PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Palaeodictyoptera

Insecta - Palaeodictyoptera

Taxonomy

Palaeodictyoptera was named by Goldenberg (1877).

It was assigned to Pterygogenea by Handlirsch (1906); to Palaeoptera by Martynov (1928), Carpenter (1992) and Brauckmann and Gröning (1996); to Palaeodictyopterida by Grimaldi and Engel (2005); to Insecta by Atkinson (2005); and to Palaeodictyopteroida by Bechly (2007).

Subtaxa

Achaeoptilus, Aenigmatopsis, Althansia, Anagesthes, Anthracosta, Asiodictya, Bathytaptus, Bojoptera, Breyerioidea, Brongniartites, Brongniartula, Commentryites (syn. Commentrya), Compsoneura (syn. Zeilleria), Delafondia, Dictyoneurina, Diexodus, Elmoboriidae, Eubleptidae, Eugereonoidea, Eukulojidae, Eumecoptera, Eurydictyella (syn. Eurydictya), Eurythmopteryx, Frankenholziina, Gegenemene, Haplophlebium, Heolidae, Heolus, Homoiopteridae (syn. Thesoneuridae, Roechlingiidae, Scepasmidae), Homothetidae, Idoptilus, Kansasia, Kansasiidae, Mammia, Mecynoptera, Mecynopteridae, Mecynostomites, Megaptilidae (syn. Anaxionidae, Lithoptilidae), Mongolianidae, Monsteropterum, Neopalaeodictyoptera, Neosecoptera, P. amissa, P. macrophlebia, Palaiotaptus, Palapteris, Paraboltonia, Paramecynostoma, Paramegaptilus, Platephemera, Propalingenia, Pseudomecynostoma, Psychroptilidae, Pteronidia, Pteronidiidae, Pyebrodia, Rochdalia, Saarlandia, Saarlandiidae, Sabitaptus, Schedoneura, Severinopsis, Spilaptera sutcliffei, Spilapterites, Spilapteroidea, Straeleniellidae, Synaptoneura, Titanophasma, Turnbullia

Synonyms

Neosecoptera was named by Riek (1976).

It was considered an invalid subgroup of Palaeodictyoptera by Carpenter (1992).

It was assigned to Megasecoptera by Riek (1976).
Dictyoneurida was named by Handlirsch (1906).

It was synonymized subjectively with Palaeodictyoptera by Carpenter (1992).

It was assigned to Insecta by Sinitshenkova (1981).
Protohemiptera was named by Handlirsch (1906). It is not extant.

It was synonymized subjectively with Palaeodictyoptera by Carpenter (1992).

It was assigned to Insecta by Handlirsch (1906) and McKeown (1937).
Heolidae was named by Handlirsch (1906). Its type is Heolus.

It was considered an invalid subgroup of Palaeodictyoptera by Carpenter (1992).

It was assigned to Palaeodictyoptera by Handlirsch (1906), Handlirsch (1906), Handlirsch (1919) and Handlirsch (1922).
Spilaptera sutcliffei was named by Bolton (1916). Its type specimen is MM L.8197, a wing, and it is an impression. Its type locality is Sparth Bottoms, Rochdale (MM coll), which is in a Westphalian A delta plain shale in the United Kingdom. It is the type species of Spilapterites.

It was recombined as Spilapterites sutcliffei by Handlirsch (1922); it was considered a nomen dubium by Kukalová (1969).
Termitidium amissum was named by Goldenberg (1877). It is an impression. Its type locality is Altenwald mine, which is in a Westphalian C terrestrial shale/bituminous coal in the Lower Saarbrucker layer Formation of Germany.

It was recombined as Dictyoneura amissa by Handlirsch (1906) and von Schlechtendal (1913); it was considered a nomen dubium by Handlirsch (1919), Handlirsch (1922), Waterlot (1934) and Guthörl (1934).
Dictyoneura macrophlebia was named by Handlirsch (1906). It is an impression. Its type locality is Altenwald mine, which is in a Westphalian C terrestrial shale/bituminous coal in the Lower Saarbrucker layer Formation of Germany.

