Basic info Taxonomic history Classification Included Taxa
Morphology Ecology and taphonomy External Literature Search Age range and collections

Adephaga

Insecta - Coleoptera

Synonyms
Synonymy list
YearName and author
1806Adephaga Clairville p. 2
1815Dytiscoidea Leach p. 84
1981Adephaga Crowson p. 694
1992Adephaga Carpenter
1995Adephaga Lawrence and Newton, Jr.
2002Tritarsusidae Hong p. 102
2002Tritarsus Hong p. 104
2002Tritarsus latus Hong p. 105 figs. 2.6.1-10
2007Dytiscoidea Ghosh et al. p. 1336
2011Tritarsidae Bouchard et al.
2011Adephaga Bouchard et al. p. 99
2022Adephaga Boudinot et al.
2022Adephaga Cai et al.
2022Tritarsusidae Cai et al.

Is something missing? Join the Paleobiology Database and enter the data

RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—EcdysozoaAguinaldo et al. 1997
—Panarthropoda
phylumArthropodaLatreille 1829
—Deuteropoda
—Mandibulata
—PancrustaceaZrzavý and Štys 1997
—AltocrustaceaRegier et al. 2010
superclassAllotriocarida
—Athalassocarida
—LabiocaridaSchwentner et al. 2017
superclassHexapodaLatreille 1825
—Cercofilata
RankNameAuthor
classInsecta
—Dicondylia
—Paranotalia
subclassPterygota()
—NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
—Holometabola
—Aparaglossata
—Neuropteriformia
—Coleopterida
orderColeoptera
—Mesocoleoptera
—Metacoleoptera
suborderAdephaga
suborderAdephaga

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Subor. Adephaga Clairville 1806
show all | hide all
G. †Amarodes Handlirsch 1906
+
†Amarodes pseudozabrus Deichmüller 1886
G. †Amphoxyne Bode 1953
+
†Amphoxyne darbiensis Ponomarenko 1986
†Amphoxyne lineata Bode 1953
+
Invalid names: Cryptauchenia simplex Bode 1953 [synonym]
†Amphoxyne minuta Bode 1953
+
Invalid names: Coilotrachelus lineatus Bode 1953 [synonym], Cricotrachelites rotundatus Bode 1953 [synonym], Loxoncus procerus Bode 1953 [synonym]
Invalid names: Coilotrachelus Bode 1953 [synonym], Cricotrachelites Bode 1953 [synonym], Loxostelidotus Bode 1953 [synonym]
Fm. Carabidae Latreille 1802 [ground beetle]
+
G. †Antemnacrassa Gomez Pallerola 1979
+
†Antemnacrassa albomaculata Nikolajev 2004
†Antemnacrassa albosulcata Nikolajev 2004
†Antemnacrassa geminata Nikolajev 2004
†Antemnacrassa incerta Ponomarenko 1990
†Antemnacrassa kazakhstanica Nikolajev 2007
†Antemnacrassa latitibialis Nikolajev 1992
†Antemnacrassa maculata Nikolajev 1992
†Antemnacrassa magna Nikolajev 2004
†Antemnacrassa monreali Gomez Pallerola 1979
†Antemnacrassa nebulosa Nikolajev 2004
†Antemnacrassa nigrimontana Nikolajev 1992
†Antemnacrassa picturata Nikritin 1977
†Antemnacrassa punctata Ponomarenko 1986
†Antemnacrassa sulcata Nikritin 1977
†Antemnacrassa vitimensis Nikritin 1977
G. Apotomopterus Hope 1838
G. †Ataktosites Haupt 1956
+
†Ataktosites palustris Haupt 1956
G. Bembidium Latreille 1802
+
Subg. Bembidium (Campa) Motschulsky 1844
+
Subg. Bembidium (Testidium) Motschulsky 1864
†Bembidium absolutum Giebel 1856
†Bembidium cyaneomicans Piton 1940
†Bembidium exoletum Scudder 1876
†Bembidium expletum Scudder 1900
†Bembidium florissantensis Wickham 1913
†Bembidium fragmentum Scudder 1890
†Bembidium glaciatum Scudder 1890
†Bembidium haywardi Scudder 1900
†Bembidium infernum Heer 1856
†Bembidium laevigatum Förster 1891
†Bembidium obductum Scudder 1900
†Bembidium praeteritum Scudder 1900
†Bembidium saportanum Oustalet 1874
†Bembidium tumulorum Scudder 1900
†Bembidium vanum Scudder 1900
†Bembidium vestigium Scudder 1900
Subfm. Brachininae Bonelli 1810
+
Tr. Brachinini Bonelli 1810
+
Subtr. Brachinina Bonelli 1810
+
G. Brachinus Weber 1801
G. †Brachinites Fritsch 1882
+
†Brachinites truncatus Fritsch 1882
Subfm. Broscinae Hope 1838
+
Tr. Broscini Hope 1838
+
Subtr. Broscina Hope 1838
+
G. Miscodera Eschscholtz 1830
Subtr. Nothobroscina Roig-Juñent 2000
+
G. Mecodema Blanchard 1843
G. †Carabidium Westwood 1854
+
†Carabidium dejeanianum Westwood 1854
Subfm. Carabinae Latreille 1802
+
G. †Atrirabus Hong and Wang 1990
+
†Atrirabus shandongensis Hong and Wang 1990
†Atrirabus tuanwangensis Hong and Wang 1990
Tr. Carabini Latreille 1802
+
G. Calosoma Weber 1801
+
Subg. Calosoma (Campalita) Motschulsky 1865
†Calosoma agassizi Barthelemy-Lapommeraye 1846
†Calosoma brunneum Hong 1985
†Calosoma calvini Wickham 1909
†Calosoma caraboides Heer 1860
†Calosoma catenulatum Heer 1860
†Calosoma ceresti Nel 1987
†Calosoma cockerelli Wickham 1910
†Calosoma emmonsi Scudder 1900
†Calosoma escheri Heer 1860
†Calosoma fernquisti Cockerell 1925
†Calosoma grasui Paucă and Ciobanu 1978
†Calosoma heeri Scudder 1895
Calosoma inquisitor Linnaeus 1758
†Calosoma jaccardi Heer 1860
†Calosoma kimi Lee and Nam 2021
Calosoma maderae Fabricius 1775
†Calosoma nauckianum Heer 1860
Calosoma semilaeve LeConte 1852
Calosoma sycophanta Linnaeus 1758 [forest caterpillar hunter]
G. Carabus Linnaeus 1758
+
Subg. Carabus (Apotomopterus) Hope 1838
Subg. Carabus (Archicarabus) Seidlitz 1887
Subg. Carabus (Aulonocarabus) Reitter 1896
Subg. Carabus (Damaster) Kollar 1836
Subg. Carabus (Hemicarabus) Gehin 1876
Subg. Carabus (Homoeocarabus) Reitter 1896
Subg. Carabus (Megadontus) Solier 1848
Subg. Carabus (Melancarabus) Thomson 1875
Subg. Carabus (Ohomopterus) Reitter 1896
Subg. Carabus (Tachypus) Weber 1801
Subg. Carabus (Tanaocarabus) Reitter 1896
†Carabus agassizi Barthelemy-Lapommeraye 1846
Carabus clathratus Linnaeus 1761
Carabus clatratus Linnaeus 1761
†Carabus foveolatus Piton 1935
Carabus granulatus Linnaeus 1758
Carabus hortensis Linnaeus 1758
†Carabus jeffersoni Scudder 1900
Carabus kollari Palliardi 1825
Carabus maderae Fabricius 1775
†Carabus maeandroides Lomnicki 1894
†Carabus mecothoracus Zhang 1989
†Carabus neli Deuve 1998
†Carabus novalensis Omboni 1886
†Carabus ovalus Hong and Wang 1986
†Carabus praearvensis Lomnicki 1894
Invalid names: Calosomini Bonelli 1810 [synonym]
Tr. Cychrini Perty 1830
+
G. Cychrus Fabricius 1794
+
Cychrus caraboides Linnaeus 1758
