PBDB Taxon

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Trigonotarbida

Arachnida - Trigonotarbida

Taxonomy

Trigonotarbi was named by Petrunkevitch (1949).

It was corrected as Trigonotarbida by Dunlop (1996), Brauckmann et al. (2003), Shultz (2007), Garwood and Dunlop (2014), Poschmann et al. (2016).

It was assigned to Arachnida by Petrunkevitch (1949), Dunlop (1996), Brauckmann et al. (2003); and to Pantetrapulmonata by Shultz (2007), Garwood and Dunlop (2014), Poschmann et al. (2016).

Synonyms

Anthracomartus woodruffi was named by Scudder (1893). It is an impression. Its type locality is Pawtucket, which is in a Westphalian terrestrial horizon in Rhode Island.

It was recombined as Trigonomartus woodruffi by Petrunkevitch (1913), Petrunkevitch (1949), Petrunkevitch (1953); it was considered a nomen dubium by Dunlop et al. (2013).
Termes hageni was named by Goldenberg (1873). Its type specimen is Naturhist. Ver. Bonn, 17, a wing, and it is an impression. Its type locality is Altenwald mine, which is in a Westphalian C terrestrial shale/bituminous coal in the Lower Saarbrucker layer Formation of Germany.

It was recombined as Palaeodictyopteron hageni by Handlirsch (1906), von Schlechtendal (1913), Handlirsch (1919); it was considered a nomen dubium by Guthörl (1934), Carpenter (1948), Petrunkevitch (1953), Dunlop et al. (2013).
Anthracomarti was named by Karsch (1882).

It was reranked as the suborder Anthracomarti by Haase (1890); it was replaced with Trigonotarbida by Dunlop et al. (2013).

It was assigned to Opiliones by Haase (1890); to Arachnoidea by Guthörl (1934); and to Arachnida by Petrunkevitch (1945).
Termes buchi was named by Goldenberg (1873). It is not extant. Its type specimen is Naturhist. Ver. Bonn, 18, an exoskeleton. Its type locality is Jaegersfreude mine, which is in a Westphalian C lacustrine shale/coal in the Upper Saarbrucker layer Formation of Germany.

It was recombined as Anthracomartus buchi by Guthörl (1934), Petrunkevitch (1953); it was considered a nomen dubium by Dunlop et al. (2013).
Elaverimartus was named by Petrunkevitch (1953).

It was considered a nomen dubium by Dunlop et al. (2013).
Elaverimartus pococki was named by Petrunkevitch (1953) [Dunlop et al. 2013 argue this is a nomen dubium due to incomplete preservation]. Its type specimen is NMS G.1957.1.4994, an exoskeleton, and it is a compression fossil. Its type locality is Ellismuir, Baillieston, which is in a Westphalian A terrestrial siliciclastic in the Kiltoung Coal Formation of the United Kingdom.

It was considered a nomen dubium by Dunlop et al. (2013), Dunlop et al. (2013).
Eurymartus spinulosus was named by Matthew (1895). Its type specimen is NBMG 3004, an exoskeleton, and it is an impression. Its type locality is Fern Ledges, Plant bed 2, Carleton, Saint John, which is in a Westphalian A estuary/bay shale in the Lancaster Formation of Canada.

It was considered a nomen dubium by Dunlop et al. (2013).
Eurymarti was named by Matthew (1895).

It was synonymized subjectively with Trigonotarbida by Dunlop et al. (2013).
Eurymartidae was named by Matthew (1895).

It was considered an invalid subgroup of Trigonotarbida by Dunlop et al. (2013).

It was assigned to Eurymarti by Matthew (1895).
Eophrynus scharfi was named by Scharf (1924). Its type specimen is Geiseltalmuseum SCHA1924.8.MLU and is an impression. Its type locality is Ottostollen mine tip, Ilfeld, which is in an Asselian fluvial-lacustrine siliciclastic in the Netzkater Formation of Germany.

It was considered a nomen dubium by Rössler et al. (2003).