It was considered a nomen nudum by Goldenberg (1885); it was considered a nomen dubium by Handlirsch (1919), Handlirsch (1922), Waterlot (1934) and Guthörl (1934).
Mecynopteridae was named by Handlirsch (1904).

It was synonymized subjectively with Stenodictyopteridae by Laurentiaux (1953); it was considered an invalid subgroup of Palaeodictyoptera by Carpenter (1992).

It was assigned to Palaeodictyoptera by Handlirsch (1906), Handlirsch (1919), Handlirsch (1922) and Pruvost (1930).
Pteronidiae was named by Bolton (1912). Its type is Pteronidia.

It was corrected as Pteronidiidae by Handlirsch (1919), Bolton (1921) and Handlirsch (1922); it was considered an invalid subgroup of Palaeodictyoptera by Carpenter (1992).

It was assigned to Palaeodictyoptera by Handlirsch (1919), Bolton (1921) and Handlirsch (1922).
Saarlandiidae was named by Guthörl (1930). It is not extant. Its type is Saarlandia.

It was considered an invalid subgroup of Palaeodictyoptera by Carpenter (1992).

It was assigned to Palaeodictyoptera by Guthörl (1930) and Guthörl (1934).
Kansasiidae was named by Demoulin (1954). Its type is Kansasia.

It was considered an invalid subgroup of Palaeodictyoptera by Carpenter (1992).

It was assigned to Archodonata by Demoulin (1954).
Spilapterites was named by Handlirsch (1922). Its type is Spilaptera sutcliffei.

It was considered a nomen dubium by Kukalová (1969).

It was assigned to Palaeodictyoptera by Handlirsch (1922).
Hemiodonata was named by Zalessky (1946).

It was synonymized subjectively with Palaeodictyoptera by Carpenter (1992).

Synonymy list

Year	Name and author
1877	Palaeodictyoptera Goldenberg p. 8
1877	Termitidium amissum Goldenberg p. 17 figs. Pl 1, fig 6
1904	Mecynopteridae Handlirsch p. 7
1906	Dictyoneurida Handlirsch
1906	Palaeodictyoptera Handlirsch p. 61
1906	Dictyoneura amissa Handlirsch p. 78 figs. Pl X fig 5
1906	Dictyoneura macrophlebia Handlirsch p. 78 figs. Pl 10, fig 6
1906	Mecynopteridae Handlirsch p. 82
1906	Heolidae Handlirsch p. 94
1906	Protohemiptera Handlirsch p. 387
1906	Heolidae Handlirsch p. 677
1913	Dictyoneura amissa von Schlechtendal p. 128 figs. Pl 10, fig 2
1913	Dictyoneura macrophlebica von Schlechtendal p. 129 figs. Pl 10, fig 1
1916	Spilaptera sutcliffei Bolton p. 53 figs. Pl 4, fig 1
1919	Mecynopteridae Handlirsch p. 524
1919	Pteronidiidae Handlirsch p. 524
1919	Heolidae Handlirsch p. 527
1921	Pteronidiidae Bolton
1921	Spilaptera sutcliffei Bolton p. 54 figs. 14; Pl III fig 3
1922	Mecynopteridae Handlirsch p. 35
1922	Pteronidiidae Handlirsch p. 35
1922	Heolidae Handlirsch p. 42
1922	Spilapterites Handlirsch p. 57
1922	Spilapterites sutcliffei Handlirsch p. 57
1928	Palaeodictyoptera Martynov p. 4
1930	Saarlandiidae Guthörl p. 154
1930	Mecynopteridae Pruvost p. 147
1934	Saarlandiidae Guthörl p. 85
1937	Protohemiptera McKeown p. 31
1946	Hemiodonata Zalessky p. 63
1954	Kansasiidae Demoulin p. 334
1976	Neosecoptera Riek p. 230
1981	Dictyoneurida Sinitshenkova p. 121
1992	Palaeodictyoptera Carpenter p. 26
1996	Palaeodictyoptera Brauckmann and Gröning
2005	Palaeodictyoptera Atkinson
2005	Palaeodictyoptera Grimaldi and Engel
2007	Palaeodictyoptera Bechly

Is something missing? Join the Paleobiology Database and enter the data

Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Ecdysozoa
—	Panarthropoda
phylum	Arthropoda	Latreille 1829
superclass	Hexapoda	Latreille 1825

