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Hymenoptera
Taxonomy
Hymenoptera was named by Linnaeus (1758). It is extant.
It was assigned to Insecta by Linnaeus (1758), Schreber (1759), Linnaeus (1761), Müller (1764), Linnaeus (1767), Strøm (1768), Drury (1770), Forster (1771), Müller (1776), Harris (1780), Latreille (1796), Cockerell (1921) and Hong and Wang (1985); to Endoneoptera by Kukalová-Peck and Lawrence (2004); to Endopterygota by Carpenter (1992) and Beutel and Pohl (2006); to Holometabola by Misof et al. (2014); and to Hymenopterida by Engel (2022).
It was assigned to Insecta by Linnaeus (1758), Schreber (1759), Linnaeus (1761), Müller (1764), Linnaeus (1767), Strøm (1768), Drury (1770), Forster (1771), Müller (1776), Harris (1780), Latreille (1796), Cockerell (1921) and Hong and Wang (1985); to Endoneoptera by Kukalová-Peck and Lawrence (2004); to Endopterygota by Carpenter (1992) and Beutel and Pohl (2006); to Holometabola by Misof et al. (2014); and to Hymenopterida by Engel (2022).
Synonyms
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Synonymy list
Year | Name and author |
---|---|
1758 | Hymenoptera Linnaeus p. 341 |
1759 | Hymenoptera Schreber p. 14 |
1761 | Hymenoptera Linnaeus p. 385 |
1763 | Aculeata Scopoli p. 273 |
1764 | Hymenoptera Müller p. 67 |
1767 | Hymenoptera Linnaeus p. 917 |
1768 | Hymenoptera Strøm p. 344 |
1770 | Hymenoptera Drury |
1771 | Hymenoptera Forster p. 77 |
1776 | Hymenoptera Müller p. 147 |
1780 | Hymenoptera Harris p. 95 |
1796 | Hymenoptera Latreille p. 105 |
1798 | Piezata Fabricius p. 209 |
1836 | Cleptis stenstrupii Beck p. 219 |
1867 | Symphyta Gerstaecker p. 2 |
1897 | Chalastogastra Konow |
1921 | Hymenoptera Cockerell |
1923 | Chalastogastra Meunier p. 85 |
1937 | Pseudisobrachium oligocenicum Theobald pp. 397, 398 figs. Pl XXVIII fig 16 |
1955 | Symphyta Riek p. 656 |
1980 | Siricomorpha Rasnitsyn |
1983 | Mesopamphila Hong |
1985 | Symphyta Hong and Wang p. 7 |
1985 | Hymenoptera Hong and Wang p. 16 |
1985 | Shandongodes necrodes Zhang p. 35 figs. Pl 6, fig 28; text-fig 17 |
1988 | Pseudisobrachium oligocenicum Sorg p. 74 |
1992 | Hymenoptera Carpenter |
1992 | Symphyta Carpenter |
2000 | Symphyta Schlüter p. 76 |
2004 | Hymenoptera Kukalová-Peck and Lawrence |
2006 | Hymenoptera Beutel and Pohl |
2012 | Pseudisobrachium oligocenicum Azevedo and Azar p. 217 |
2014 | Hymenoptera Misof et al. |
2015 | Siricomorpha Archibald and Rasnitsyn |
2018 | Pseudisobrachium oligocenicum Azevedo et al. p. 125 |
2022 | Hymenoptera Engel |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Or. Hymenoptera Linnaeus 1758
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Unr. Aculeati Latreille 1802
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Unr. Nemoglossata Latreille 1802
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Fm. Apiariae Latreille 1802
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Invalid names: Apes Latreille 1802 [empty], Bombi Latreille 1802 [empty], Ceratinae Latreille 1802 [empty], Eucerae Latreille 1802 [empty], Megachiles Latreille 1802 [empty], Parasitae Latreille 1802 [empty], Xylocopae Latreille 1802 [empty]
Unr. Platiglossata Latreille 1802
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Invalid names: Andrenetae Latreille 1802 [empty], Crabronites Latreille 1802 [empty], Formicariae Latreille 1802 [empty], Philantores Latreille 1802 [empty], Scolietae Latreille 1802 [empty], Sphegimae Latreille 1802 [empty]
Superfm. Anaxyeloidea Martynov 1925
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Fm. Anaxyelidae Martynov 1925
+
Subfm. †Anaxyelinae Martynov 1925
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G. †Anaxyela Martynov 1925
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†Anaxyela destructa Rasnitsyn 1969
†Anaxyela nana Rasnitsyn 1968
†Anaxyela parvula Rasnitsyn 1963
G. †Brachysyntexis Rasnitsyn 1969
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†Brachysyntexis acuta Kopylov et al. 2020
†Brachysyntexis brachyura Rasnitsyn 1968
†Brachysyntexis brevicornis Kopylov et al. 2020
†Brachysyntexis laticella Kopylov et al. 2020
†Brachysyntexis micrura Rasnitsyn 1969
†Brachysyntexis minuta Kopylov et al. 2020
†Brachysyntexis nova Rasnitsyn 1969
†Brachysyntexis robusta Zhang and Rasnitsyn 2006
†Brachysyntexis rohweri Kopylov et al. 2020
†Brachysyntexis sinensis Kopylov et al. 2020
†Brachysyntexis tenebrosa Kopylov 2018
†Brachysyntexis tigris Kopylov 2018
G. †Kulbastavia Rasnitsyn 1968
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†Kulbastavia grandis Kopylov 2018
†Kulbastavia macrura Rasnitsyn 1963
G. †Sphenosyntexis Rasnitsyn 1969
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†Sphenosyntexis antonovi Rasnitsyn 1963
†Sphenosyntexis pallicornis Rasnitsyn 1969
G. †Syntexyela Rasnitsyn 1968
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†Syntexyela asiatica Rasnitsyn 1968