G. Scaphinotus Dejean 1826
+
Subg. Scaphinotus (Brennus) Motschulsky 1866
Subg. Scaphinotus (Nomaretus) Leconte 1853
Scaphinotus interruptus Menetries 1843
†Scaphinotus minor Horn 1876
†Scaphinotus wheatleyi Horn 1876
G. Sphaeroderus Dejean 1831 [small snail eating beetle]
+
Sphaeroderus nitidicollis Guerin-Meneville 1829
Tr. Pamborini Hope 1838
+
G. Maoripamborus Brookes 1944
+
G. †Carabites Ponomarenko 1977
+
†Carabites caraboides Ponomarenko 1989
†Carabites nigriventris Ponomarenko 1986
G. †Carabites Heer 1852
+
†Carabites anthracinus Heer 1852
†Carabites arapahoensis Cockerell 1918
†Carabites capellinii Cecconi 1892
†Carabites creta Ponomarenko 1977
†Carabites cuneatus Cockerell 1920
†Carabites dubius Geinitz 1894
†Carabites ellipticus Haupt 1956
†Carabites eocenicus Cockerell 1921
†Carabites exanimus Scudder 1892
†Carabites exilis Heer 1862
†Carabites gardneri Cockerell 1920
†Carabites geinitzi Handlirsch 1906
†Carabites harringtoni Cockerell 1936
†Carabites islandicus Heer 1868
†Carabites kincaidi Cockerell 1908
†Carabites laesicollis Heer 1862
†Carabites parallelus Cockerell 1920
†Carabites peracutus Cockerell 1920
†Carabites rugosus Meunier 1898
†Carabites russelli Cockerell 1928
†Carabites schueli Cockerell 1926
†Carabites scoticus Cockerell 1921
†Carabites vitimensis Ponomarenko 1977
G. †Carabopteron Martynov 1926
+
†Carabopteron rugosum Ponomarenko 1985
G. †Cavicarabus Hong 1985
+
†Cavicarabus lucensus Hong 1985
Unr. Celeripedes Latreille 1802
+
Invalid names: Barbati Latreille 1802 [empty], Crepitantes Latreille 1802 [empty], Longipalpati Latreille 1802 [empty], Melanchlaeni Latreille 1802 [empty], Metallici Latreille 1802 [empty]
G. †Cicindelites Meunier 1898
+
†Cicindelites armissanti Meunier 1898
Subfm. †Conjunctiinae Ponomarenko 1977
+
G. †Conjunctia Ponomarenko 1977
+
†Conjunctia longa Zhang 1997
†Conjunctia prodroma Ponomarenko 1977
G. †Mesorabus Ponomarenko 1977
+
†Mesorabus elongatus Ponomarenko 1977
Subfm. Elaphrinae Latreille 1802
+
G. Blethisa Bonelli 1810
+
Blethisa catenaria Brown 1944
Blethisa inexpectata Goulet and Smetana 1983
Blethisa multipunctata Linnaeus 1758
†Blethisa pleistocenica Lomnicki 1894
Invalid names: Helobium Leach 1815 [objective synonym]
G. Diacheila Motschulsky 1844
+
Diacheila arctica Gyllenhal 1810
†Diacheila matthewsi Böcher 1995
Diacheila polita Faldermann 1835
G. Elaphrus Fabricius 1775
+
Elaphrus americanus Dejean 1831
Elaphrus angusticollis Sahlberg 1844
Elaphrus californicus Mannerheim 1843
Elaphrus cupreus Duftschmid 1812
Elaphrus finitimus Casey 1920
Elaphrus lapponicus Gyllenhal 1810
Elaphrus lecontei Crotch 1876
†Elaphrus pliocenicus Piton 1939
Elaphrus purpurans Hausen 1891
Elaphrus riparius Linnaeus 1758
Elaphrus sibiricus Motschulsky 1846
Invalid names: Elaphrus irregularis Scudder 1890 [nomen dubium], Elaphrus ruscarius foveatus Pierce 1948 [synonym]
G. †Elaphrotites Haupt 1956
+
†Elaphrotites densus Haupt 1956
G. †Fangshania Hong 1981
+
†Fangshania punctata Hong 1981
G. †Feronites Frič 1884
+
†Feronites velenovskyi Frič 1884
G. †Fonsecacarabus Martins-Neto and Mendes 2002
+
†Fonsecacarabus placidus Martins-Neto and Mendes 2002
Subfm. Gehringiinae Darlington 1933
+
Tr. Gehringiini Darlington 1933
+
Subtr. Gehringiina Darlington 1933
+
G. Gehringia Darlington 1933
G. †Glenopterus Heer 1847
+
G. †Hadrocephalus Handlirsch 1906
+
†Hadrocephalus anglicus Handlirsch 1906
†Hadrocephalus liasinus Handlirsch 1906
†Hadrocephalus minor Handlirsch 1906
Subfm. Harpalinae Bonelli 1810
+
Tr. Abacetini Chaudoir 1873
+
G. Loxandrus Leconte 1852
G. †Aethocarabus Ren 1995
+
Tr. Chlaeniini Brullé 1834
+
Subtr. Chlaeniina Brullé 1834
+
G. Chlaenius Bonelli 1810
+
Invalid names: Shanwangicarabus Hong 1984 [synonym]
Tr. Galeritini Kirby 1825
+
Subtr. Galeritina Kirby 1825
+
G. Galerita Fabricius 1801
Tr. Harpalini Bonelli 1810
+
Subtr. Anisodactylina Lacordaire 1854
+
G. Anisodactylus Dejean 1829
Invalid names: Eurytrichi Le Conte 1847 [synonym]
Subtr. Harpalina Bonelli 1810
+
G. Cratacanthus Dejean 1829
G. Euryderus Le Conte 1846
+
Invalid names: Nothopus Leconte 1852 [objective synonym]
G. Harpalus Latreille 1802
G. Ophonus Dejean 1821
G. Selenophorus Dejean 1829
Invalid names: Dapti Le Conte 1847 [synonym]
Subtr. Stenolophina Kirby 1837
+
G. Bradycellus Erichson 1837
G. Dicheirotrichus Jacquelin du Val 1857
G. Stenolophus Dejean 1821
Tr. Helluonini Hope 1838
+
Subtr. Omphrina Jedlicka 1941
+
G. Creagris Nietner 1857
G. Helluomorpha De Laporte 1834
Tr. Lebiini Bonelli 1810
+
G. †Agatoides Motschulsky 1856
+
†Agatoides carinulatus Motschulsky 1856
Subtr. Calleidina Chaudoir 1873
+
G. Plochionus Latreille and Dejean 1821
Subtr. Cymindidina Laporte 1834
+
G. Cymindis Latreille 1806
G. †Cymindoides Motschulsky 1856
+
†Cymindoides sculptipennis Motschulsky 1856
Subtr. Dromiusina Bonelli 1810
+
G. †Antephilorhizus Kirichenko-Babko et al. 2021
G. Dromius Bonelli 1810
+
Invalid names: Lebina Germar 1813 [synonym]
G. Microlestes Schmidt-Göbel 1846
G. Paradromius Fowler 1887
G. Syntomus Hope 1838
Subtr. Lebiina Bonelli 1810
+
G. Lebia Latreille 1802 [colorful foliage ground beetle]
G. †Miolebidia Zhang 1989
+
†Miolebidia gaura Zhang 1989
Subtr. Pericalina Hope 1838
+
G. Coptodera Dejean 1825
G. †Protoscalidion Schaufuss 1889
+
†Protoscalidion rugiae Schaufuss 1889
Invalid names: Brachinus repressus Scudder 1900 [nomen dubium]
Tr. Licinini Bonelli 1810
+
Subtr. Dicaelina Laporte 1834
+
G. Dicaelus Bonelli 1813
G. Diplocheila Brullé 1834
+
Invalid names: Rembus Latreille 1817 [replaced]
Subtr. Licinina Bonelli 1810
+
G. Badister Clairville 1806
Tr. Odacanthini Laporte 1834
+
Invalid names: Casnoniae LeConte 1861 [synonym]
Tr. Panagaeini Bonelli 1810
+
Subtr. Panagaeina Bonelli 1810
+
G. Panagaeus Latreille 1802
Tr. Pseudomorphini Hope 1838
+
G. †Burmapseudomorphus Beutel et al. 2020
+
†Burmapseudomorphus planus Beutel et al. 2020
Tr. Pterostichini Bonelli 1810