Rank	Name	Author
class	Insecta
—	Dicondylia
—	Paranotalia
subclass	Pterygota	()
—	Neopterygota	Crampton 1924
superorder	Palaeodictyopterida	(Bechly 1996)
order	Palaeodictyoptera
order	Palaeodictyoptera

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Or. †Palaeodictyoptera Goldenberg 1877

show all | hide all

G. †Achaeoptilus Scudder 1881

†Achaeoptilus ingens Scudder 1881

G. †Aenigmatopsis Handlirsch 1937

†Aenigmatopsis regularis Bolton 1921

G. †Althansia Guthörl 1934

†Althansia sahneri Guthörl 1934

G. †Anagesthes Handlirsch 1906

†Anagesthes affinis Goldenberg 1854

G. †Anthracosta Pruvost 1930

†Anthracosta duboisi Pruvost 1930

G. †Asiodictya Rohdendorf 1961

†Asiodictya rossica Rohdendorf 1961

G. †Bathytaptus Handlirsch 1906

†Bathytaptus falcipennis Handlirsch 1906

G. †Bojoptera Kukalová 1958

†Bojoptera colorata Kukalová 1958

Superfm. †Breyerioidea Handlirsch 1906

Fm. †Breyeriidae Handlirsch 1906

G. †Aviobreyeria Prokop et al. 2013

†Aviobreyeria gracilis Prokop et al. 2013

G. †Breyeria de Borre 1875

†Breyeria barborae Kukalová 1959

†Breyeria bistrata Brauckmann 1995

†Breyeria borinensis de Borre 1875

†Breyeria boulei Meunier 1910

†Breyeria brauckmanni Pinto 1992

†Breyeria delruei Laurentiaux 1949

†Breyeria janseni Zessin 2006

†Breyeria lachlani Brongniart 1894

†Breyeria limburgica Laurentiaux 1950

†Breyeria rappi Carpenter 1967

†Breyeria sharovi Sinitshenkova 1979

†Breyeria solida Brauckmann and Herd 2000

†Breyeria triramosa Sinitshenkova 1979

†Breyeria vrankeni Laurentiaux-Vieira and Laurentiaux 1964

Invalid names: Borrea Brongniart 1894 [synonym], Breyeriopsis Laurentiaux 1949 [synonym], Pseudoborrea Handlirsch 1919 [synonym]

G. †Hasala Brauckmann 1995

†Hasala inferiorsaxonica Brauckmann 1995

G. †Jugobreyeria Brauckmann et al. 1985

†Jugobreyeria sippelorum Brauckmann et al. 1985

G. †Megaptiloides Handlirsch 1906

†Megaptiloides brodiei Brongniart 1894

G. †Stobbsia Handlirsch 1908

†Stobbsia woodwardiana Handlirsch 1908

G. †Vernooijia Prokop et al. 2018

†Vernooijia harlemensis Brauckmann and Gröning 1996

†Vernooijia sassoonae Prokop et al. 2018

Fm. †Cryptoveniidae Bolton 1912

G. †Cryptovenia Bolton 1912

†Cryptovenia moyseyi Bolton 1912

G. †Brongniartites Handlirsch 1919

†Brongniartites radians Handlirsch 1919

G. †Brongniartula Handlirsch 1919

†Brongniartula rastrata Handlirsch 1919

G. †Commentryites Handlirsch 1922

†Commentryites reticulata Handlirsch 1919

Invalid names: Commentrya Handlirsch 1919 [replaced]

G. †Compsoneura Brongniart 1894

†Compsoneura formosa Brongniart 1894

†Compsoneura fusca Brongniart 1885

Invalid names: Zeilleria Brongniart 1885 [replaced]