†Syntexyela continentalis Zhang and Rasnitsyn 2006
†Syntexyela gracilicornis Rasnitsyn 1968
†Syntexyela inversa Rasnitsyn 1968
†Syntexyela media Rasnitsyn 1963
G. †Urosyntexis Rasnitsyn 1969
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†Urosyntexis depressa Rasnitsyn 1969
†Urosyntexis drepanura Rasnitsyn 1968
†Urosyntexis magna Rasnitsyn 1968
†Urosyntexis undosa Kopylov 2019
Invalid names: Dolichostigmatinae Rasnitsyn 1968 [synonym], Kempendajinae Rasnitsyn 1980 [synonym]
Subfm. Syntexinae Benson 1935
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G. †Cretosyntexis Rasnitsyn and Martínez-Delclòs 2000
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†Cretosyntexis montsecensis Rasnitsyn and Martínez-Delclòs 2000
G. †Eosyntexis Rasnitsyn 1990
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†Eosyntexis catalonicus Rasnitsyn and Martínez-Delclòs 2000
†Eosyntexis parva Ortega-Blanco et al. 2008
†Eosyntexis senilis Rasnitsyn 1990
†Eosyntexis tuffinae Rasnitsyn and Jarzembowski 1998
G. †Orthosyntexis Gao et al. 2021
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†Orthosyntexis elegans Gao et al. 2021
†Orthosyntexis thanti Gao et al. 2021
Invalid names: Paraxiphydriinae Gao et al. 2022 [synonym]
Fm. †Bryopompilidae Engel and Grimaldi 2006
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Invalid names: Bryopompilidae Rodriguez et al. 2016 [objective synonym]
Superfm. Pamphilioidea Cameron 1890 [sawfly]
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Fm. Megalodontesidae Konow 1897
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Subfm. †Archoxyelydinae Wang et al. 2013
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Subfm. †Decorisiricinae Wang et al. 2015
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G. †Decorisiricius Wang et al. 2015
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†Decorisiricius longus Wang et al. 2015
†Decorisiricius patulus Wang et al. 2015
G. †Limbisiricius Wang et al. 2015
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†Limbisiricius aequalis Wang et al. 2015
†Limbisiricius complanatus Wang et al. 2015
G. †Jibaissodes Ren 1995
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†Jibaissodes bellus Gao et al. 2016
†Jibaissodes giganteus Ren 1995
†Jibaissodes peichenae Wang et al. 2019
Subfm. †Praesiricinae Rasnitsyn 1968
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Invalid names: Praesiricidae Rasnitsyn 1968 [synonym]
Fm. †Mirolydidae Wang et al. 2017
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Fm. Pamphiliidae Cameron 1890
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Subfm. Cephalciinae Benson 1945
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G. Acantholyda Costa 1894
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Subg. Acantholyda (Acantholyda) Costa 1894
†Acantholyda ribesalbesensis Peñalver and Arillo 2002
G. Cephalcia Jurine 1801
Invalid names: Cephalciini Benson 1945 [invalid subgroup]
G. Cephaleia Jurine 1801
Subfm. Juralydinae Rasnitsyn 1977
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G. †Dolicholyda Zhuang et al. 2022
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†Dolicholyda angusta Zhuang et al. 2022
†Dolicholyda confluens Zhuang et al. 2022
†Dolicholyda obtusata Zhuang et al. 2022
G. †Scabolyda Wang et al. 2014
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†Scabolyda incompleta Wang et al. 2014
†Scabolyda latusa Zhuang et al. 2022
†Scabolyda orientalis Wang et al. 2014
†Scabolyda tenuis Zhuang et al. 2022
Subfm. Pamphiliinae Cameron 1890
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G. Neurotoma Konow 1897 [Leaf-rolling sawfly]
G. Pamphilius Latreille 1802
Invalid names: Neurotomini Benson 1945 [invalid subgroup]
Invalid names: Lydites Newman 1834 [replaced]
G. †Prolyda Rasnitsyn 1968
+
†Prolyda depressa Rasnitsyn 1969
†Prolyda dimidia Wang et al. 2016
†Prolyda elegantula Wang et al. 2016
†Prolyda karatavica Rasnitsyn 1968
†Prolyda xyelocera Rasnitsyn 1968
Fm. †Xyelydidae Rasnitsyn 1968
+
G. †Ferganolyda Rasnitsyn 1983
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†Ferganolyda charybdis Rasnitsyn et al. 2006
†Ferganolyda chungkuei Rasnitsyn et al. 2006
†Ferganolyda cubitalis Rasnitsyn 1983
†Ferganolyda eucalla Wang et al. 2015
†Ferganolyda insolita Wang et al. 2015
†Ferganolyda radialis Rasnitsyn 1983
†Ferganolyda scylla Rasnitsyn et al. 2006
†Ferganolyda sogdiana Rasnitsyn 1983
G. †Fissilyda Wang et al. 2015
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†Fissilyda alba Wang et al. 2015
†Fissilyda compta Wang et al. 2015
†Fissilyda parilis Wang et al. 2015
G. †Medilyda Wang and Rasnitsyn 2016
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†Medilyda distorta Wang and Rasnitsyn 2016
†Medilyda procera Wang and Rasnitsyn 2016
G. †Mesolyda Rasnitsyn 1963
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†Mesolyda jurassica Rasnitsyn 1963