+
G. Cyclotrachelus Chaudoir 1838
+
Subg. Cyclotrachelus (Evarthrus) Leconte 1853
†Cyclotrachelus tenebricus Scudder 1900
Subtr. Pterostichina Bonelli 1810
+
G. Abax Bonelli 1810
G. Myas Dejean 1828
G. Platysma Bonelli 1810
G. Plocamostethus Britton 1940
G. Poecilus Bonelli 1810
+
Invalid names: Feronia Latreille 1816 [synonym]
G. Pterostichus Bonelli 1810
G. Stomis Clairville 1806
Invalid names: Feroniens Dejean 1825 [synonym]
G. †Sinis Heer 1862
+
†Sinis brevicollis Heer 1862
Tr. Sphodrini Laporte 1834
+
Subtr. Calathina Laporte 1834
+
G. Calathus Bonelli 1810
G. †Quasicalathus Schmidt and Will 2022
Subtr. Sphodrina Laporte 1834
+
G. Pristonychus Dejean 1828
G. Sphodrus Clairville 1806
Subtr. Synuchina Lindroth 1956
+
G. Synuchus Gyllenhal 1810
Tr. Zabrini Bonelli 1810
+
Subtr. Amarina Zimmerman 1832
+
G. Amara Bonelli 1810
Subtr. Zabrina Bonelli 1810
+
G. Zabrus Clairville 1806
Tr. Zuphiini Bonelli 1810
+
Subtr. Zuphiina Bonelli 1810
+
G. Polistichus Bonelli 1810
Invalid names: Dryptini Bonelli 1810 [empty], Licininae Bonelli 1810 [synonym]
G. †Harpalomimes Handlirsch 1906
+
†Harpalomimes burmeisteri Giebel 1856
G. †Hebeicarabus Hong 1984
+
G. †Lithocantharis Lin 1997
+
Subfm. Loricerinae Bonelli 1810
+
Tr. Loricerini Bonelli 1810
+
G. †Cretoloricera Liu et al. 2023
+
†Cretoloricera electra Liu et al. 2023
G. Loricera Latreille 1802
+
Loricera decempunctata Eschscholtz 1833
†Loricera electrica Klausnitzer 2003
†Loricera groehni Cai et al. 2017
Loricera pilicornis Fabricius 1775
Invalid names: Loriceritae Bonelli 1810 [empty]
G. †Meileyingia Hong 1987
+
†Meileyingia spinosa Hong 1987
G. †Metagrilium Handlirsch 1906
+
†Metagrilium westwoodi Handlirsch 1906
Subfm. Migadopinae Chaudoir 1861
+
G. †Cretomigadops Liu et al. 2022
+
†Cretomigadops bidentatus Liu et al. 2022
Tr. Migadopini Chaudoir 1861
+
Subtr. Migadopina Chaudoir 1861
+
G. Migadops Waterhouse 1842
Subfm. Nebriinae Laporte 1834
+
G. †Ledouxnebria Deuve 1998
+
†Ledouxnebria brisaci Deuve 1998
Tr. Nebriini Laporte 1834
+
G. †Archaeonebria Kavanaugh and Schmidt 2019
+
†Archaeonebria inexspectata Schmidt and Kavanaugh 2019
G. Leistus Frölich 1799
G. Nebria Latreille 1802
+
Nebria brevicollis Fabricius 1792
†Nebria fossilis Piton 1935
Nebria frigida Sahlberg 1844
Nebria metallica Fischer von Waldheim 1822
Nebria nivalis Paykull 1790
†Nebria pluto Heer 1862
Nebria rufescens Strøm 1768
+
Invalid names: Nebria gyllenhali Schönherr 1806 [synonym]
Nebria sahlbergi Fischer von Waldheim 1828
†Nebria tisiphone Oustalet 1874
Tr. Notiophilini Motschulsky 1850
+
G. Notiophilus Duméril 1805
+
Notiophilus aeneus Herbst 1806
Notiophilus aquaticus Linnaeus 1758
Notiophilus biguttatus Fabricius 1779
Notiophilus palustris Duftschmid 1812
†Notiophilus vancouveri Pierce 1950
Tr. Opisthiini Dupuis 1912
+
G. Opisthius Kirby 1837
+
Tr. Pelophilini Kavanaugh 1996
+
G. Pelophila Dejean 1821
+
Pelophila borealis Paykull 1790
G. †Nebrorabus Ponomarenko 1989
+
†Nebrorabus baculum Ponomarenko 1989
†Nebrorabus capitatus Ponomarenko 1989
†Nebrorabus nebrioides Ponomarenko 1989
G. †Neothanes Scudder 1890
+
†Neothanes europaeus Ponomarenko 2019
†Neothanes testeus Scudder 1878
G. †Notokistus Handlirsch 1906
+
†Notokistus brodiei Handlirsch 1906
Subfm. Omophroninae Bonelli 1810
+
G. †Cretomophron Rosová et al. 2023
+
†Cretomophron mutilum Rosová et al. 2023
Subfm. Paussinae Latreille 1806 [ant nest beetle]
+
G. †Cerapterites Wasmann 1926
+
†Cerapterites primaevus Wasmann 1926
Tr. Paussini Latreille 1806
+
Subtr. †Arthropteritina Luna de Carvalho 1961
+
G. †Arthropterites Wasmann 1926
Subtr. Carabidomemnina Wasmann 1928
+
G. Eohomopterus Wasmann 1919
Subtr. Cerapterina Billberg 1820
+
G. Arthropterus M'Leay 1838
G. †Protocerapterus Wasmann 1926
Subtr. †Eopaussina Luna de Carvalho 1951
+
G. †Eopaussus Wasmann 1926
Subtr. Paussina Latreille 1806
+
G. Homopterus Westwood 1841
G. Paussus Linnaeus 1775
G. Pleuarthropterus Wasmann 1929
G. †Paussoides Motschulsky 1856
+
†Paussoides mengei Motschulsky 1856
G. †Paussopsis Cockerell 1911
+
†Paussopsis nearctica Cockerell 1911
G. †Succinarthropterus Kolbe 1926
+
†Succinarthropterus helmi Schaufuss 1896
†Succinarthropterus kolbei Wasmann 1926
Invalid names: Arthropterillus Wasmann 1926 [synonym]
Invalid names: Ozaenidae Hope 1838 [synonym], Ozaenini [empty]
Subfm. Platyninae Bonelli 1810
+
Tr. Platynini Bonelli 1810
+
G. Agonum Bonelli 1810
+
Subg. Agonum (Europhilus) Chaudoir 1859
Subg. Agonum (Olisares) Motschulsky 1865
Subg. Agonum (Platynomicrus) Casey 1920
†Agonum abstractum Scudder 1900
†Agonum antiquum Piton 1940
Agonum micans Nicolai 1822
Agonum muelleri Herbst 1784
Agonum quinquepuncatum Motschulsky 1844
Agonum sahlbergi Chaudoir 1850
Agonum sexpunctatum Linnaeus 1758
Agonum versutum Sturm 1824
G. Anchomenus Bonelli 1810
+
†Anchomenus bipunctatus Förster 1891
†Anchomenus orphanus Heer 1847
G. Blackburnia Sharp 1878
+
Subg. Blackburnia (Blackburnia) Sharp 1878
Subg. Blackburnia (Colpocaccus) Sharp 1903
Subg. Blackburnia (Metromenus) Sharp 1884
Invalid names: Disenochus Blackburn 1878 [synonym]
G. Oxypselaphus Chaudoir 1843
+
G. Paranchus Lindroth 1974
+
Paranchus albipes Fabricius 1794
Subtr. Platynina Bonelli 1810
+
G. Limodromus Motschulsky 1850
G. Platynus Bonelli 1810
+
Subg. Platynus (Batenus) Motschulsky 1865
†Platynus caesus Scudder 1890
†Platynus calvini Wickham 1917
†Platynus casus Scudder 1890
†Platynus desuetus Scudder 1890
†Platynus dilapidatus Scudder 1895
†Platynus dissipatus Scudder 1890
†Platynus exterminatus Scudder 1900
†Platynus florissantensis Wickham 1913
Platynus funebris LeConte 1854
†Platynus halli Scudder 1890
†Platynus harttii Scudder 1890
†Platynus hindei Scudder 1890
†Platynus insculptipennis Wickham 1917
†Platynus interglacialis Scudder 1900
†Platynus interitus Scudder 1900
†Platynus longaevus Scudder 1900
†Platynus pleistocenicus Wickham 1917
†Platynus senex Scudder 1878