Synonymy list

Year	Name and author
1873	Termes buchi Goldenberg p. 12 figs. Pl 2, fig 9
1873	Termes hageni Goldenberg p. 12 figs. Pl 2, fig 7b
1882	Anthracomarti Karsch p. 560
1890	Anthracomarti Haase p. 650
1893	Anthracomartus woodruffi Scudder p. 9 figs. Pl I f
1895	Eurymartus spinulosus Matthew p. 276 figs. Pl 2, fig 9
1895	Eurymarti Matthew p. 277
1895	Eurymartidae Matthew p. 277
1906	Palaeodictyopteron hageni Handlirsch p. 62
1913	Trigonomartus woodruffi Petrunkevitch p. 105 figs. Pl 9, fig 52
1913	Palaeodictyopteron hageni von Schlechtendal p. 99 figs. Pl 1, fig 7; pl 8, fig 1
1919	Palaeodictyopteron hageni Handlirsch p. 536
1924	Eophrynus scharfi Scharf p. 433 fig. 95
1934	Anthracomarti Guthörl
1934	Anthracomartus buchi Guthörl
1945	Anthracomarti Petrunkevitch p. 46
1949	Trigonotarbi Petrunkevitch p. 235
1949	Trigonomartus woodruffi Petrunkevitch p. 257
1953	Anthracomartus buchi Petrunkevitch p. 68
1953	Trigonomartus woodruffi Petrunkevitch p. 90
1953	Elaverimartus Petrunkevitch p. 92
1953	Elaverimartus pococki Petrunkevitch p. 92 figs. 96, 171
1996	Trigonotarbida Dunlop
2003	Trigonotarbida Brauckmann et al. p. 46
2007	Trigonotarbida Shultz
2014	Trigonotarbida Garwood and Dunlop p. 22
2016	Trigonotarbida Poschmann et al. p. 36

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Ecdysozoa	Aguinaldo et al. 1997
—	Panarthropoda
phylum	Arthropoda	Latreille 1829
—	Deuteropoda
subphylum	Chelicerata	Heymons 1901

Rank	Name	Author
—	Euchelicerata	Weygoldt and Paulus 1979
—	Prosomapoda
—	Planaterga
—	Dekatriata
—	Sclerophorata	Kamenz et al. 2011
class	Arachnida	Lamarck 1801
—	Pantetrapulmonata
order	Trigonotarbida	()
order	Trigonotarbida	()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Or. †Trigonotarbida Petrunkevitch 1949

show all | hide all

G. †Aenigmatarbus Poschmann et al. 2016

†Aenigmatarbus rasteli Poschmann et al. 2016

Fm. †Anthracomartidae Haase 1890

G. †Anthracomartus Karsch 1882

†Anthracomartus bohemica Fritsch 1895

†Anthracomartus carcinoides Fritsch 1899

Invalid names: Arthrolycosa salticoides Fritsch 1899 [synonym], Promygale rotundata Fritsch 1899 [synonym]

†Anthracomartus elegans Fritsch 1899

†Anthracomartus granulatus Fritsch 1904

†Anthracomartus hindi Pocock 1911

Invalid names: Cleptomartus planus Petrunkevitch 1949 [synonym], Cryptomartus meyeri Guthörl 1964 [synonym]

†Anthracomartus janae Opluštil 1986

†Anthracomartus kustae Petrunkevitch 1953

†Anthracomartus minor Kušta 1885

Invalid names: Anthracomartus socius Kušta 1889 [synonym]

†Anthracomartus nyranensis Petrunkevitch 1953

†Anthracomartus palatinus von Ammon 1901

†Anthracomartus priesti Pocock 1911

Invalid names: Anthracomartus denuiti Pruvost 1922 [synonym], Cleptomartus plautus Petrunkevitch 1949 [synonym]

†Anthracomartus radvanicensis Oplustil 1985

†Anthracomartus triangularis Petrunkevitch 1913

†Anthracomartus trilobitus Scudder 1884

†Anthracomartus voelkelianus Karsch 1882

Invalid names: Brachylycosa Fritsch 1904 [synonym], Cleptomartus Petrunkevitch 1949 [synonym], Coryphomartus Petrunkevitch 1945 [synonym], Cryptomartus Petrunkevitch 1945 [synonym], Oomartus Petrunkevitch 1953 [synonym], Perneria Fritsch 1904 [synonym], Pleomartus Petrunkevitch 1945 [synonym], Promygale Fritsch 1899 [synonym]