G. †Delafondia Langiaux and Parriat 1975

†Delafondia granulata Langiaux and Parriat 1975

Subor. †Dictyoneurina Handlirsch 1906

Superfm. †Calvertielloidea Martynov 1932

Fm. †Calvertiellidae Martynov 1931

G. †Calvertiella Tillyard 1925

†Calvertiella permiana Tillyard 1925

G. †Carrizopteryx Kukalová-Peck 1976

†Carrizopteryx arroyo Kukalová-Peck 1976

†Carrizopteryx lucasi Sinitshenkova 2004

G. †Lodetiella Béthoux et al. 2007

†Lodetiella magnifica Béthoux et al. 2007

G. †Moravia Kukalová 1964

†Moravia convergens Kukalová 1964

†Moravia grandis Carpenter 1979

G. †Moraviptera Kukalová 1955

†Moraviptera reticulata Kukalová 1955

G. †Sharovia Sinitshenkova 1977

†Sharovia sojanica Sinitshenkova 1977

†Sharovia tshekardica Ivanov 2000

G. †Xiaheyanella Fu et al. 2015

†Xiaheyanella orta Fu et al. 2015

Invalid names: Mongolodictyidae Sinitshenkova 1992 [synonym]

Superfm. †Dictyoneuroidea Handlirsch 1906

Fm. †Dictyoneuridae Handlirsch 1906

G. †Arltia Guthörl 1934

†Arltia carnica Brauckmann and Hahn 1983

†Arltia eurycubitalis Guthörl 1940

†Arltia mioanali Guthörl 1940

†Arltia obsoleta Goldenberg 1877

†Arltia schmitzi Goldenberg 1877

†Arltia willerti Guthörl 1930

G. †Cleffia Guthörl 1931

†Cleffia palatina Guthörl 1940

†Cleffia sarana Guthörl 1931

Invalid names: Pseudocleffia Guthörl 1940 [synonym]

G. †Dictyoneura Goldenberg 1854

†Dictyoneura higginsii Handlirsch 1906

†Dictyoneura kemperi Brauckmann and Koch 1983

†Dictyoneura libelluloides Goldenberg 1854

Invalid names: Dictyophlebia protogaea Goldenberg 1852 [replaced]

†Dictyoneura nigra Kliver 1883

†Dictyoneura rugosa Handlirsch 1906

Invalid names: Dictyophlebia Goldenberg 1852 [nomen oblitum]

G. †Dictyoneurula Handlirsch 1906

†Dictyoneurula gracilis Kliver 1886

G. †Goldenbergia Scudder 1885

†Goldenbergia elongata Goldenberg 1877

†Goldenbergia formosa Goldenberg 1854

Invalid names: Termes deeheni Goldenberg 1873 [synonym]

†Goldenbergia glomerata Halle 1933

G. †Hongius Özdikmen 2008

†Hongius qiligouensis Hong 1985

Invalid names: Palaeoneura Hong 1985 [replaced]

G. †Kallenbergia Guthörl 1930

†Kallenbergia handlirschi Guthörl 1930

G. †Macrodictya Guthörl 1940

†Macrodictya stenomediali Guthörl 1940

G. †Microdictya Brongniart 1894

†Microdictya hamyi Brongniart 1894

†Microdictya handlirschi Bolton 1925

†Microdictya lacroixi Meunier 1910

†Microdictya latipennis Handlirsch 1919

†Microdictya vaillanti Brongniart 1894

Invalid names: Stenodictya gaudryi Brongniart 1894 [synonym]

†Microdictya villeneuvei Meunier 1908

Invalid names: Heeria Brongniart 1894 [replaced]

G. †Polioptenus Scudder 1885

†Polioptenus bertrandi Waterlot 1934

†Polioptenus chanderisi Waterlot 1934

†Polioptenus decheni Goldenberg 1854

†Polioptenus elegans Goldenberg 1877

†Polioptenus mancomediali Guthörl 1940

Invalid names: Acanthodictyon Handlirsch 1906 [synonym]

G. †Rotundopteris Guthörl 1940

†Rotundopteris multimediali Guthörl 1940

G. †Saarodictyum Guthörl 1939

†Saarodictyum abnormis Guthörl 1939

G. †Sagenoptera Handlirsch 1906

G. †Schmidtopteron Brauckmann and Hahn 1978

†Schmidtopteron adictyon Brauckmann and Hahn 1978

G. †Siberiodictya Sharov and Sinitshenkova 1977

†Siberiodictya gigas Sharov and Sinitshenkova 1977

G. †Stenodictya Brongniart 1894

†Stenodictya agnita Meunier 1908

Invalid names: Stenodictya fayoli Meunier 1909 [synonym]