†Platynus subgelidus Wickham 1917
†Platynus tartareus Scudder 1900
G. †Praeanchodemus Schmidt et al. 2017
+
†Praeanchodemus punctaticeps Schmidt et al. 2017
G. †Procarabites Handlirsch 1906
Subfm. Protopaussinae Gestro 1892
+
G. Protopaussus Gestro 1892
+
†Protopaussus pristinus Nagel 1997
Subfm. †Protorabinae Ponomarenko 1977
+
G. †Cordorabus Ponomarenko 1977
+
†Cordorabus antennatus Ponomarenko 1977
†Cordorabus minimus Ponomarenko 1977
†Cordorabus notatus Ponomarenko 1977
†Cordorabus striatus Ponomarenko 1986
†Cordorabus vittatus Ponomarenko 1980
†Cordorabus wuqiensis Hong 1995
G. †Cretorabus Ponomarenko 1977
+
†Cretarabus medius Hong and Wang 1990
†Cretorabus capitatus Ponomarenko 1977
†Cretorabus latus Ponomarenko 1977
†Cretorabus orientalis Ponomarenko 1989
†Cretorabus ovalis Ponomarenko 1989
†Cretorabus rasnitsyni Wang and Zhang 2011
†Cretorabus sulcatus Ponomarenko et al. 2005
G. †Lirabus Hong 1992
+
†Lirabus granulatus Hong 1992
G. †Lithorabus Ponomarenko 1977
+
†Lithorabus incertus Ponomarenko 1977
G. †Magnirabus Hong and Wang 1990
+
†Magnirabus furvus Hong and Wang 1990
G. †Ovrabites Ponomarenko 1977
+
†Ovrabites incertus Ponomarenko 1993
†Ovrabites jurassicus Ponomarenko 1977
†Ovrabites ovalis Ponomarenko 1977
G. †Penecupes Ren 1995
+
†Penecupes rapax Ren 1995
G. †Protorabus Ponomarenko 1977
+
†Protorabus crassus Ponomarenko 1989
†Protorabus elongatus Ponomarenko 1986
†Protorabus kobdoensis Ponomarenko 1986
†Protorabus magnus Ponomarenko 1977
†Protorabus minisculus Zhang 1997
†Protorabus nigrimonticola Ponomarenko 1977
†Protorabus planus Ponomarenko 1977
†Protorabus tsagensis Ponomarenko 1989
G. †Rhopalochlaenius Zhang et al. 1994
+
†Rhopalochlaenius phalanthus Zhang et al. 1994
Subfm. Scaritinae Bonelli 1810
+
Tr. Clivinini Rafinesque 1815
+
G. Camptodontus Dejean 1826
Subtr. Clivinina Rafinesque 1815
+
G. Clivina Latreille 1802
Tr. Dyschiriini Kolbe 1880
+
G. Dyschiriodes Jeannel 1941
+
Dyschiriodes aeneus Dejean 1825
Dyschiriodes luticola Chaudoir 1850
Dyschiriodes nigricornis Motschulsky 1844
+
Invalid names: Dyschirius septentrionum Munster 1923 [synonym]
G. Dyschirius Bonelli 1810
+
Dyschirius montanus Le Conte 1879
+
Invalid names: Dyschirius arenosus Stephens 1827 [synonym]
Invalid names: Helophorus rigescens Scudder 1890 [nomen dubium]
G. †Dyschiriomimus Yablokov-Khnzorian 1960
+
†Dyschiriomimus stackelbergi Yablokov-Khnzorian 1960
Tr. †Palaeoaxinidiini McKay 1991
+
G. †Palaeoaxinidium McKay 1991
+
Tr. Scaritini Bonelli 1810
+
Subtr. Scaritina Bonelli 1810
+
G. Scarites Fabricius 1775
Invalid names: Promecognathinae Le Conte 1853 [synonym]
G. †Sinocarabus Hong 1982
+
G. Stereocerus Kirby 1837
+
Stereocerus rubripes Motschulsky 1860
G. †Sunocarabus Hong 1987
+
†Sunocarabus brunneus Hong 1987
Subfm. Trechinae Bonelli 1810
+
Tr. Patrobini Kirby 1837
+
Subtr. Patrobina Kirby 1837
+
G. Patrobus Dejean 1821
G. †Trechinites Heer 1862
+
†Trechinites oblongus Heer 1862
Unr. Trechitae Bonelli 1810
+
G. †Balticeler Schmidt and Maddison 2021
+
†Balticeler kerneggeri Schmidt and Maddison 2021
Tr. Bembidiini Stephens 1827
+
G. Asaphidion Des Gozis 1886
G. Bembidion Latreille 1802
G. Ocys Stephens 1828
G. Sinechostictus Motschulsky 1864
Invalid names: Bembidiina Stephens 1827 [invalid subgroup], Tachyina Motschulsky 1862 [empty]
Tr. Pogonini Laporte 1834
+
G. Pogonus Dejean 1821
Tr. Tachyini Motschulsky 1862
+
Subtr. Tachyina Motschulsky 1862
Tr. Trechini Bonelli 1810
+
G. †Antarctotrechus Ashworth and Erwin 2016
G. Trechiama Jeannel 1927
Subtr. Trechina Bonelli 1810
Invalid names: Bembidion succini Giebel 1856 [nomen dubium], Patrobinae Kirby 1837 [synonym]
G. †Trechoides Motschulsky 1856
+
†Trechoides fasciatus Motschulsky 1856
G. †Tremembecarabus Martins-Neto 1998
+
†Tremembecarabus rotundus Martins-Neto 1998
G. †Wongyekokia Lin 1997
+
G. †Wuchangicarabus Hong 1985
+
†Wuchangicarabus latus Hong 1985
G. †Yunnanocarabus Lin 1977
+
Invalid names: Cratacanthus florissantensis Wickham 1917 [nomen dubium], Fossores Latreille 1802 [empty], Loricera exita Scudder 1900 [nomen dubium], Rhysodinae Laporte 1840 [empty], Siagoninae Bonelli 1813 [empty], Trechus fractus Wickham 1912 [nomen dubium]
Infraor. Carabiformia Lameere 1900
+
Superfm. Caraboidea Latreille 1802
+
G. †Beipiaocarabus Hong 1983
+
G. †Carabilarva Ponomarenko 1985
+
†Carabilarva jurassica Ponomarenko 1985
†Carabilarva robusta Makarov 1995
†Carabilarva triassica Makarov and Prokin 2013
G. †Eurycoleus Hong 1982
+
†Eurycoleus arcuatus Hong 1982
†Eurycoleus foveolatus Hong 1984
†Eurycoleus parvus Hong 1982
G. †Glottocoleus Hong 1982
+
G. †Glyptocoleus Hong 1982
+
G. †Grypocoleus Hong 1982
+
G. †Jiangxicoleus Hong 1988
+
G. †Leptocoleus Hong 1982
+
†Leptocoleus lenis Hong 1982
G. †Mesocoleus Hong 1982
+
G. †Obesofemoria Hong 1982
+
G. †Phyllocoleus Hong 1982
+
G. †Pleurocoleus Hong 1982
+
G. †Procalosoma Meunier 1895
+
†Procalosoma giardi Meunier 1895
†Procalosoma incertum Ponomarenko 2014
†Procalosoma major Handlirsch 1906
†Procalosoma mimor Handlirsch 1906
†Procalosoma minor Handlirsch 1906
G. †Vago Hong 1982
+
†Vago oblonga Hong 1982
G. †Yumenocoleus Hong 1982
+
†Yumenocoleus lineatus Hong 1982
†Yumenocoleus longus Hong 1982
†Yumenocoleus tenuis Hong 1988
Invalid names: Unda chonggezhuangensis Wang 1998 [nomen dubium], Unda pandurata Ren 1995 [nomen dubium]
G. †Chimaerocoleus Ponomarenko 1985
+
†Chimaerocoleus aenigma Ponomarenko 1985
Fm. Cicindelidae Latreille 1802 [tiger beetle]
+
Tr. Cicindelini Latreille 1802
+
Subtr. Cicindelina Latreille 1802
+
G. Cicindela Linnaeus 1758
+
Cicindela hybrida Linnaeus 1758
Cicindela oregona LeConte 1857
Subtr. Iresiina Rivalier 1971
+
G. †Palaeoiresina Wiesner et al. 2017
+
†Palaeoiresina cassolai Wiesner et al. 2017
G. †Cretotetracha Zhao et al. 2019
+
†Cretotetracha grandis Zhao et al. 2019
G. †Goriresina Matalin et al. 2021
+
†Goriresina fungifora Matalin et al. 2021