†Mesolyda lingyuanensis Ren 1995
†Mesolyda sibirica Rasnitsyn 1983
Invalid names: Alloxyelula Ren 1995 [synonym]
G. †Novalyda Gao et al. 2013
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†Novalyda cretacica Gao et al. 2013
†Novalyda decora Wang et al. 2015
G. †Sagulyda Rasnitsyn 1983
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†Sagulyda arcuata Rasnitsyn 1983
†Sagulyda ferganica Rasnitsyn 1983
†Sagulyda magna Rasnitsyn 1983
G. †Strenolyda Wang and Rasnitsyn 2016
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†Strenolyda marginalis Wang and Rasnitsyn 2016
†Strenolyda retrorsa Wang and Rasnitsyn 2016
G. †Strophandria Rasnitsyn 1968
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†Strophandria grossa Rasnitsyn 1968
†Strophandria moderata Rasnitsyn 1983
Invalid names: Megalodontoidea Konow 1897 [synonym]
Superfm. Tenthredinoidea Latreille 1802 [wood wasp]
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Fm. Argidae Konow 1890 [argid sawfly]
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Subfm. Arginae Konow 1890
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Subfm. Sterictiphorinae Rohwer 1911
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G. Didymia Lepeletier and Serville 1828
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†Didymia davisi Smith 1992
†Didymia dominicana Smith 1992
†Didymia ebena Smith 1992
†Didymia poinari Smith 1992
†Didymia protera Smith 1992
G. Sterictiphora Billberg 1820
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†Sterictiphora konowi Rohwer 1908
Invalid names: Schizocerus Berthold 1827 [synonym]
Fm. Athaliidae Rohwer 1911
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Fm. Blasticotomidae Thomson 1871
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G. Blasticotoma Klug 1834
G. Runaria Malaise 1931
Invalid names: Xyelotomidae Rasnitsyn 1968 [synonym]
Fm. Cimbicidae Kirby 1837 [cimbicid sawfly]
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Subfm. Abiinae Thomson 1871
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G. Abia Leach 1817
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Abia lonicerae Linnaeus 1758
†Abia maculosa Zhang 1989
†Abia paurocephala Zhang 1989
†Abia shandongensis Hong and Wang 1985
†Abia shanwangensis Hong 1984
Invalid names: Procimbex Hong 1984 [synonym], Zaraea Leach 1817 [synonym]
Subfm. †Cenocimbicinae Archibald and Rasnitsyn 2023
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G. †Allenbycimbex Archibald and Rasnitsyn 2023
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†Allenbycimbex morrisae Archibald and Rasnitsyn 2023
G. †Leptostigma Archibald and Rasnitsyn 2023
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†Leptostigma alaemacula Archibald and Rasnitsyn 2023
†Leptostigma brevilatum Archibald and Rasnitsyn 2023
†Leptostigma fasciatum Archibald and Rasnitsyn 2023
†Leptostigma longiclava Archibald and Rasnitsyn 2023
†Leptostigma longipallidum Archibald and Rasnitsyn 2023
†Leptostigma longitenebricum Archibald and Rasnitsyn 2023
†Leptostigma proxivena Archibald and Rasnitsyn 2023
Subfm. Cimbicinae Kirby 1837
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G. Cimbex Olivier 1790
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†Cimbex chromoptera Zhang 1989
†Cimbex miocenica Riou 1992
†Cimbex turoliana Riou 1992
Invalid names: Clavellaria Olivier 1789 [replaced], Clavellarius Olivier 1789 [invalid subgroup]
G. Pseudoclavellaria Schultz 1906
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†Pseudoclavellaria autochthona Zhang 1989
†Pseudoclavellaria bicolor Zhang 1989
†Pseudoclavellaria longiclava Zhang 1989
†Pseudoclavellaria molpa Zhang et al. 1994
G. †Sinocimbex Zhang et al. 1994
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†Sinocimbex pellucidus Zhang et al. 1994
†Sinocimbex silaceus Zhang et al. 1994
Subfm. Corynidinae Benson 1938
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Subfm. Pachylostictinae Benson 1938
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Fm. Diprionidae Rohwer 1910 [conifer sawfly]
+
Subfm. Diprioninae Rohwer 1910
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G. Diprion Schrank 1802
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Invalid names: Lophyrus Latreille 1802 [synonym], Pteronus Jurine 1801 [invalid subgroup]
Subfm. Monocteninae MacGillivray 1906
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Fm. †Electrotomidae Rasnitsyn 1977
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G. †Electrotoma Rasnitsyn 1977
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†Electrotoma kopylovi Rasnitsyn and Manukyan 2023
†Electrotoma succini Rasnitsyn 1977
Fm. Tenthredinidae Latreille 1802
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Subfm. Allantinae Newman 1834
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Tr. Empriini Rohwer 1911