†Stenodictya arnaudi Brongniart 1894

†Stenodictya fritschi Brongniart 1894

†Stenodictya grandissima Meunier 1911

†Stenodictya klebsi Meunier 1908

†Stenodictya laurentiauxi Kukalová 1970

†Stenodictya lobata Brongniart 1890

†Stenodictya lusitanica Correia et al. 2014

†Stenodictya minima Brongniart 1894

†Stenodictya oustaleti Brongniart 1894

†Stenodictya parisiana Kukalová 1970

†Stenodictya perrieri Brongniart 1894

†Stenodictya pygmaea Meunier 1911

†Stenodictya spinosa Brongniart 1890

†Stenodictya thevenini Meunier 1908

†Stenodictya vasseuri Meunier 1914

Invalid names: Scudderia Brongniart 1890 [replaced]

G. †Stenodictyoneura Leriche 1911

†Stenodictyoneura belgica Leriche 1911

G. †Stilbocrocis Handlirsch 1906

†Stilbocrocis heeri Goldenberg 1854

†Stilbocrocis lanceolata Guthörl 1940

†Stilbocrocis pulchra Guthörl 1940

Invalid names: Longivenapteris Guthörl 1940 [synonym]

Invalid names: Dictyoneuridae humboldtiana Goldenberg 1854 [nomen dubium], Dictyoneuridae kliveri Handlirsch 1906 [nomen dubium], Eugereon heeri Handlirsch 1906 [nomen dubium], Stenodictyidae Laurentiaux 1952 [synonym], Stenodictyopteridae Brongniart [replaced]

G. †Titanodictya Handlirsch 1906

†Titanodictya jucunda Scudder 1885

†Titanodictya rowlandi Sinitshenkova 2004

G. †Diexodus Handlirsch 1911

†Diexodus debilis Handlirsch 1911

Fm. †Elmoboriidae Carpenter 1976

G. †Elmoboria Carpenter 1976

†Elmoboria piperi Carpenter 1976

G. †Oboria Kukalová 1960

†Oboria longa Kukalová 1960

Fm. †Eubleptidae Handlirsch 1906

G. †Eubleptus Handlirsch 1906

†Eubleptus danielsi Handlirsch 1906

Invalid names: Athymodictya parva Handlirsch 1911 [synonym]

†Eubleptus maculosus Carpenter 1992

Invalid names: Athymodictya Handlirsch 1911 [synonym]

Superfm. †Eugereonoidea Sinitshenkova 2002

Fm. †Archaemegaptilidae Handlirsch 1919

G. †Archaemegaptilus Meunier 1908

†Archaemegaptilus blakelyi Beckemeyer and Engel 2018

†Archaemegaptilus kiefferi Meunier 1908

†Archaemegaptilus schloesseri Brauckmann et al. 2003

G. †Osnabrugapteron Rosová et al. 2022

†Osnabrugapteron rasnitsyni Rosová et al. 2022

Fm. †Eugereonidae Handlirsch 1906

G. †Dictyoptilus Brongniart 1894

†Dictyoptilus peromapteroides Meunier 1908

†Dictyoptilus renaulti Brongniart 1894

†Dictyoptilus sepultus Meunier 1910

Invalid names: Cockerellia Meunier 1908 [synonym], Cockerelliella Meunier 1909 [synonym]

G. †Eugereon Dohrn 1866

†Eugereon boeckingi Dohrn 1866

G. †Peromaptera Brongniart 1894

†Peromaptera filholi Brongniart 1894

G. †Sandiella Carpenter 1970

†Sandiella herbigi Brauckmann et al. 2023

†Sandiella readi Carpenter 1970

G. †Valdeania Teixeira 1941

†Valdeania medeirosi Teixeira 1941

Invalid names: Cockerelliellidae Handlirsch 1919 [synonym], Dictyoptilidae Lameere 1917 [synonym], Peromapteridae Handlirsch 1906 [synonym]

Fm. †Graphiptilidae Handlirsch 1906

G. †Graphiptilus Brongniart 1894

†Graphiptilus heeri Brongniart 1894

Invalid names: Graphiptilus williamsoni Brongniart 1894 [synonym]