G. †Hujia Song et al. 2022
Tr. Manticorini Laporte 1834
+
G. Omus Eschscholtz 1829
+
Omus audouini Reiche 1838
Invalid names: Amblycheilini Csiki 1903 [synonym]
Tr. Megacephalini Laporte 1834
+
G. Tetracha Hope 1838
+
Tetracha carolina Linnaeus 1763
Invalid names: Megacephalina Laporte 1834 [empty]
G. †Opsis Handlirsch 1906
+
†Opsis bavarica Handlirsch 1906
G. †Oxycheilopsis Cassola and Werner 2004
+
†Oxycheilopsis cretacicus Cassola and Werner 2004
G. †Palaeopronyssiformia Wiesner et al. 2017
+
†Palaeopronyssiformia groehni Wiesner et al. 2017
Invalid names: Oxycheilini Thomson 1857 [empty]
Fm. †Colymbotethidae Ponomarenko 1993
+
G. †Colymbotethis Ponomarenko 1993
+
†Colymbotethis antecessor Ponomarenko 1993
Fm. †Coptoclavidae Ponomarenko 1961
+
G. †Agrascapha Lin 1992
+
†Agrascapha curta Lin 1992
G. †Amblycephalonius Bode 1953
+
+
Invalid names: Barycephalus nudatus Bode 1953 [synonym], Tricyrtus tenuistriatus Bode 1953 [synonym]
Invalid names: Barocephalus Carpenter 1986 [synonym], Barycephalus Bode 1953 [synonym], Tricyrtus Bode 1953 [synonym]
Subfm. †Charonoscaphinae Ponomarenko 1977
+
G. †Charonoscapha Ponomarenko 1977
+
†Charonoscapha grossa Ponomarenko 1977
†Charonoscapha ovata Ponomarenko 1977
G. †Charonoscaphidia Ponomarenko 1977
+
†Charonoscaphidia elongata Ponomarenko 1977
G. †Coptoclavia Ping 1928
+
†Coptoclavia africana Teixeira 1975
Subfm. †Coptoclavinae Ponomarenko 1961
+
G. †Bolbonectes Ponomarenko 1987
+
†Bolbonectes intermedius Ponomarenko 1987
†Bolbonectes occidentalis Ponomarenko 1993
†Bolbonectus lithographicus Ponomarenko and Martínez-Delclòs 2000
G. †Coptoclava Ping 1928
+
†Coptoclava longipoda Ping 1928
G. †Hoyaclava Soriano et al. 2007
+
†Hoyaclava buscalionae Soriano et al. 2007
G. †Megacoptoclava Ponomarenko and Martínez-Delclòs 2000
+
†Megacoptoclava longiurogomphia Ponomarenko and Martínez-Delclòs 2000
Subfm. †Coptoclaviscinae Soriano et al. 2007
+
G. †Coptoclavella Ponomarenko 1980
+
†Coptoclavella elegans Ponomarenko 1980
†Coptoclavella inexpecta Soriano et al. 2007
†Coptoclavella jurassica Ponomarenko 2014
†Coptoclavella minor Ponomarenko 1980
†Coptoclavella purbeckensis Ponomarenko et al. 2005
†Coptoclavella striata Ponomarenko 1986
†Coptoclavella vittata Ponomarenko 1986
G. †Coptoclavisca Ponomarenko 1987
+
†Coptoclavisca nigricollinus Ponomarenko 1987
G. †Holcoptera Handlirsch 1906
+
†Holcoptera alisonae Kelly et al. 2017
†Holcoptera giebeli Handlirsch 1906
†Holcoptera pigmentatus Kelly et al. 2017
†Holcoptera rasnitsyni Ponomarenko and Fedorenko 2022
†Holcoptera schlotheimi Giebel 1856
+
Invalid names: Holcoptera confluens Cockerell 1915 [synonym]
†Holcoptera solitensis Kelly et al. 2017
Invalid names: Holcoelytrum Handlirsch 1906 [synonym], Stargelytron Ponomarenko 2015 [synonym]
G. †Euroscapha Lin 1992
+
Subfm. †Hispanoclavinae Soriano et al. 2007
+
G. †Hispanoclavina Soriano et al. 2007
+
†Hispanoclavina diazromerali Soriano et al. 2007
†Hispanoclavina gratshevi Soriano et al. 2007
Subfm. †Timarchopsinae Wang et al. 2010
+
G. †Daohugounectes Wang et al. 2009
+
†Daohugounectes primitivus Wang et al. 2009
G. †Protonectes Prokin and Ponomarenko 2013
+
†Protonectes germanicus Prokin and Ponomarenko 2013
G. †Timarchopsis Brauer et al. 1889
+
†Timarchopsis aquaticus Ponomarenko 1977
†Timarchopsis cyrenaicus Ponomarenko 1977
†Timarchopsis czekanowskii Brauer et al. 1889
†Timarchopsis gigas Ponomarenko 1977
†Timarchopsis gobiensis Ponomarenko 1987
†Timarchopsis latus Ponomarenko 1977
†Timarchopsis longus Ponomarenko 2014
†Timarchopsis mongolicus Ponomarenko 1985
†Timarchopsis sainshandensis Ponomarenko 1987
Invalid names: Necronectes Ponomarenko 1977 [synonym]
Invalid names: Necronectinae Ponomarenko 1977 [replaced]
G. †Curculionites Oppenheim 1888
+
†Curculionites striatus Oppenheim 1888
Fm. Dytiscidae Leach 1815 [predaceous diving beetle]
+
Subfm. Agabinae Thomson 1867
+
Tr. Agabini Thomson 1867
+
G. Hydrotrupes Sharp 1882
+
†Hydrotrupes prometheus Gómez and Damgaard 2014
G. Platambus Thomson 1859
+
Platambus optatus Sharp 1884
Platambus stygius Régimbart 1899
G. Agabus Leach 1817
+
Subg. Agabus (Agabus) Leach 1817
+
†Agabus charon Wickham 1912
†Agabus congeneroides Lomnicki 1894
†Agabus corticeus Lomnicki 1894
†Agabus florissantensis Wickham 1913
†Agabus hochi Statz 1939
†Agabus latissimus Ponomarenko 2019
†Agabus lithax Říha 1960
†Agabus niedzwiedzkii Lomnicki 1894
†Agabus parvulus Lomnicki 1894
†Agabus perditus Scudder 1900
†Agabus praelugens Wickham 1917
†Agabus rathbuni Scudder 1900
†Agabus reductus von Heyden and von Heyden 1866
†Agabus reitteri Lomnicki 1894
†Agabus rottensis Nilsson 2001
+
Invalid names: Agabus latus Statz 1939 [replaced]
†Agabus savagei Wickham 1917
†Agabus ungeri Heer 1847
Invalid names: Necticus Hope 1838 [synonym]
G. Ilybius Erichson 1832
+
Ilybius apicalis Sharp 1873
†Ilybius boryslavicus Lomnicki 1894
Ilybius fenestratus Fabricius 1781
†Ilybius gratshevi Ponomarenko 2019
Ilybius subaeneus Erichson 1837
Ilybius vittiger Gyllenhal 1827
G. †Angaragabus Ponomarenko 1963
+
†Angaragabus jurassicus Ponomarenko 1963
Subfm. Colymbetinae Erichson 1837
+
Tr. Colymbetini Erichson 1837
+
G. Colymbetes Clairville 1806
+
†Colymbetes aemulus Heer 1862
Colymbetes fuscus Linnaeus 1758
†Colymbetes miocaenicus Říha 1974
Colymbetes paykulli Erichson 1837
Invalid names: Cymatopterus Dejean 1833 [synonym]
G. Rhantus Dejean 1833
+
Rhantus exsoletus Forster 1771
†Rhantus praesuturellus Lomnicki 1894
†Rhantus villumi Prokin et al. 2024
Subfm. Copelatinae Van den Branden 1885
+
G. Copelatus Erichson 1832
+
†Copelatus aphroditae Balke 2003
†Copelatus chiapas Hájek et al. 2025
†Copelatus convexus Förster 1891
†Copelatus fossilis Říha 1974
†Copelatus ponomarenkoi Říha 1974
†Copelatus predaveterus Miller 2003
Invalid names: Copelatini Van den Branden 1885 [empty]
Subfm. Coptotominae Van den Branden 1885
+
Tr. Coptotomini Van den Branden 1885