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G. Empria Lepeletier de Saint-Fargeau and Audinet-Serville 1828
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Invalid names: Leucempria Enslin 1913 [synonym]
G. Eriocampa Hartig 1837
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†Eriocampa bruesi Rohwer 1908
†Eriocampa celata Cockerell 1915
†Eriocampa disjecta Cockerell 1922
†Eriocampa pristina Cockerell 1910
†Eriocampa scudderi Brues 1908
†Eriocampa synthetica Cockerell 1911
†Eriocampa tulameenensis Rice 1968
G. †Hylotomites Meunier 1914
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G. †Palaeotaxonus Brues 1908
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†Palaeotaxonus trivittatus Rohwer 1908
†Palaeotaxonus typicus Brues 1908
†Palaeotaxonus vetus Cockerell 1917
G. Pseudosiobla Ashmead 1898
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†Pseudosiobla campbelli Rice 1968
†Pseudosiobla megoura Cockerell 1907
†Pseudosiobla misera Brues 1908
G. †Rovnotaxonus Vilhelmsen and Perkovsky 2024
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†Rovnotaxonus aristovi Vilhelmsen and Perkovsky 2024
Subfm. Heterarthrinae Benson 1952
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G. Caliroa Costa 1859
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†Caliroa micrarche Cockerell 1916
†Caliroa mima Cockerell 1917
†Caliroa revelata Cockerell 1909
Invalid names: Eriocampoides Konow 1890 [synonym]
G. Fenusa Leach 1817
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†Fenusa parva Brues 1906
†Fenusa primula Rohwer 1908
Invalid names: Lithoryssus Brues 1906 [synonym]
Subfm. Nematinae
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G. †Florissantinus Zhelochovtzev and Rasnitsyn 1972
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†Florissantinus angulatus Zhelochovtzev and Rasnitsyn 1972
G. Mesoneura Hartig 1837
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†Mesoneura vexabilis Brues 1908
Invalid names: Lisconeura Rohwer 1908 [synonym]
G. Nematus Panzer 1801
Invalid names: Hoplocampini Konow 1890 [invalid subgroup]
G. Rhogogaster Konow 1884
Subfm. Selandriinae Thomson 1871
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Subfm. Tenthredininae Latreille 1802
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G. Tenthredo Linnaeus 1758
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Subg. Tenthredo (Tenthredella) Rohwer 1910
†Tenthredo avia Brues 1908
†Tenthredo infossa Brues 1908
†Tenthredo meunieri Taeger et al. 2009
+
Invalid names: Tenthredo fasciata Meunier 1922 [replaced], Tenthredo gracilis Statz 1936 [replaced]
†Tenthredo miocenica Zhang and Zhang 1990
†Tenthredo petrae Zhang and Zhang 1990
†Tenthredo primordialis Piton 1940
†Tenthredo rhysia Zhang 1989
†Tenthredo saxorum Rohwer 1908
†Tenthredo submersa Cockerell 1907
Invalid names: Tenthredo gervaisi Heer 1861 [nomen nudum]
Invalid names: Tenthredinini Latreille 1802 [empty]
Invalid names: Nematus ellipticus Fritsch 1901 [nomen dubium]
Invalid names: Pteronus prodigus Brues 1908 [nomen dubium]
Unr. Terebrantes Latreille 1802
+
Unr. Ventropedonculati Latreille 1802
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Fm. Ichneumonides Latreille 1802 [ichneumon wasp]
+
Invalid names: Ichneumonides Latreille 1802 [empty], Sphegii Latreille 1802 [empty]
Invalid names: Cleptiosa Latreille 1802 [empty]
Unr. Ventrosessiles Latreille 1802
+
Invalid names: Tenthredines Latreille 1802 [empty], Urocerata Latreille 1802 [empty]
Unr. Unicalcarida Schulmeister et al. 2002
+
Subor. Apocrita Gerstaecker 1867 [wasp]
+
Unr. Aculeata Latreille 1802
+
Superfm. Apoidea Latreille 1802 [apoid wasp]
+
Fm. †Allommationidae Rosa and Melo 2021
Fm. Ampulicidae Shuckard 1840 [cockroach wasp]
Unr. Anthophila Latreille 1804 [bee]
Fm. Astatidae Lepeletier 1845
Fm. †Burmasphecidae Rosa and Melo 2023
Fm. †Cirrosphecidae Antropov 2000
Fm. Crabronidae Latreille 1802
Fm. Dasypodaidae Sagemehl 1882
Fm. Heterogynaidae Nagy 1969
Fm. Pemphredonidae Dahlbom 1835 [aphid wasp]
Fm. Philanthidae Latreille 1802
Fm. Psenidae Costa 1858
Fm. Sphecidae Latreille 1802 [digger wasp]
Fm. †Spheciellidae Rosa and Melo 2021
Fm. †Temnogynidae Rosa and Melo 2024
G. †Trigampulex Antropov 2000
Invalid names: Ammoplanidae Evans 1959 [empty], Bembicidae Latreille 1802 [empty], Sphecoidea Latreille 1802 [synonym]
Superfm. †Bethylonymoidea Rasnitsyn 1975
+
Fm. †Bethylonymidae Rasnitsyn 1975
Fm. †Trifionychidae Lepeco and Melo 2022
Superfm. Chrysidoidea Latreille 1802
+
Fm. Chrysididae Latreille 1802 [cuckoo wasp]
Fm. †Chrysobythidae Melo and Lucena 2019
Fm. Dryinidae Haliday 1833
Fm. Embolemidae Forster 1856
Fm. †Plumalexiidae Brothers 2011
Fm. †Radiophronidae Ortega-Blanco et al. 2010
Fm. Sclerogibbidae Ashmead 1902 [sclerogibbid wasp]
Fm. Scolebythidae Evans 1963
Invalid names: Bethyloidea [synonym], Plumariidae [empty]
G. †Dolichomyrma Dlussky 1975
+
†Dolichomyrma latipes Dlussky 1975