G. †Brachypyge Woodward 1878

†Brachypyge carbonis Woodward 1878

G. †Maiocercus Pocock 1911

†Maiocercus celticus Pocock 1902

Invalid names: Maiocercus orbicularis Gill 1911 [synonym]

Invalid names: Brachypygidae Pocock 1911 [synonym], Coryphomartidae Petrunkevitch 1945 [synonym], Eurymartus Matthew 1895 [nomen dubium], Eurymartus latus Matthew 1895 [nomen dubium], Pleomartidae Petrunkevitch 1945 [synonym], Promygalidae Fritsch 1904 [synonym]

G. †Anthracophrynus Andrée 1913

†Anthracophrynus tuberculatus Andrée 1913

Fm. †Anthracosironidae Pocock 1903

G. †Anthracosiro Pocock 1903

†Anthracosiro fritschii Pocock 1903

Invalid names: Anthracosiro elongatus Waterlot 1934 [synonym]

†Anthracosiro woodwardi Pocock 1903

Invalid names: Anthracosiro corsini Pruvost 1926 [synonym], Anthracosiro latipes Gill 1909 [synonym]

G. †Arianrhoda Dunlop and Selden 2004

†Arianrhoda bennetti Dunlop and Selden 2004

G. †Vratislavia Fritsch 1904

†Vratislavia silesiaca Römer 1878

Fm. †Aphantomartidae Petrunkevitch 1945

G. †Alkenia Størmer 1970

†Alkenia mirabilis Størmer 1970

G. †Aphantomartus Pocock 1911

†Aphantomartus areolatus Pocock 1911

Invalid names: Aphantomartus pococki Pruvost 1912 [synonym], Eophrynus waechteri Guthörl 1938 [synonym], Trigonomartus dorlodoti Pruvost 1930 [synonym]

†Aphantomartus ilfeldicus Scharf 1924

†Aphantomartus pustulatus Scudder 1884

Invalid names: Cleptomartus ploetzensis Simon 1971 [synonym], Kreischeria villeti Pruvost 1912 [synonym]

Invalid names: Phrynomartus Petrunkevitch 1945 [synonym], Trigonomartus Petrunkevitch 1913 [synonym]

G. †Doubravatarbus Hradská et al. 2022

†Doubravatarbus krafti Hradská et al. 2022

Invalid names: Trigonomartidae Petrunkevitch 1945 [synonym]

Fm. †Archaeomartidae Poschmann and Dunlop 2010

G. †Archaeomartus Størmer 1970

†Archaeomartus levis Størmer 1970

Invalid names: Archaeomartus tuberculatus Størmer 1970 [synonym]

G. †Areomartus Petrunkevitch 1913

†Areomartus ovatus Petrunkevitch 1913

Fm. †Eophrynidae Karsch 1882

G. †Eophrynus Woodward 1871

†Eophrynus prestvicii Buckland 1837

†Eophrynus udus Brauckmann et al. 1985

†Euphrynus salmi Stur 1877

Invalid names: Duma Giebel 1856 [nomen oblitum]

G. †Nyranytarbus Harvey and Selden 1995

†Nyranytarbus hofmanni Fritsch 1899

†Nyranytarbus longipes Fritsch 1899

Invalid names: Hemiphrynus Fritsch 1899 [replaced]

G. †Petrovicia Fritsch 1904

†Petrovicia proditoria Fritsch 1904

G. †Planomartus Petrunkevitch 1953

†Planomartus krejcii Kušta 1884

Invalid names: Anthracomartus affinis Kušta 1885 [synonym]

G. †Pleophrynus Petrunkevitch 1945

†Pleophrynus ensifer Petrunkevitch 1945

†Pleophrynus hawesi Dunlop et al. 2014

†Pleophrynus verrucosus Pocock 1911

Invalid names: Eophrynus jugatus Ambrose and Romano 1972 [synonym], Eophrynus warei Dix and Pringle 1930 [synonym]