+
G. Coptotomus Say 1830
+
†Coptotomus balticus Hendrich and Balke 2020
G. †Cretodytes Ponomarenko 1977
+
†Cretodytes incertus Prokin et al. 2013
†Cretodytes latipes Ponomarenko 1977
Subfm. Cybisterinae Sharp 1880
+
G. Cybister Curtis 1827
+
†Cybister agassizi Heer 1862
†Cybister atavus Heer 1862
†Cybister fractus Říha 1974
†Cybister imperfectus Říha 1974
†Cybister mancus Říha 1974
†Cybister rotundatus Říha 1974
G. Megadytes Sharp 1882
+
Subg. Megadytes (Paramegadytes) Trémouilles and Bachmann 1980
+
†Megadytes nicoleti Heer 1862
Subfm. Dytiscinae Leach 1815
+
Tr. Aciliini Thomson 1867
+
G. Acilius Leach 1817
+
†Acilius florissantensis Wickham 1909
†Acilius oeningensis Heer 1847
†Acilius praesulcatus Lomnicki 1894
G. Graphoderus Dejean 1833
+
†Graphoderus heeri Nilsson 2001
+
Invalid names: Dytiscus areolatus Heer 1862 [replaced]
†Graphoderus mirabilis Říha 1974
Tr. †Ambarticini Yang et al. 2019
+
G. †Ambarticus Yang et al. 2019
+
†Ambarticus myanmaricus Yang et al. 2019
Tr. Dytiscini Leach 1815
+
G. Dytiscus Linnaeus 1758
+
†Dytiscus avunculus von Heyden 1862
Dytiscus lapponicus Gyllenhal 1808
†Dytiscus latahensis Wickham 1931
†Dytiscus lavateri Heer 1847
Dytiscus marginalis Linnaeus 1758
†Dytiscus miocenicus Lewis and Gundersen 1987
†Dytiscus zersii Sordelli 1882
G. †Miodytiscus Wickham 1911
+
†Miodytiscus hirtipes Wickham 1911
Tr. Eretini Crotch 1873
+
G. Eretes Laporte 1833
+
†Eretes antiquus Oustalet 1870
Invalid names: Eunectes Erichson 1832 [replaced]
Tr. Hydaticini Sharp 1880
+
G. Hydaticus Leach 1817
+
Subg. Hydaticus (Hydaticus) Leach 1817
Subfm. Hydroporinae Aubé 1836
+
Tr. Bidessini Sharp 1880
+
G. Bidessus Sharp 1882
Invalid names: Bidessus laminarum Wickham 1914 [nomen dubium], Hydroporus sedimentorum Wickham 1914 [nomen dubium]
G. †Glessaria Koch and Berendt 1854
+
†Glessaria rostrata Koch and Berendt 1854
Tr. Hydroporini Aubé 1836
+
Subtr. Deronectina Galewski 1994
+
G. Boreonectes Angus 2010
G. Deronectes Sharp 1882
G. Nectoporus Guignot 1950
G. Oreodytes Seidlitz 1887
G. Hydrocolus Roughley and Larson 2000
+
Hydrocolus oblitus Aubé 1838
G. Hydroporus Clairville 1806
+
†Hydroporus carstengroehni Balke et al. 2010
†Hydroporus clessini Lomnicki 1894
Hydroporus discretus Fairmaire and Brisout 1859
†Hydroporus edenianus Lesne 1926
†Hydroporus inanimatus Scudder 1900
†Hydroporus inundatus Scudder 1900
†Hydroporus lapideus Říha 1974
Hydroporus lapponum Gyllenhal 1808
Hydroporus morio Aubé 1838
Hydroporus notabilis LeConte 1850
+
Invalid names: Hydroporus arcticus Thomson 1856 [synonym]
Hydroporus palustris Linnaeus 1761
Hydroporus planus Fabricius 1781
†Hydroporus pleistocenicus Lomnicki 1894
†Hydroporus praedorsalis Lomnicki 1894
†Hydroporus praenigrita Lomnicki 1894
†Hydroporus praenivalis Lomnicki 1894
†Hydroporus sandbergeri Lomnicki 1894
†Hydroporus sectus Scudder 1900
†Hydroporus statzi Nilsson 2001
+
Invalid names: Hydroporus macularis Statz 1939 [replaced]
Hydroporus striola Gyllenhal 1826
†Hydroporus subarcticus Lomnicki 1894
†Hydroporus trimaculatus Theobald 1937
G. Nebrioporus Régimbart 1906
+
†Nebrioporus benzeli Heer 1862
Invalid names: Potamonectes Zimmerman 1921 [synonym]
G. †Pteroporus Guignot 1933
+
†Pteroporus antiquus Heer 1862
Tr. Hygrotini Portevin 1929
+
G. Clemnius Villastrigo et al. 2017
+
Subg. Clemnius (Clemnius) Villastrigo et al. 2017
G. Hygrotus Stevens 1828
+
Subg. Hygrotus (Coelambus) Thomson 1860
Subg. Hygrotus (Hygrotus) Stephens 1828
Subg. Hygrotus (Leptolambus) Villastrigo et al. 2017
†Hygrotus cribrarius Scudder 1900
†Hygrotus derelictus Scudder 1900
†Hygrotus disjectus Scudder 1900
†Hygrotus dzieduszyckii Lomnicki 1894
†Hygrotus ganglbaueri Lomnicki 1894
†Hygrotus infernalis Scudder 1900
†Hygrotus latefasciatus Lomnicki 1894
†Hygrotus miocenus Wickham 1912
†Hygrotus niedzwiedzkii Lomnicki 1894
†Hygrotus ozokeriticus Lomnicki 1894
†Hygrotus picipoides Lomnicki 1894
Tr. Hyphydrini Gistel 1848
+
G. Hyphydrus Illiger 1802
Tr. Methlini Van den Branden 1885
+
G. Methles Sharp 1882
+
†Methles caucasicus Říha 1974
G. †Procoelambus Theobald 1937
+
†Procoelambus macrocephalus Theobald 1937
Tr. †Schistomerini Palmer 1957
+
G. †Schistomerus Palmer 1957
+
Tr. Vatellini Sharp 1880
+
G. †Calicovatellus Miller and Lubkin 2001
+
†Calicovatellus petrodytes Miller and Lubkin 2001
G. Derovatellus Sharp 1882
Subfm. Laccophilinae Gistel 1848
+
G. Japanolaccophilus Sato 1972
+
†Japanolaccophilus beatificus Balke and Hendrich 2019
Tr. Laccophilini Gistel 1848
+
G. †Electruphilus Balke et al. 2019
+
†Electruphilus wendeli Balke et al. 2019
G. Laccophilus Leach 1817
+
†Laccophilus corystes Zhang 1989
†Laccophilus pusulatus Zhang 1989
†Laccophilus strigatus von Schlechtendal 1894
Invalid names: Palaeogyrinus von Schlechtendal 1894 [synonym]
Invalid names: Palaeogyrininae von Schlechtendal 1894 [synonym]
Subfm. †Liadytiscinae Prokin and Ren 2010
+
G. †Liadroporus Prokin and Ren 2010
+
†Liadroporus elegans Prokin and Ren 2010
Tr. †Liadytiscini Prokin and Ren 2010
+
G. †Liadytiscus Prokin and Ren 2010
+
†Liadytiscus cretaceus Prokin and Ren 2010
†Liadytiscus latus Prokin and Ren 2010
†Liadytiscus longitibialis Prokin and Ren 2010
G. †Liadyxianus Prokin et al. 2013
+
†Liadyxianus kirejtshuki Prokin et al. 2013
Tr. †Mesoderini Prokin et al. 2013
+
G. †Mesoderus Prokin and Ren 2010
+
†Mesoderus magnus Prokin and Ren 2010
†Mesoderus ovatus Prokin et al. 2013
†Mesoderus punctatus Prokin et al. 2013
†Mesoderus ventralis Prokin and Ren 2010
G. †Mesodytes Prokin et al. 2013
+
†Mesodytes rhantoides Prokin et al. 2013
G. †Palaeodytes Ponomarenko 1987
+
†Palaeodytes baissiensis Prokin et al. 2013
†Palaeodytes gutta Ponomarenko 1987
†Palaeodytes incompleta Ponomarenko et al. 2005
†Palaeodytes sibiricus Ponomarenko 1987
G. †Sinoporus Prokin and Ren 2010
+
†Sinoporus lineatus Prokin and Ren 2010
Invalid names: Aubehydrinae Guignot 1942 [empty], Necticus palustris Aymard 1856 [nomen nudum]