†Dolichomyrma longiceps Dlussky 1975
Superfm. Formicoidea Latreille 1802
+
G. †Camelomecia Barden and Grimaldi 2016
G. †Camelosphecia Boudinot et al. 2020
Fm. Formicidae Latreille 1802 [ant]
Superfm. Pompiloidea Latreille 1804
+
Fm. †Burmusculidae Zhang and Rasnitsyn 2018
Fm. Myrmosidae Fox 1894
Fm. Sapygidae Latreille 1810
Superfm. Scolioidea Latreille 1802
+
G. †Oryctopterus Rasnitsyn 1975
Fm. Scoliidae Latreille 1802 [hairy flower wasp]
Invalid names: Bradynobaenidae Saussure 1892 [empty]
Fm. †Stigmaphronidae Kozlov 1975
+
G. †Allocotidus Muesebeck 1963
G. †Aphrostigmon Rasnitsyn 1991
G. †Burmaphron Engel and Grimaldi 2009
G. †Elasmomorpha Kozlov 1975
G. †Elasmophron Engel and Grimaldi 2009
G. †Hippocoon Kozlov 1975
G. †Libanophron Engel and Grimaldi 2009
G. †Stigmaphron Kozlov 1975
G. †Tagsmiphron Engel and Grimaldi 2009
Superfm. Vespoidea Latreille 1802
+
Fm. †Falsiformicidae Rasnitsyn 1975
Fm. Rhopalosomatidae
Fm. Tiphiidae Leach 1815
Fm. Vespidae Latreille 1802
Invalid names: Bocchus cenomanianus Olmi et al. 2010 [nomen dubium], Bombus pusillus Menge 1856 [nomen dubium]
G. †Apocritites Rasnitsyn and Jarzembowski 1998
+
†Apocritites distinctus Rasnitsyn and Jarzembowski 1998
Fm. †Armaniidae Dlussky 1983
+
G. †Archaeopone Dlussky 1975
+
†Archaeopone kzylzharica Dlussky 1975
†Archaeopone taylori Dlussky 1983
G. †Armania Dlussky 1983
+
†Armania capitata Dlussky 1999
†Armania curiosa Dlussky 1983
†Armania pristina Dlussky 1999
†Armania robusta Dlussky 1983
Invalid names: Armaniella Dlussky 1983 [synonym]
G. †Orapia Dlussky et al. 2004
+
†Orapia minor Dlussky et al. 2004
†Orapia rayneri Dlussky et al. 2004
G. †Pseudarmania Dlussky 1983
+
†Pseudarmania aberrans Dlussky 1983
†Pseudarmania rasnitsyni Dlussky 1983
Infraor. Brachycleistogastromorpha Kovalev 1994
+
Invalid names: Cleistogastroidea Rasnitsyn 1975 [empty]
Superfm. Ceraphronoidea Haliday 1833
+
Fm. Ceraphronidae Haliday 1833
+
G. Aphanogmus Thomson 1858
G. Ceraphron Jurine 1807
Invalid names: Ceraphronini Haliday 1833 [empty]
Fm. Megaspilidae Ashmead 1893
+
G. Lagynodes Forster 1841
Subfm. Lagynodinae Masner and Dessart 1967
Subfm. Megaspilinae Ashmead 1893
Invalid names: Calliceratidae [synonym]
Superfm. Evanioidea Latreille 1802
+
Unr. Neoevanioides Engel 2006
+
Fm. †Anomopterellidae Rasnitsyn 1975
Unr. Aulaciformes Grimaldi and Engel 2005
Unr. Evaniiformes Grimaldi and Engel 2005
Unr. †Protoevanioides Jouault et al. 2021
+
Fm. †Othniodellithidae Engel and Huang 2016
Fm. †Praeaulacidae Rasnitsyn 1972
Invalid names: Asiapterites Hong 2002 [nomen nudum], Euryopterites Hong 2002 [nomen nudum], Oblongigastrula Hong 2002 [nomen nudum]
Superfm. Ichneumonoidea Latreille 1802
+
Fm. Braconidae Nees ab Esenbeck 1811
+
Subfm. Agathidinae Haliday 1833
G. Aivalykus Nixon 1938
G. Alysia Latreille 1804
Subfm. Alysiinae Leach 1815
Subfm. Aphidiinae Haliday 1833
Subfm. Apozyginae Mason 1978
G. Austrohelcon Turner 1918
Subfm. Betylobraconinae Tobias 1979
Subfm. Braconinae Nees ab Esenbeck 1811
Subfm. Cardiochilinae Ashmead 1900
Subfm. Charmontinae van Achterberg 1979
Subfm. Cheloninae Nees 1816
G. Coeloreuteus Roman 1910
G. †Digastrotheca Brues 1933
Subfm. †Diospilitinae Tobias 1987
Subfm. Doryctinae Förster 1862
G. †Electrohelcon Brues 1933
G. †Eobracon Cockerell 1921
Subfm. Euphorinae Förster 1862
Subfm. Exothecinae Förster 1862
Subfm. Helconinae Förster 1862
Subfm. Homolobinae van Achterberg 1979
G. Hormiopterus Giraud 1869
Subfm. Ichneutinae Förster 1862
G. Iphiaulax Förster 1862
Subfm. Masoninae van Achterberg 1995
Subfm. †Megalyrhyssalinae Belokobylskij and Jouault 2021
Subfm. Microgastrinae Förster 1862
Subfm. Microtypinae Szepligeti 1908
Subfm. Miracinae
Subfm. Opiinae Blanchard 1845
G. Opius Wesmael 1835
Subfm. Orgilinae Ashmead 1900
G. Phanomeris Förster 1862
Subfm. †Protobraconinae van Achterberg and Chen 2020
Subfm. †Protorhyssalinae Basibuyuk et al. 1999
G. †Pyramidibracon van Achterberg and Chen 2020
G. Rhaconotus Ruthe 1854
Subfm. Rhyssalinae Förster 1862
Subfm. Rogadinae Förster 1862
G. Semirhytus Szepligeti 1902
Subfm. Sigalphinae Haliday 1833
G. Sigalphus Latreille 1802
G. †Sinobracon Hong 1974
G. Snellenius Westwood 1882
G. †Stephanorhyssalus Belokobylskij and Jouault 2021
G. †Vertibracon van Achterberg and Chen 2023
G. Xenarcha Förster 1862
Invalid names: Eoichneumonidae Jell and Duncan 1986 [synonym], Fushunobracon Hong 2002 [nomen nudum], Microgasterinae Förster 1862 [empty], Seneciobraconinae Engel and Huang 2018 [empty], Xuaulacites Hong 2002 [nomen nudum]
Fm. Ichneumonidae Latreille 1802 [ichneumon wasp]
+
G. Absyrtus Holmgren 1858
G. Acoenitus Latreille 1809
G. †Carinibus Spasojevic et al. 2018
G. †Eopimpla Cockerell 1921
G. †Epicharopimpla Zhang et al. 1994
Subfm. Hybrizoninae Blanchard 1845
+
Invalid names: Ghilarovitinae Kasparyan 1988 [synonym], Paxylommatinae Förster 1862 [synonym]
Unr. Ichneumoniformes
G. †Ichneumonites Heer 1867