Invalid names: Acrokreischeria Petrunkevitch 1953 [synonym]

G. †Pocononia Petrunkevitch 1953

†Pocononia whitei Ewing 1930

G. †Somaspidion Jux 1982

†Somaspidion hammapheron Jux 1982

G. †Stenotrogulus Fritsch 1904

†Stenotrogulus salmii Stur 1877

Invalid names: Cyclotrogulus sturii Fritsch 1904 [synonym]

Invalid names: Cyclotrogulus Fritsch 1904 [synonym], Pseudoeophrynus Pribyl 1958 [synonym]

Invalid names: Hemiphrynidae Fritsch 1904 [synonym], Pleophrynidae Petrunkevitch 1945 [synonym]

Fm. †Kreischeriidae Haase 1890

G. †Anzinia Petrunkevitch 1953

†Anzinia thevenini Pruvost 1919

G. †Gondwanarachne Pinto and Hunicken 1980

†Gondwanarachne argentinensis Pinto and Hünicken 1980

G. †Hemikreischeria Fritsch 1904

†Hemikreischeria geinitzi Thevenin 1902

Invalid names: Hemikreischeria thevenini Fritsch 1904 [objective synonym]

G. †Kreischeria Geinitz 1882

†Kreischeria bohemica Fritsch 1895

†Kreischeria wiedei Geinitz 1882

G. †Pseudokreischeria Petrunkevitch 1953

†Pseudokreischeria pococki Gill 1924

Invalid names: Eophrynus varius Petrunkevitch 1949 [synonym]

Fm. †Lissomartidae Dunlop 1995

G. †Lissomartus Petrunkevitch 1949

†Lissomartus carbonarius Petrunkevitch 1913

†Lissomartus schucherti Petrunkevitch 1913

G. †Namurotarbus Poschmann and Dunlop 2010

†Namurotarbus roessleri Dunlop and Brauckmann 2006

Fm. †Palaeocharinidae Hirst 1923

G. †Aculeatarbus Shear et al. 1987

†Aculeatarbus depressus Shear et al. 1987

G. †Gelasinotarbus Shear et al. 1987

†Gelasinotarbus bifidus Shear et al. 1987

†Gelasinotarbus bonamoae Shear et al. 1987

†Gelasinotarbus fimbriunguis Shear et al. 1987

†Gelasinotarbus heptops Shear et al. 1987

†Gelasinotarbus reticulatus Shear et al. 1987

G. †Gigantocharinus Shear 2000

†Gigantocharinus szatmaryi Shear 2000

G. †Gilboarachne Shear et al. 1987

†Gilboarachne griersoni Shear et al. 1987

G. †Palaeocharinus Hirst 1923

†Palaeocharinus calmani Hirst 1923

†Palaeocharinus hornei Hirst 1923

†Palaeocharinus kidstoni Hirst 1923

†Palaeocharinus rhyniensis Hirst 1923

†Palaeocharinus scourfieldi Hirst 1923

†Palaeocharinus tuberculatus Fayers et al. 2005

Invalid names: Palaeocharinoides Hirst 1923 [synonym]

G. †Spinocharinus Poschmann and Dunlop 2011

†Spinocharinus steinmeyeri Poschmann and Dunlop 2011

G. †Palaeocteniza Hirst 1923

†Palaeocteniza crassipes Hirst 1923

G. †Palaeotarbus Dunlop 1999

†Palaeotarbus jerami Dunlop 1996

Invalid names: Eotarbus Dunlop 1996 [replaced]