Fm. Gyrinidae Latreille 1810 [whirligig beetle]
+
G. †Anagyrinus Handlirsch 1906
+
†Anagyrinus atavus Heer 1864
G. †Coptogyrinus Handlirsch 1906
+
†Coptogyrinus scutellatus Handlirsch 1906
Subfm. Gyrininae Latreille 1810
+
G. †Chimerogyrus Gustafson et al. 2020
+
†Chimerogyrus gigagalea Gustafson et al. 2020
Tr. Enhydrusini Branden 1882
+
Invalid names: Dineutina Desmarest 1851 [empty], Dineutini Desmarest 1851 [replaced]
Tr. Gyrinini Latreille 1810
+
G. †Gyrinoides Motschulsky 1856
+
†Gyrinoides limbatus Motschulsky 1856
G. Gyrinus Geoffroy 1762
+
Gyrinus confinis Le Conte 1868
Gyrinus marinus Gyllenhal 1808
+
Invalid names: Gyrinus praemarinus Lomnicki 1894 [synonym]
Gyrinus minutus Fabricius 1798
Gyrinus opacus Sahlberg 1819
Gyrinus praeopacus Lomnicki 1894
Tr. Orectochilini Kolbe 1880
+
G. Gyretes Brullé 1835
+
†Gyretes giganteus Piton 1940
G. Orectochilus Dejean 1833
+
Orectochilus villosus Müller 1776
G. †Gyrinopsis Handlirsch 1906
+
†Gyrinopsis antiquus Heer 1864
G. †Protogyrininus Hatch 1927
+
†Protogyrininus sculpturatus Mjöberg 1905
+
Invalid names: Protogyrininus confinis Hatch 1927 [synonym]
G. †Tunguskagyrus Yan et al. 2018
+
†Tunguskagyrus planus Yan et al. 2018
†Tunguskagyrus yani Ponomarenko 2021
Invalid names: Spanglerogyrinae Folkerts 1979 [empty]
Subfm. Haliplidae Aubé 1836 [crawling water beetle]
+
G. †Cretihaliplus Ren et al. 1995
+
†Cretihaliplus chifengensis Ren et al. 1995
†Cretihaliplus sidaojingensis Ren et al. 1995
G. Haliplus Latreille 1802
+
†Haliplus cretaceus Prokin and Ponomarenko 2013
Haliplus obliquus Fabricius 1787
Haliplus ruficollis De Geer 1774
G. Peltodytes Regimbart 1878
Fm. Hygrobiidae Regimbart 1878
+
G. Hygrobia Latreille 1804
+
†Hygrobia cretzschmari von Heyden and von Heyden 1866
Invalid names: Paelobius Schoenherr 1808 [synonym]
Invalid names: Paelobiidae Erichson 1837 [synonym]
Fm. †Liadytidae Ponomarenko 1977
+
G. †Liadytes Ponomarenko 1963
+
†Liadytes aspidytoides Prokin et al. 2013
†Liadytes avus Ponomarenko 1963
†Liadytes crassus Ponomarenko 1977
†Liadytes dajensis Ponomarenko 1987
†Liadytes longus Ponomarenko 1977
†Liadytes major Ponomarenko 1985
G. †Ovidytes Ren et al. 1995
+
†Ovidytes gaoi Ren et al. 1995
G. †Longurius Hong 1982
+
†Longurius xigeduensis Hong 1982
G. †Memptus Handlirsch 1906
+
†Memptus ganglbaueri Ponomarenko 1985
†Memptus guttula Ponomarenko 1985
†Memptus handlirschi Ponomarenko 1985
†Memptus redtenbacheri Handlirsch 1906
+
Invalid names: Memptus braueri Handlirsch 1906 [synonym]
Fm. Noteridae Thomson 1860 [burrowing water beetle]
+
Subfm. Noterinae Thomson 1860
+
Tr. Noterini Thomson 1860
+
G. Canthydrus Sharp 1882
G. Hydrocanthus Say 1823
G. Noterus Clairville 1806
+
Noterus crassicornis Müller 1776
Noterus japonicus Sharp 1873
Invalid names: Hydrocanthini Sharp 1880 [synonym]
Subfm. Phreatodytinae Ueno 1957
+
G. Phreatodytes Ueno 1957
G. †Oryctites Oppenheim 1888
+
†Oryctites fossilis Oppenheim 1888
Fm. †Parahygrobiidae Ponomarenko 1977
+
G. †Parahygrobia Ponomarenko 1977
+
†Parahygrobia natans Ponomarenko 1977
Fm. Trachypachidae Thomson 1857 [false ground beetle]
+
G. †Apermunda Kirejtshuk and Ansorge 2022
+
†Apermunda minor Ponomarenko and Volkov 2013
G. †Denudirabus Ren 1995
+
G. †Dundorabus Ponomarenko 1989
+
†Dundorabus glabrus Ponomarenko 1989
Subfm. †Eodromeinae Ponomarenko 1977
+
G. †Ademosynoides Dunstan 1923
+
†Ademosynoides abnormis Dunstan 1923
†Ademosynoides alternata Dunstan 1923
†Ademosynoides angusta Tillyard 1916
†Ademosynoides japonicus Fujiyama 1973
†Ademosynoides magnifica Dunstan 1923
†Ademosynoides minor Handlirsch 1906
†Ademosynoides obtusa Tillyard 1916
†Ademosynoides striatella Dunstan 1923
G. †Coreoeicos Bode 1953
+
†Coreoeicos dilatatus Bode 1953
G. †Dolichorabus Ponomarenko 1985
+
†Dolichorabus longipes Ponomarenko 1985
G. †Eodromeus Ponomarenko 1977
+
†Eodromeus antiquus Ponomarenko 1977
†Eodromeus daohugouensis Wang et al. 2012
†Eodromeus dissectus Ponomarenko 1977
†Eodromeus major Ponomarenko 1977
†Eodromeus mongolicus Ponomarenko 1989
†Eodromeus robustus Wang et al. 2012
†Eodromeus sternalis Ponomarenko 1977
†Eodromeus sulcatus Ponomarenko 1989
G. †Evertus Ponomarenko 1986
+
†Evertus cornutus Ponomarenko 1986
†Evertus notatus Ponomarenko 1986
†Evertus pervalimand Jia and Ren 2011
Invalid names: Fortiseode Jia and Ren 2011 [synonym]
G. †Karadromeus Ponomarenko 1977
+
†Karadromeus capitatus Ponomarenko 1986
†Karadromeus elongatus Ponomarenko 1977
†Karadromeus erensis Ponomarenko 1986
†Karadromeus gobiensis Ponomarenko 1980
†Karadromeus latus Ponomarenko 1977
†Karadromeus mongolicus Ponomarenko 1977
†Karadromeus rostratus Ponomarenko 1977
†Karadromeus verrucosus Ponomarenko 1989
G. †Karatoma Ponomarenko 1977
+
†Karatoma agilis Ponomarenko 1977
†Karatoma raptor Ponomarenko 1977
G. †Liassodites Kirejtshuk and Ansorge 2022
+
†Liassodites obsti Kirejtshuk and Ansorge 2022
G. †Necronectulus Ponomarenko 1977
+
†Necronectulus avus Ponomarenko 1977
†Necronectulus lazarus Lee et al. 2024
G. †Novunda Kirejtshuk and Ansorge 2022
+
†Novunda angulata Ponomarenko 1977
†Novunda chifengensis Wang et al. 2012
†Novunda cursoria Ponomarenko 1977
†Novunda microplata Ponomarenko 1977
†Novunda pachycephala Ponomarenko 1989
Invalid names: Unda Ponomarenko 1977 [replaced]
G. †Permunda Ponomarenko and Volkov 2013
+
†Permunda nana Ponomarenko and Volkov 2013
G. †Petrodromeus Ponomarenko and Volkov 2013
+
†Petrodromeus asiaticus Martynov 1936
G. †Platycoxa Ponomarenko 1977
+
†Platycoxa armata Ponomarenko 1977
†Platycoxa jurassica Ponomarenko 1977
G. †Prosynactus Bode 1953
+
†Prosynactus gracilis Bode 1953
+
Invalid names: Cardioides incisus Bode 1953 [synonym], Cyclotrachelus exsecatus Bode 1953 [synonym], Cyphospheron virgatus Bode 1953 [synonym], Liassocarabites pulcher Bode 1953 [synonym], Mesalocistus constrictus Bode 1953 [synonym], Rhabdotus cingulatus Bode 1953 [synonym]
