragosus Hedlund and Norris 1968
†Retitricolpites herrerae Van der Hammen and Garcia de Mutis 1966
†Retitricolpites incisus González-Guzmán 1967
†Retitricolpites jardinei Kedves 1999
†Retitricolpites maledictus González-Guzmán 1967
†Retitricolpites marginatus van Hoeken-Klinkenberg 1966
†Retitricolpites maturus González-Guzmán 1967
†Retitricolpites microreticulatus Van der Hammen 1954
†Retitricolpites minutus Kedves 1999
†Retitricolpites obtusus van Hoeken-Klinkenberg 1966
†Retitricolpites operculatus Herngreen 1973
†Retitricolpites perditus González-Guzmán 1967
†Retitricolpites polaris Gonzalez 1966
†Retitricolpites robustus Sah and Kar 1970
†Retitricolpites salardae Kedves 1999
†Retitricolpites simplex González-Guzmán 1967
†Retitricolpites sitholeyii Ramanujam 1966
†Retitricolpites virgeus Brenner 1962
†Retitricolpites vulgaris Pierce 1961
G. †Scabratriporites Van der Hammen 1956
+
†Scabratriporites annellus van Hoeken-Klinkenberg 1964
†Scabratriporites moderatus González-Guzmán 1967
†Scabratriporites redundans González-Guzmán 1967
†Scabratriporites simpliformis van Hoeken-Klinkenberg 1966
G. †Schrankipollis Doyle et al. 1990
+
†Schrankipollis kachaikensis Barreda and Archangelsky 2006
†Schrankipollis mawhoubensis Doyle et al. 1990
†Schrankipollis reticulatus Regali and Santos 1996
G. †Senectotetradites Dettmann 1973
+
G. †Stephanoporopollenites Thomson 1953
+
G. †Striatriporites van Hoeken-Klinkenberg 1966
+
†Striatriporites nigeriensis van Hoeken-Klinkenberg 1966
G. †Translucentipollis Chlonova 1961
+
G. †Tricolporoidites Pacltova 1971
+
G. †Trifossapollenites Rouse 1957
+
†Trifossapollenites constatus Dutta and Sah 1970
†Trifossapollenites ivoirensis Jardiné and Magloire 1965
G. †Triporites Van der Hammen 1954
G. †Triporotetradites van Hoeken-Klinkenberg 1964
+
†Triporotetradites hoekenii Salard-Cheboldaeff 1978
†Triporotetradites letouzeyi Salard-Cheboldaeff 1978
†Triporotetradites scabratus van Hoeken-Klinkenberg 1964
G. †Verrustephanocolporites Salard-Cheboldaeff 1978
+
†Verrustephanocolporites complanatus Salard-Cheboldaeff 1978
G. †Verrutricolpites Pierce 1961
+
†Verrutricolpites irregularis González-Guzmán 1967
†Verrutricolpites triangulus Sah and Kar 1970
†Verrutricolpites unicus González-Guzmán 1967
Ph. †Moresnetiophyta Doweld 2001
+
Cl. †Moresnetiopsida Doweld 2001
+
Or. †Pullarithecales Doweld 1998
Invalid names: Moresnetiales Doweld 2001 [empty]
Ph. †Noeggerathiophyta Doweld 2001
+
G. †Koilosphenus Bohlin 1971
+
†Koilosphenus cuneifolius Bohlin 1971
Ph. †Peltaspermophyta Doweld 2001
+
Cl. †Arberiopsida Doweld 2001
+
Or. †Arberiales Meyen 1982
+
Invalid names: Glossopteridales Melville 1960 [replaced]
Or. †Vojnovskyales Neuburg 1965
Cl. †Dictyopteridiopsida Doweld 2001
+
Or. †Dictyopteridiales Doweld 2001
Or. †Lidgettoniales Doweld 2001
Invalid names: Glossopteridopsida Plumstead 1956 [synonym], Rigbyales Doweld 2001 [empty]
G. †Hermanophyton Arnold 1962
+
†Hermanophyton kirkbyorum Arnold 1962
G. †Lanceolatus Plumstead 1952
+
†Lanceolatus derouxides Plumstead 1952
Cl. †Peltaspermopsida Doweld 2001
+
G. †Aneimites Dawson 1860
Or. †Peltaspermales Delevoryas 1979
G. †Psaliangium Remy 1953
Or. †Sporophyllitales Doweld 2001
Or. †Trichopityales Doweld 2001
Or. †Umkomasiales Doweld 2001
G. †Reticulopteris Gothan 1941
+
†Reticulopteris muensteri Eichwald 1840
Invalid names: Pentoxylopsida Pant 1959 [empty]
Unr. †Pteridophytanae Doweld 2001
+
Ph. †Botryopteridiophyta Doweld 2001
+
Cl. †Botryopteridiopsida Doweld 2001
+
Or. †Botryopteridiales Doweld 2001
Or. †Neomariopteridales Doweld 2001
Invalid names: Senftenbergiales Doweld 2001 [empty]
Cl. †Urnatopteridopsida Doweld 2001
+
Or. †Crossothecales Doweld 2001
Or. †Discopteridales Doweld 2001
Or. †Urnatopteridales Doweld 2001
Invalid names: Stauropteridopsida Doweld 2001 [empty]
Ph. †Cladoxylophyta Doweld 2001
+
G. †Asteropteris Dawson 1880
+
Ph. Equisetophyta Scott 1900
+
Cl. †Asterocalamitopsida Doweld 2001
+
Or. †Asterocalamitales Nakai 1943
Or. †Autophyllitales Doweld 2001
Cl. †Calamitopsida Endlicher 1841
+
Or. †Calamitales Eichwald 1852
Invalid names: Calamocarpales Doweld 2001 [empty]
Cl. Equisetopsida Agardh 1825
+
Subcl. †Anthocerotidae Engler and Prantl 1893
G. †Chlamidostachys White 1937
Or. †Gondwanostachyales Doweld 2001
G. †Paraphyllanthoxylon Bailey 1924
+
Invalid names: Glochidioxylon Ramanujam 1956 [synonym]
Subcl. Pinidae Cronquist et al. 1966
Or. †Sphenophyllales Seward 1898
Invalid names: Bryidae [empty], Echinostachyales Doweld 2001 [empty], Equisetophyceae [replaced]
G. †Parasphenophyllum Asama 1970
+
G. †Paratrizygia Asama 1970
+
†Paratrizygia kobonensis Kobatake 1970
Ph. Marattiophyta Heintze 1927
+
Cl. Marattiopsida Doweld 2001
+
Or. †Asterothecales Doweld 2001
Invalid names: Ophioglossophyta Doweld 2001 [empty], Sphenophyllophyta Doweld 2001 [empty]
Invalid names: Psylophytanae Doweld 2001 [empty]
Fm. †Rhyniaceae Kidston and Lang 1920
+
G. †Palaeostigma Kraeusel and Dolianiti 1957
+
†Palaeostigma sewardi Kraeusel and Dolianiti 1957
G. †Steganotheca Edwards 1970
+
†Steganotheca striata Edwards 1970
G. †Taeniophyton Weyland and Berendt 1968
+
†Taeniophyton inopinatum Weyland and Berendt 1968
Cl. †Rhyniopsida Kryshtofovich 1925
+
Invalid names: Rhyniales Nemejc 1950 [empty], Rhyniophyta Cronquist et al. 1966 [replaced]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
No ecological data are available