G. †Permotarbus Dunlop and Rößler 2013

†Permotarbus schuberti Dunlop and Rößler 2013

Fm. †Trigonotarbidae Petrunkevitch 1949

G. †Trigonotarbus Pocock 1911

†Trigonotarbus arnoldi Petrunkevitch 1955

†Trigonotarbus carbonarius Petrunkevitch 1913

†Trigonotarbus johnsoni Pocock 1911

†Trigonotarbus stoermeri Schultka 1991

G. †Tynecotarbus Hradská and Dunlop 2013

†Tynecotarbus tichaveki Hradská and Dunlop 2013

Invalid names: Anthracomarti Karsch 1882 [replaced], Anthracomartus hageni Goldenberg 1873 [nomen dubium], Elaverimartus Petrunkevitch 1953 [nomen dubium], Elaverimartus pococki Petrunkevitch 1953 [nomen dubium], Eophrynus scharfi Scharf 1924 [nomen dubium], Eurymarti Matthew 1895 [synonym], Eurymartidae Matthew 1895 [invalid subgroup], Eurymartus spinulosus Matthew 1895 [nomen dubium], Termes buchi Goldenberg 1873 [nomen dubium], Trigonomartus woodruffi Scudder 1893 [nomen dubium]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

chitinc

Grouping:

solitaryc

Environment:

terrestrialc

Locomotion:

actively mobilec

Created:

2017-04-17 09:55:58

Modified:

2017-04-17 09:55:58

Source: c = class

Reference: Bush and Bambach 2015

Age range: base of the Pridoli to the top of the Artinskian or 422.70000 to 283.30000 Ma

Collections (115 total)