Unr. Labeniformes
Subfm. †Labenopimplinae Kopylov 2010
G. Lapton Nees 1815
G. †Lithapechtis Cockerell 1921
Subfm. Lycorininae Cushman and Rohwer 1920
G. †Megatryphon Cockerell 1924
G. †Mesopimpla Cockerell 1919
G. †Mesornatus Spasojevic et al. 2018
Subfm. Microleptinae Townes 1958
G. †Miolyta Zhang 1989
Subfm. †Novichneumoninae Li et al. 2016
Unr. Ophioniformes Gauld 1985
Subfm. Orthopelmatinae Schmiedeknecht 1910
Subfm. †Palaeoichneumoninae Kopylov 2009
G. †Paratilgidopsis Martins-Neto 1998
Subfm. †Pherhombinae Kasparyan 1988
Unr. Pimpliformes Wahl 1990
G. †Polychorella Zhang 1985
Subfm. Stilbopinae Townes and Townes 1949
Subfm. †Tanychorinae Rasnitsyn 1980
G. †Taubatehymen Martins-Neto 1998
G. †Tilgidopsis Cockerell 1921
Subfm. †Townesitinae Kasparyan 1994
G. †Trymectus Spasojevic et al. 2018
G. †Tryphopimpla Kopylov 2010
Unr. Xoridiformes
Invalid names: Eucerotinae Seyrig 1934 [empty], Ichneumonidae gabbroensis Handlirsch 1907 [nomen dubium], Liaoichneumon Hong 2002 [nomen nudum], Oxytorinae Thomson 1883 [empty], Polysphincta atavina Cockerell 1921 [nomen dubium]
G. †Rasnitsynia Pagliano and Scaramozzino 1990
+
†Rasnitsynia huadongensis Zhang 1985
Invalid names: Oligoneuroides Zhang 1985 [replaced]
Fm. †Khutelchalcididae Rasnitsyn et al. 2004
+
Fm. †Kuafuidae Rasnitsyn and Zhang 2010
+
Superfm. Megalyroidea Schletterer 1889
+
Fm. Megalyridae Schletterer 1889
+
Subfm. Megalyrinae Schletterer 1889
Subfm. †Megazarinae Perrichot 2009
Invalid names: Dinapsini Waterston 1922 [invalid subgroup]
Superfm. †Panguoidea Li et al. 2019
+
Infraor. Proctotrupomorpha
+
Superfm. Cynipoidea Latreille 1802
+
Fm. †Archaeocynipidae Rasnitsyn and Kovalev 1988
Fm. Cynipidae Latreille 1802
Fm. Figitidae Forster 1869
Fm. †Gerocynipidae Liu and Engel 2007
Fm. Ibaliidae Forster 1869
Fm. Liopteridae Ashmead 1895
Fm. †Protimaspidae Liu and Engel 2007
Fm. †Stolamissidae Liu and Engel 2007
Invalid names: Archaeocynipoidea Rasnitsyn and Kovalev 1988 [synonym], Diplolepididae Latreille 1802 [empty]
Unr. Microprocta Rasnitsyn and Öhm-Kühnle 2020
+
Unr. Bipetiolarida Engel 2005
Superfm. Chalcidoidea Leach 1815 [chalcid wasp]
Fm. †Chalscelionidae Rasnitsyn and Öhm-Kühnle 2020
Superfm. Diaprioidea Haliday 1833
Fm. †Jurapriidae Rasnitsyn 1983
Superfm. Platygastroidea Haliday 1833
Fm. †Trupochalcididae Kozlov 1975
+
Invalid names: Cretacoformicidae Rasnitsyn and Öhm-Kühnle 2019 [synonym]
Superfm. Proctotrupoidea Latreille 1802
+
G. †Amitchellia Rasnitsyn and Jarzembowski 1998
Fm. Heloridae Forster 1856
Fm. †Mesoserphidae Kozlov 1970
G. †Paraulacus Ping 1928
Fm. †Peleserphidae Zhang et al. 2017
Fm. Peradeniidae Naumann and Masner 1985
Fm. Roproniidae Viereck 1916
Invalid names: Austroniidae Kozlov 1975 [empty], Fushunoxyela Hong 2002 [nomen nudum], Hemerocynipites Hong 2002 [nomen nudum], Monomachidae Ashmead 1902 [empty], Sinocynipites Hong 2002 [nomen nudum]
Invalid names: Cynipomorpha Kovalev 1994 [synonym]
Superfm. Stephanoidea Leach 1815
+
Fm. †Ephialtitidae Handlirsch 1906
+
Subfm. †Ephialtitinae Handlirsch 1906
G. †Kotaphialtites Rasnitsyn 2008
Subfm. †Symphytopterinae Rasnitsyn 1980
Invalid names: Karataidae Rasnitsyn 1977 [synonym]
Fm. †Ohlhoffiidae Jouault et al. 2020
+
G. †Cretephialtites Rasnitsyn and Ansorge 2000
G. †Myanmephialtites Jouault et al. 2020
G. †Ohlhoffia Jouault et al. 2020
Fm. Stephanidae Leach 1815 [crown wasp]
+
G. †Aphanostephanus Ge and Tan 2023
Subfm. †Lagenostephaninae Ge and Tan 2023
G. †Phoriostephanus Engel and Huang 2016
Subfm. Schlettereriinae Orfila 1949
G. Stephanus Panzer 1805
G. †Tichostephanus Engel 2019
Invalid names: Eostephanitidae Hong 2002 [invalid subgroup], Ephialtitoidea Handlirsch 1906 [synonym]
Superfm. Trigonalyoidea Cresson 1887
+
Fm. Trigonalidae Cresson 1887
+
G. †Darbigonalus Rasnitsyn 1986
G. †Eotrigonalis Poinar 2005
G. Trigonalys Westwood 1835
Invalid names: Apiaria lapidea Germar 1842 [nomen dubium], Asiacynipites Hong 2002 [nomen nudum], Dendrocerus dubitatus Brues 1937 [nomen dubium], Eucynipites Hong 2002 [nomen nudum], Evaniomorpha Rasnitsyn 1988 [invalid subgroup], Stephanomorpha Leach 1815 [invalid subgroup]
Superfm. Cephoidea Newman 1834 [stem sawfly]
+
Fm. Cephidae Newman 1834 [stem sawfly]
+
Subfm. Cephinae Newman 1834
+
G. Cephus Latreille 1802
G. †Electrocephus Konow 1897
G. Janus Stephens 1835
G. †Mesocephus Rasnitsyn 1968
Fm. †Sepulcidae Rasnitsyn 1968
+
Subfm. †Ghilarellinae Rasnitsyn 1988
+
G. †Ghilarella Rasnitsyn 1988
G. †Meiaghilarella Rasnitsyn and Martínez-Delclòs 2000
Subfm. †Parapamphiliinae Rasnitsyn 1968
+
G. †Micramphilius Rasnitsyn 1993
G. †Pamparaphilius Rasnitsyn 1993
G. †Parabakharius Rasnitsyn 1993
G. †Parapamphilius Rasnitsyn 1968
G. †Shurabisca Rasnitsyn 1968
G. †Yananphilius Jouault 2024
Subfm. †Trematothoracinae Rasnitsyn 1988
+
G. †Prosyntexis Sharkey 1990
G. †Thoracotrema Rasnitsyn 1988
Subfm. †Xyelulinae Rasnitsyn 1993
+