Age range

Maximum range based only on fossils: base of the Ludfordian to the top of the Late/Upper Pleistocene or 425.60000 to 0.01170 Ma
Minimum age of oldest fossil (stem group age): 423.0 Ma

Collections (4899 total)

Oldest occurrences

Time interval Ma Country or state Original ID and collection number
Silurian443.8 - 419.2Norway (Svalbard) Zosterophyllum sp. (20839)
Gorstian - Pridoli427.4 - 419.2Bolivia (Tarija dept.) Steganotheca sp. (11158)
Ludlow - Pridoli427.4 - 419.2Canada (Nunavut) Zosterophyllum b (10599) Zosterophyllum sp. (53920)
Ludfordian425.6 - 423.0Canada (Nunavut) Zosterophyllum sp. (11426)
Ludfordian - Pridoli425.6 - 419.2Ukraine Zosterophyllum myretonianum (14147)
Ludfordian - Pridoli425.6 - 419.2United Kingdom (Wales) Renalia sp., Steganotheca striata (10635)
Ludfordian - Silurian425.6 - 419.2United Kingdom (Wales) Steganotheca striata (10634)
Pridoli423.0 - 419.2United Kingdom (England) Rhyniopsida indet. (11218)
Pridoli423.0 - 419.2Ukraine Zosterophyllum sp. (13873)
Pridoli423.0 - 419.2Ukraine (Podolia) Zosterophyllum myretonianum (52702)
Pridoli423.0 - 419.2China (Xinjiang) Zosterophyllum sp. (10931)
Pridoli - Helderbergian423.0 - 393.3China (Xinjiang) Zosterophyllum sp. (27917)
Pridoli - Helderbergian423.0 - 393.3Australia (New South Wales) Tracheophyta indet. (24903)
Lochkovian419.2 - 410.8Australia (New South Wales) Tracheophyta indet. (24907)
Lochkovian419.2 - 410.8Belgium (Liege?) Zosterophyllum fertile, Zosterophyllum spp. (11722)
Lochkovian419.2 - 410.8United Kingdom (England) Rhyniopsida indet. (11112) Zosterophyllum fertile (10679)
Lochkovian419.2 - 410.8United Kingdom (Scotland) Zosterophyllum fertile (10705) Zosterophyllum myretonianum (26755 26756 26757)
Lochkovian419.2 - 410.8United Kingdom (Wales) Zosterophyllum fertile (10997)
Devonian - Early/Lower Devonian419.2 - 393.3Canada (New Brunswick) Pertica dalhousii (23007)
Early/Lower Devonian419.2 - 393.3China (Sichuan) Zosterophyllum yunnanicum, Zosterophyllum sichuanense, Zosterophyllum myretonianum (10907)