Time interval	Ma	Country or state	Original ID and collection number
Pridoli	422.7 - 419.62	United Kingdom (England)	Trigonotarbida indet. (type locality: 11128)
Lochkovian	419.62 - 413.02	United Kingdom	Arianrhoda bennetti (153478)
Pragian	413.02 - 410.62	United Kingdom (Scotland)	Palaeocharinus kidstoni (10721) Palaeocharinus sp. (153498) Palaeocharinus sp., Palaeocharinoides hornei, Palaeocharinus rhyniensis, Palaeocharinus scourfieldi, Palaeocharinus calmani (172777) Palaeocharinus sp., Palaeocteniza crassipes (172778) Palaeocharinus tuberculatus (153492)
Pragian	413.02 - 410.62	Germany	Spinocharinus steinmeyeri, Archaeomartus levis (155760) Trigonotarbida indet. (152107)
Early/Lower Emsian	410.62 - 399.5	Germany	Alkenia mirabilis (155774) Archaeomartus levis, Alkenia mirabilis (155757) Trigonotarbus stoermeri (172343)
Early/Lower Emsian	410.62 - 399.5	Germany (Alken)	Alkenia mirabilis, Archaeomartus tuberculatus (27049)
Early/Lower Givetian	387.95 - 385.9	USA (New York)	Gelasinotarbus fimbriunguis, Gilboarachne griersoni, Gelasinotarbus reticulatus, Gelasinotarbus bonamoae (153480)
Givetian	387.95 - 382.31	USA (New York)	Gelasinotarbus fimbriunguis, Gilboarachne griersoni, Gelasinotarbus reticulatus, Gelasinotarbus bonamoae (21172) Gilboarachne griersoni, Gelasinotarbus reticulatus, Gelasinotarbus bonamoae, Gelasinotarbus bifidus, Gelasinotarbus heptops, Aculeatarbus depressus (28246)
Late/Upper Famennian	365.2 - 358.86	USA (Pennsylvania)	Gigantocharinus szatmaryi (35044)
Chadian	346.7 - 339.9	USA (Virginia)	Trigonomartus whitei (155455)
Asbian	339.9 - 335.9	Germany	Cryptomartus sp. (156764)
Brigantian	335.9 - 330.3	Germany	Anthracomartus sp., Trigonomartus pustulatus (153594)
Brigantian - Serpukhovian	335.9 - 323.4	Canada (Nova Scotia)	Anthracomartus sp. (155663)
Arnsbergian	326.9 - 323.6	Czech Republic	Euphrynus salmi (38971)
Morrowan	323.4 - 318.6	USA (Arkansas)	Anthracomartus trilobitus (153485)
Alportian	322.8 - 321.8	Austria	Aphantomartus pustulatus (153488)
Marsdenian	321.3 - 320.0	Germany (Ruhr)	Eophrynus udus (155259) Trigonotarbus johnsoni (152121)
Westphalian A	318.7 - 316.9	Canada (New Brunswick)	Eurymartus latus, Eurymartus spinulosus (114468) Eurymartus sp. (114467)
Westphalian A	318.7 - 316.9	Canada (Nova Scotia)	Anthracomartus triangularis (146858)
Westphalian A	318.7 - 316.9	United Kingdom (Lancashire)	Maiocercus orbicularis (153490)
Westphalian A	318.7 - 316.9	United Kingdom	Aphantomartus areolatus (146813) Nemastomoides elaveris (146854)
Westphalian A	318.7 - 316.9	France (Nord-Pas-de-Calais)	Anthracomartus sp., Maiocercus orbicularis (155425) Aphantomartus areolatus (153494) Maiocercus orbicularis (155440)
Westphalian A	318.7 - 316.9	Germany	Cryptomartus meyeri (155773)
Westphalian A	318.7 - 316.9	Belgium	Anthracomartus denuiti (155639) Brachypyge carbonis (155637) Brachypyge orbicularis (155638) Trigonomartus dorlodoti (155635)
Westphalian A	318.7 - 316.9	United Kingdom (England)	Cleptomartus plautus, Cryptomartus hindi (127287) Cryptomartus hindi (34905) Maiocercus celticus (34944) Maiocercus celticus, Cryptomartus sp. (124367)
Westphalian A	318.7 - 316.9	Poland	Architarbus silesiacus (153496) Trigonotarbus sp., Anthracomartus voelkelianus, Anthracomartus granulatus (146849)
Westphalian A	318.7 - 316.9	United Kingdom (Staffordshire)	Anthracosiro woodwardi (155452)
Westphalian	318.7 - 307.4	USA (Rhode Island)	Anthracomartus woodruffi (112700)
Westphalian	318.7 - 307.4	France (Nord-Pas-de-Calais)	Kreischeria geinitzi (153493)
Westphalian B	316.9 - 314.6	United Kingdom (Staffordshire)	Anthracomartus hindi, Anthracomartus priesti (34908) Anthracomartus hindi, Anthracomartus priesti, Cleptomartus plautus (34910) Anthracomartus hindi, Anthracomartus priesti, Trigonotarbus johnsoni, Cleptomartus plautus (34907) Anthracomartus hindi, Trigonotarbus johnsoni, Cryptomartus hindi, Anthracomartus priesti (34935) Cryptomartus hindi, Cryptomartus priesti (155449) Eophrynus prestvicii (153486) Trigonotarbus johnsoni, Cleptomartus plautus, Cleptomartus planus, Cryptomartus priesti (34906)