G. †Neoxyelula Rasnitsyn 1993
G. †Onokhoius Rasnitsyn 1990
G. †Xyelula Rasnitsyn 1969
Superfm. Orussoidea Newman 1834 [wood wasp]
+
Fm. †Burmorussidae Zhang et al. 2020
+
Fm. Orussidae Newman 1834 [parasitic wood wasp]
+
Fm. †Paroryssidae Martynov 1925
+
G. †Microryssus Rasnitsyn 1968
+
†Microryssus antennatus Rasnitsyn 1968
†Microryssus brachyurus Rasnitsyn 1968
†Microryssus crassipes Rasnitsyn 1968
†Microryssus minus Rasnitsyn 1968
†Microryssus robustus Rasnitsyn 1968
†Microryssus subtilis Rasnitsyn 1968
G. †Praeoryssus Rasnitsyn 1968
+
†Praeoryssus gracilis Rasnitsyn 1968
†Praeoryssus venosus Rasnitsyn 1968
Invalid names: Karatavitoidea Rasnitsyn 1963 [synonym]
Superfm. Siricoidea Billberg 1820
+
Fm. †Protosiricidae Rasnitsyn and Zhang 2004
+
Fm. †Pseudosiricidae Handlirsch 1906
+
G. †Myrmicium Westwood 1854
+
†Myrmicium araripterum Freitas et al. 2020
†Myrmicium elongatum Germar 1839
†Myrmicium heerii Westwood 1854
†Myrmicium nanus Handlirsch 1906
†Myrmicium schroeteri Germar 1839
+
Invalid names: Pseudosirex antiquus Germar 1839 [synonym], Pseudosirex compressus Oppenheim 1885 [synonym], Pseudosirex darwini Weyenbergh 1873 [synonym], Pseudosirex deichmuelleri Handlirsch 1906 [synonym], Pseudosirex elegans Oppenheim 1885 [synonym], Pseudosirex gracilis Oppenheim 1885 [synonym], Pseudosirex karschi Oppenheim 1885 [synonym], Pseudosirex minimus Oppenheim 1885 [synonym], Pseudosirex separatus Handlirsch 1906 [synonym], Rhipidorhabdus schroeteri Oppenheim 1885 [synonym], Sphinx snelleni Weyenbergh 1869 [synonym]
Invalid names: Apiaria Germar 1839 [replaced], Fabellovena Oppenheim 1885 [synonym], Hagenia Weyenbergh 1869 [synonym], Pseudosirex Weyenbergh 1873 [synonym], Rhipidorhabdus Oppenheim 1885 [synonym]
Invalid names: Myrmiciidae Maa 1949 [synonym]
Fm. Siricidae Billberg 1820 [horntail]
+
Subfm. †Auliscinae Rasnitsyn 1968
+
G. †Aulisca Rasnitsyn 1968
G. †Megaulisca Rasnitsyn 1968
G. †Megura Rasnitsyn 1968
Subfm. †Gigasiricinae Rasnitsyn 1968
+
G. †Gigasirex Rasnitsyn 1968
G. †Liasirex Rasnitsyn 1968
G. †Pararchexyela Rasnitsyn 1968
Invalid names: Pararchexyelinae Rasnitsyn 1968 [synonym]
Subfm. Siricinae Billberg 1820
+
G. †Eourocerus Archibald and Rasnitsyn 2022
G. †Eoxeris Maa 1949
G. †Urocerites Heer 1867
G. Xeris Costa 1894
G. Xoanon Semenov 1921
G. †Ypresiosirex Archibald and Rasnitsyn 2015
Invalid names: Orussomorpha Rasnitsyn 1980 [invalid subgroup]
Superfm. Xyeloidea Newman 1834 [sawfly]
+
Fm. †Syspastoxyelidae Engel and Huang 2016
+
G. †Striaexyela Zheng 2019
+
†Striaexyela longicornis Zheng 2019
†Striaexyela simpla Wang et al. 2021
Fm. Xyelidae Newman 1834 [sawfly]
+
Subfm. †Archexyelinae Rasnitsyn 1964
+
G. †Asioxyela Rasnitsyn 1964
+
†Asioxyela grandipennis Rasnitsyn 1964
†Asioxyela parvula Rasnitsyn 1964
†Asioxyela paurura Rasnitsyn 1969
†Asioxyela smilodon Rasnitsyn 1964
G. †Ferganoxyela Rasnitsyn 1969
+
†Ferganoxyela destructa Rasnitsyn 1969
†Ferganoxyela sogdiana Rasnitsyn 1969
G. †Leioxyela Rasnitsyn 1969
+
†Leioxyela antiqua Rasnitsyn 1964
†Leioxyela grandis Rasnitsyn 1969
†Leioxyela lata Rasnitsyn 1964
†Leioxyela mitis Rasnitsyn 1969
†Leioxyela mollis Rasnitsyn 1969
G. †Madygenius Rasnitsyn 1969
+
†Madygenius extraradius Rasnitsyn 1969
†Madygenius primitivus Rasnitsyn 1969
G. †Oryctoxyela Rasnitsyn 1969
+
†Oryctoxyela anomala Rasnitsyn 1969
†Oryctoxyela triassica Rasnitsyn 1969
G. †Triassoxyela Rasnitsyn 1964
+
†Triassoxyela foveolata Rasnitsyn 1964
†Triassoxyela kirgizica Rasnitsyn 1964
†Triassoxyela orycta Rasnitsyn 1969
†Triassoxyela sharovi Denisova et al. 2023
G. †Xaxexis Pagliano and Scaramozzino 1990
+
†Xaxexis longhuaensis Hong 1984
Invalid names: Euryxyela Hong 1984 [replaced]
G. †Xiphoxyela Rasnitsyn 1969
+
†Xiphoxyela procrusta Rasnitsyn 1969
†Xiphoxyela striata Rasnitsyn 1964
G. †Xyelinus Rasnitsyn 1964
+
†Xyelinus angustiradius Rasnitsyn 1964
†Xyelinus majus Rasnitsyn 1969
†Xyelinus scherbachov Denisova et al. 2023
Subfm. †Daohugoinae Rasnitsyn and Zhang 2004
+
G. †Daohugoa Rasnitsyn and Zhang 2004
+
†Daohugoa bella Wang et al. 2019
†Daohugoa longa Wang et al. 2019
†Daohugoa tobiasi Rasnitsyn and Zhang 2004
Subfm. Macroxyelinae Ashmead 1898
+
Tr. †Angaridyelini Rasnitsyn 1966
+
G. †Angaridyela Rasnitsyn 1966
G. †Baissoxyela Rasnitsyn 1969
G. †Ceratoxyela Zhang and Zhang 2000
G. †Evacuxyela Dai et al. 2022
G. †Leptoxyela Dai et al. 2022
G. †Lethoxyela Zhang and Zhang 2000
G. †Liaoxyela Zhang and Zhang 2000
G. †Nigrimonticola Rasnitsyn 1966
G. †Ophthalmoxyela Rasnitsyn 1966
Tr. †Ceroxyelini Rasnitsyn 1966
+
G. †Ceroxyela Rasnitsyn 1966
G. †Isoxyela Zhang and Zhang 2000
G. †Sinoxyela Zhang and Zhang 2000
Tr. †Gigantoxyelini Rasnitsyn 1969
+
G. †Abrotoxyela Gao et al. 2009
G. †Chaetoxyela Rasnitsyn 1966
G. †Gigantoxyela Rasnitsyn 1966
G. †Heteroxyela Zhang and Zhang 2000
G. †Magnaxyela Zheng et al. 2020
G. †Shartexyela Rasnitsyn 2008
Tr. Macroxyelini Ashmead 1898
+
G. †Anthoxyela Rasnitsyn 1977