†Prosynactus procerus Bode 1953
+
Invalid names: Dysmorphus molestus Bode 1953 [synonym]
†Prosynactus scissus Bode 1953
+
Invalid names: Cardioides longealatus Bode 1953 [synonym], Euenarthrus mandibulatus Bode 1953 [synonym], Liassocarabites praefrictus Bode 1953 [synonym], Megelytrites mutilatus Bode 1953 [synonym], Trapezotrachelus longus Bode 1953 [synonym], Trigonocephalites sulcatus Bode 1953 [synonym]
Invalid names: Critotrachelus Carpenter 1986 [synonym], Cyclotrachelus Bode 1953 [synonym], Dysmorphus Bode 1953 [synonym], Euenarthrus Bode 1953 [synonym], Leptynticus Bode 1953 [synonym], Liassocarabites Bode 1953 [synonym], Megelytrites Bode 1953 [synonym], Mesalocistus Bode 1953 [synonym], Rhabdotus Bode 1953 [synonym], Tetragonotrachelus Bode 1953 [synonym], Trapezotrachelus Bode 1953 [synonym], Trigonocephalites Bode 1953 [synonym]
G. †Psacodromeus Ponomarenko 1977
+
†Psacodromeus crassus Ponomarenko 1977
†Psacodromeus gutta Ponomarenko 1977
†Psacodromeus minor Ponomarenko 1986
†Psacodromeus ovalis Ponomarenko 1977
†Psacodromeus recta Lee et al. 2024
†Psacodromeus rugosus Ponomarenko 1977
G. †Sinodromeus Wang et al. 2012
+
†Sinodromeus liutiaogouensis Wang et al. 2012
G. †Sogdodromeus Ponomarenko 1977
+
†Sogdodromeus altus Ponomarenko 1977
G. †Tauredon Handlirsch 1910
+
†Tauredon horni Handlirsch 1910
G. †Xishanocarabus Hong 1984
+
†Xishanocarabus parvus Hong 1984
G. †Eodromeites Ponomarenko 1985
+
†Eodromeites minimus Ponomarenko 1985
†Eodromeites udensis Ponomarenko 1985
Subfm. Trachypachinae Thomson 1857
+
Tr. Trachypachini Thomson 1857
+
G. Trachypachus Motschulsky 1845
+
Trachypachus holmbergi Mannerheim 1853
Invalid names: Karadromeus xiangfanggouensis Hong 1995 [nomen dubium]
G. †Triadogyrus Ponomarenko 1977
+
†Triadogyrus sternalis Ponomarenko 1977
Fm. †Triaplidae Ponomarenko 1977
+
G. †Avocatinus Ponomarenko 1969
+
†Avocatinus elongatus Ponomarenko 1969
G. †Catinius Ponomarenko 1968
+
†Catinius artus Zhang 1997
†Catinius ovatus Zhang 1997
†Catinius pelta Ponomarenko 1968
G. †Catinoides Ponomarenko 1969
+
†Catinoides rotundatus Ponomarenko 1969
G. †Clypeus Hong 1983
+
†Clypeus korlaensis Hong 1983
G. †Macrocatinius Ponomarenko 1969
+
†Macrocatinius brachycephalus Ponomarenko 1969
G. †Tomiaplus Ponomarenko 2021
+
†Tomiaplus minimus Ponomarenko 2021
†Tomiaplus sibiricus Volkov 2013
G. †Triaplus Ponomarenko 1977
+
†Triaplus laticoxa Ponomarenko 1977
†Triaplus macroplatus Ponomarenko 1977
G. †Triassocatinius Ponomarenko 1969
+
†Triassocatinius brachynotus Ponomarenko 1969
†Triassocatinius glabratus Ponomarenko 1969
Invalid names: Catiniidae Ponomarenko 1968 [replaced]
Invalid names: Amphizoidae Leconte 1853 [empty], Dytiscoidea Leach 1815 [invalid subgroup], Pseudomorphinae Hope 1838 [empty], Tritarsusidae Hong 2002 [invalid subgroup]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Bush and Bambach 2015, Kiessling 2004

Age range

Maximum range based only on fossils: base of the Changhsingian to the top of the Holocene or 254.14000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 251.902 Ma

Collections (701 total)

Oldest occurrences

Time interval Ma Country or state Original ID and collection number
Changhsingian254.14 - 251.902Russian Federation (Evenki Autonomous Okrug) Tunguskagyrus planus, Tunguskagyrus yani (118027) Tunguskagyrus yani (105049)
Griesbachian251.902 - 250.5Russian Federation (Kemerovo) Ademosynoides asiaticus (105040) Tomiaplus minimus, Triaplus sibiricus, Permunda nana, Petrodromeus asiaticus, Petrodromeus minor (104713)
Early/Lower Triassic251.902 - 246.7India (Madhya Pradesh) Protodytiscus johillaensis (151843)
Bithynian244.99 - 244.24Germany (Bavaria) Stargelytron altus (115969)
Bithynian244.99 - 244.24Germany Stargelytron altus (175694)
Ladinian241.464 - 237.0Kyrgyzstan (Osh) Avocatinus elongatus, Triaplus laticoxa (110763) Avocatinus elongatus, Triassocatinius brachynotus, Catinoides rotundatus, Triaplus laticoxa (106090) Macrocatinius brachycephalus (113406) Triaplus laticoxa (106088) Triaplus macroplatus (110767 123145) Triaplus macroplatus, Sogdodromeus altus (110765) Triassocatinius glabratus, Triaplus laticoxa (113468)
Longobardian239.48 - 237.0China (Shaanxi) Dytiscoidea indet. (183209)
Carnian237.0 - 227.3Japan Ademosynoides japonicus (155678)
Carnian237.0 - 227.3China (Xinjiang) Agrascapha curta, Euroscapha rectabdomena, Euroscapha hoplopoda, Ademosynoides minor (156325)
Late/Upper Triassic237.0 - 201.4Angola Coptoclavia africana (210174)
Tuvalian233.6 - 227.3Germany Coptoclavidae indet. (163985) Protonectes germanicus, Carabilarva triassica, Coptoclavidae indet. (153474) Stargelytron larissae (175695)
Norian227.3 - 205.7Kazakhstan Colymbotethis antecessor (113404) Necronectulus avus (114782) Platycoxa jurassica (105055)
Norian227.3 - 205.7Ukraine (Kharkiv) Triadogyrus sternalis, Holcoptera rasnitsyni (105057)
Norian227.3 - 205.7Australia (Queensland) Ademosyne minor (111854) Ademosyne minor, Ademosynoides striatella, Ademosyne obtusa, Ademosyne angusta (111777) Ademosynoides alternata, Ademosynoides magnifica, Ademosynoides abnormis (111558)
Norian227.3 - 205.7USA (North Carolina) Holcoptera solitensis (189406 189407)
Sevatian211.18 - 205.7Italy (Lombardia) Holcoptera schlotheimi (94207)
Rhaetian205.7 - 201.4United Kingdom (England) Harpalinae indet., Harpalinae indet. 1 (114643) Holcoptera giebeli, Holcoptera schlotheimi (113961) Holcoptera pigmentatus (113957) Holcoptera schlotheimi (114644 115223 115246 187761)
Rhaetian - Early/Lower Hettangian205.7 - 199.5United Kingdom (England) Holcoptera confluens, Coleoptera indet. 4, Coleoptera indet. 5, Coleoptera indet. 6, Coleoptera indet. 3 (124688)
Angulata201.4 - 199.5Switzerland Gyrinites antiquus, Gyrinus atavus, Carabites anthracinus (131766)
Early/Lower Hettangian201.4 - 199.5United Kingdom (England) Holcoptera schlotheimi (113956 113958 123986)