Westphalian B	316.9 - 314.6	United Kingdom (England)	Anthracomartus hindi (125071) Anthracosiro sp., Anthracosiro woodwardi, Anthracosiro fritschii (127218) Anthracosiro woodwardi, Anthracosiro fritschii, Trigonotarbus johnsoni, Cleptomartus plautus, Cryptomartus priesti (125238) Anthracosiro woodwardi, Eophrynus varius (127286) Aphantomartus areolatus, Anthracosiro fritschi, Pseudokreischeria pococki (105060)
Westphalian B	316.9 - 314.6	Germany	Somaspidion hammapheron (155776)
Westphalian B	316.9 - 314.6	USA (West Virginia)	Areomartus ovatus (146859)
Westphalian B	316.9 - 314.6	France (Nord-Pas-de-Calais)	Aphantomartus pococki (153495) Kreischeria thevenini (154641)
Westphalian B	316.9 - 314.6	United Kingdom	Eophrynus carbonis (153491)
Westphalian B	316.9 - 314.6	Belgium	Anthracomartus volkelianus (155644) Brachypyge carbonis (155631)
Westphalian B	316.9 - 314.6	United Kingdom (Wales)	Eophrynus warei (172786)
Westphalian B	316.9 - 314.6	Czech Republic	Doubravatarbus krafti (226232)
Westphalian B	316.9 - 314.6	United Kingdom (Nottinghamshire)	Anthracomartus sp. (153665)
Westphalian B	316.9 - 314.6	France	Anthracosiro corsini (155645)
Westphalian B	316.9 - 314.6	United Kingdom (Durham)	Anthracosiro latipes (153489) Anthracosiro woodwardi (155451)
Moscovian	315.2 - 307.0	Czech Republic	Petrovicia proditoria (146850)
Moscovian	315.2 - 307.0	Czech Republic (Pilsen)	Hemiphrynus longipes, Hemiphrynus hofmanni, Promygale bohemica, Promygale rotundata, Promygale elegans, Arthrolycosa salticoides, Arthrolycosa carcinoides, Brachylycosa kustae, Oomartus nyranensis (128574)
Moscovian	315.2 - 307.0	Czech Republic (Central Bohemian)	Anthracomartus minor, Anthracomartus affinis, Anthracomartus krejcii, Anthracomartus socius (126152)
Westphalian C	314.6 - 309.8	Germany	Anthracophrynus tuberculatus (138364 172344) Eophrynus waechteri (172763) Termes buchi (138349)
Westphalian C	314.6 - 309.8	Czech Republic	Promygale janae (155304) Tynecotarbus tichaveki, Trigonotarbida indet. (152463)
Westphalian C	314.6 - 309.8	Ukraine	Trigonotarbida indet. (233238)
Westphalian C	314.6 - 309.8	Slovakia	Anthracomartus voelkelianus (153479)
Westphalian C	314.6 - 309.8	Germany (Saarland)	Termes hageni (122601)
Westphalian C	314.6 - 309.8	United Kingdom (England)	Curculioides prestvicii (122563)
Westphalian C	314.6 - 309.8	Belgium	Brachypyge carbonis (123824) Trigonomartus villeti (123823)
Westphalian C - Westphalian D	314.6 - 307.4	France (Nord-Pas-de-Calais)	Kreischeria villeti (151697)
Westphalian D	309.8 - 307.4	France	Anthracosiro elongatus (153153)
Westphalian D	309.8 - 307.4	USA (Indiana)	Eophrynidae indet. (226214)
Westphalian D	309.8 - 307.4	Germany	Kreischeria wiedei (35059)
Westphalian D	309.8 - 307.4	United Kingdom	Aphantomartus areolatus (141947) Eophrynus jugatus (162731) Kreischeria verrucosa (69648)
Westphalian D	309.8 - 307.4	USA (Oklahoma)	Anthracomartus sp. (226215) Aphantomartus sp. (152120)
Westphalian D	309.8 - 307.4	USA (Illinois)	Anthracomartus pustulatus, Trigonotarbus carbonarius (21981) Pleophrynus ensifer (133734) Trigonomartus pustulatus (146857) Trigonotarbus schucherti, Trigonomartus pustulatus (112630)
Stephanian	307.4 - 300.3	Italy (Sardinia)	Anthracomartidae indet. (153483)
Stephanian	307.4 - 300.3	Germany	Anthracomartus palatinus (138156)
Kasimovian	307.0 - 303.7	USA (New Mexico)	Pleophrynus hawesi (159023)
Kasimovian	307.0 - 303.7	USA (Kansas)	Anthracomartus sp. (122495)
Kasimovian - Gzhelian	307.0 - 298.9	Argentina (San Luis)	Gondwanarachne argentinensis (35057)
Stephanian B	301.4 - 300.9	France	Trigonotarbus arnoldi (153610)
Stephanian B	301.4 - 300.9	Czech Republic	Cryptomartus radvanicensis (155303)
Stephanian C	300.9 - 300.3	France	Aenigmatarbus rasteli (177540)
Stephanian C	300.9 - 300.3	Portugal	Aphantomartus pustulatus (156371)
Stephanian C	300.9 - 300.3	Germany	Cleptomartus ploetzensis (172340)
Asselian	298.9 - 293.52	Germany (Thuringia)	Brachylycosa manebachensis (172766) Eophrynus ilfeldicus, Eophrynus scharfi (155777)
Sakmarian - Artinskian	293.52 - 283.3	Germany	Permotarbus schuberti (153497)