G. †Brachyoxyela Gao et al. 2011
G. †Chionoxyela Rasnitsyn 1993
G. Megaxyela Ashmead 1898
G. †Paleoxyela Jouault et al. 2021
Tr. Xyeleciini Benson 1945
+
G. †Microxyelecia Rasnitsyn 1969
G. †Proxyelia Jouault et al. 2021
G. †Scleroxyela Zheng et al. 2021
G. †Uroxyela Rasnitsyn 1966
G. Xyelecia Ross 1932
G. †Xyelites Rasnitsyn 1966
Subfm. †Madygellinae Rasnitsyn 1969
+
G. †Madygella Rasnitsyn 1969
+
†Madygella analoga Rasnitsyn 1969
†Madygella aristovi Kopylov 2014
†Madygella bashkuevi Kopylov 2014
†Madygella humioi Oyama and Maeda 2020
†Madygella kurochkini Kopylov 2014
†Madygella levivenosa Kopylov 2014
G. †Samarkandykia Kopylov 2014
+
†Samarkandykia ryzhkovae Kopylov 2014
†Samarkandykia shmakovi Kopylov 2014
Subfm. Xyelinae Newman 1834
+
Tr. †Liadoxyelini Rasnitsyn 1966
+
G. †Anomoxyela Rasnitsyn 1966
G. †Cathayxyela Wang et al. 2014
G. †Kirghizoxyela Rasnitsyn 1966
G. †Liadoxyela Martynov 1937
G. †Lydoxyela Rasnitsyn 1966
G. †Novoxyela Zheng et al. 2023
G. †Orthoxyela Rasnitsyn 1983
G. †Platyxyela Wang et al. 2012
G. †Potrerilloxyela Lara et al. 2014
G. Pleuroneura Konow 1897
Tr. Xyelini Newman 1834
+
G. †Enneoxyela Rasnitsyn 1966
G. †Eoxyela Rasnitsyn 1965
G. †Spathoxyela Rasnitsyn 1969
G. †Xyelisca Rasnitsyn 1969
G. †Yanoxyela Ren 1995
Invalid names: Aculeata Scopoli 1763 [synonym], Cleptes steenstrupi Beck 1836 [nomen nudum], Piezata Fabricius 1798 [synonym], Pseudisobrachium oligocenicum Theobald 1937 [nomen dubium], Shandongodes necrodes Zhang 1985 [nomen dubium], Siricomorpha Rasnitsyn 1980 [invalid subgroup], Symphyta Gerstaecker 1867 [invalid subgroup]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
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Source: c = class, p = phylum | |||||
References: Kiessling 2004, Bush and Bambach 2015 |
Age range
Maximum range based only on fossils: base of the Volkhov to the top of the Holocene or 471.10000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 469.2 Ma
Minimum age of oldest fossil (stem group age): 469.2 Ma
Collections (1030 total)
Oldest occurrences | |||
Time interval | Ma | Country or state | Original ID and collection number |
---|---|---|---|
Volkhov | Russian Federation (Leningrag oblast) | Aulocopsis simplex (235818) | |
Ladinian | Kyrgyzstan (Osh) | Asioxyela parvula, Triassoxyela grandipennis, Asioxyela smilodon, Xyelinus angustiradius, Triassoxyela kirgizica, Triassoxyela orycta, Triassoxyela striata, Euryxyela euryptera, Euryxyela lata, Asioxyela antiqua (106090) Asioxyela paurura, Xyelinus majus, Xiphoxyela procrusta, Lithoxyela fenestralis, Leioxyela grandis, Leioxyela mollis, Leioxyela mitis, Madygenius extraradius, Dinoxyela armata, Oryctoxyela triassica, Madygella analoga (110763) Madygella kurochkini, Chubakka madygensis (163987) Triassoxyela foveolata (113468) Triassoxyela orycta, Ferganoxyela destructa, Ferganoxyela sogdiana, Madygenius primitivus, Oryctoxyela anomala (106088) Xyelidae indet., Xyelinus scherbachov, Microxyela minuta, Triassoxyela sharovi, Ferganoxyela sp., Ferganoxyela destructa, Samarkandykia ryzhkovae, Samarkandykia shmakovi, Madygella bashkuevi, Madygella aristovi, Madygella levivenosa (163986) | |
Carnian | South Africa (Eastern Cape) | Moltenia rieki (23142) | |
Carnian | Japan (Yamaguchi) | Xyeloidea indet., Madygella humioi (106482) | |
Carnian | Argentina | Potrerilloxyela menendezi (151962) | |
Norian | Australia (Queensland) | Archexyela crosbyi (106484) Archexyela ipswichensis (156521) | |
Sevatian | Germany | Sepulenia sp. (123417) | |
Hettangian - Sinemurian | Kyrgyzstan (Tonskiy) | Kirghizoxyela mirabilis (105087) Sogutia liassica (105084) | |
Pliensbachian | Kyrgyzstan | Liadoxyela praecox (114701) | |
Early/Lower Toarcian | Germany (Mecklenburg-Vorpommern) | Grimmaratavites mirabilis (114993) Xyelula benderi (154298) | |
Early/Lower Toarcian | Germany | Ephialtitidae indet., Xyelula benderi (114991) | |
Early/Lower Toarcian | Germany (Bavaria) | Xyelula benderi (117794) | |
Early/Lower Toarcian | Germany (Niedersachsen) | Liadobracona raduhna (139222) Sippelipterus liasinus (114995) Thilopterus lampei (154300) Xyelula benderi (154299) | |
Falciferum | Luxembourg | Symphyta indet., Pseudoxyelocerus bascharagensis (105427) | |
Toarcian | Russian Federation (Irkutsk) | Liadoxyela iensis (104995) | |
Toarcian | Germany | Liadobracona raduhna, Brigittepterus brauckmanni, Xyelula benderi (104056) | |
Toarcian | Kyrgyzstan (Batken) | Liasirex sogdianus, Shurabisca liassica, Sepulenia syricta (105124) Proapocritus praecursor, Onokhoius sogdianus, Ferganolyda cubitalis, Ferganolyda radialis, Ferganolyda sogdiana, Sagulyda ferganica, Sagulyda magna (105003) Sagulyda arcuata (110817) | |
Toarcian | India (Andhra Pradesh) | Xyelula alexandri (79229) | |
Toarcian | India (Adilabad) | Kotaphialtites frankmortoni (117538) | |
Aalenian | China (Shaanxi) | Yananphilius peizhuangensis (222545) |