Invalid names: Graphiptiloides Handlirsch 1906 [synonym]

G. †Patteiskya Laurentiaux 1958

†Patteiskya bouckaerti Laurentiaux 1958

†Patteiskya volmensis Brauckmann 1984

G. †Rhabdoptilus Brongniart 1894

†Rhabdoptilus edwardsi Brongniart 1894

Invalid names: Rhabdoptilidae Handlirsch 1919 [synonym]

Fm. †Jongmansiidae Laurentiaux 1950

G. †Jongmansia Laurentiaux 1950

†Jongmansia tuberculata Bolton 1921

G. †Katosaxoniapteron Prokop et al. 2023

†Katosaxoniapteron brauneri Prokop et al. 2023

Fm. †Lithomanteidae Handlirsch 1906

G. †Archaeophasma Matthew 1910

†Archaeophasma grandis Matthew 1910

G. †Catadyesthus Handlirsch 1906

†Catadyesthus priscus Andrée 1864

G. †Dyscritus Scudder 1868

†Dyscritus vetustus Scudder 1880

G. †Lithomantis Woodward 1876

†Lithomantis bohemica Novak 1880

†Lithomantis carbonarius Woodward 1876

†Lithomantis meyeri Brauckmann et al. 2009

†Lithomantis varius Brauckmann et al. 1985

†Lithomantis woodwardi Brongniart 1890

Invalid names: Hadroneuria Handlirsch 1906 [synonym], Lithosialis Scudder 1881 [synonym]

G. †Lusiella Laurentiaux and Teixeira 1958

†Lusiella fariai Laurentiaux and Teixeira 1958

†Lusiella greberi Laurentiaux 1949

Invalid names: Macroptera Laurentiaux 1949 [replaced]

Invalid names: Lusiellidae Laurentiaux and Teixeira 1958 [synonym], Macropteridae Laurentiaux 1949 [synonym]

Fm. †Lycocercidae Handlirsch 1906

G. †Apopappus Handlirsch 1906

†Apopappus guernei Brongniart 1894

G. †Lycocercus Handlirsch 1906

†Lycocercus goldenbergi Brongniart 1885

Invalid names: Lycocercus brongniarti Handlirsch 1906 [synonym]

†Lycocercus pictus Handlirsch 1919

Invalid names: Homoioptera brongniarti Meunier 1911 [replaced]

G. †Lycodemas Carpenter and Richardson 1971

†Lycodemas adolescens Carpenter and Richardson 1971

G. †Lycodus Carpenter 1992

†Lycodus garretti Carpenter 1992

G. †Madera Carpenter 1970

†Madera mamayi Carpenter 1970

G. †Notorachis Carpenter and Richardson 1971

†Notorachis wolfforum Carpenter and Richardson 1971

Invalid names: Apopappidae Lameere 1917 [synonym], Patteiskyidae Laurentiaux 1958 [synonym]

Fm. †Namuroningxiidae Prokop and Ren 2007

G. †Namuroningxia Prokop and Ren 2007

†Namuroningxia elegans Prokop and Ren 2007

Fm. †Polycreagridae Handlirsch 1906

G. †Polycreagra Handlirsch 1906

†Polycreagra elegans Handlirsch 1906

Fm. †Protagriidae Handlirsch 1906

G. †Protagrion Brongniart 1894

†Protagrion audouini Brongniart 1894

Fm. †Synarmogidae Handlirsch 1910

G. †Synarmoge Handlirsch 1910

†Synarmoge ferrarii Handlirsch 1910

Invalid names: Climacoptera antiqua Laurentiaux 1949 [synonym]

Invalid names: Climacoptera Laurentiaux 1949 [synonym]

Fm. †Tchirkovaeidae Sinitshenkova 1979

G. †Paimbia Sinitshenkova 1979

†Paimbia fenestrata Sinitshenkova 1979

†Paimbia ultima Sinitshenkova 1981

G. †Tchirkovaea Zalessky 1931

†Tchirkovaea guttata Zalessky 1931

G. †Eumecoptera Handlirsch 1906

†Eumecoptera laxa Goldenberg 1877

G. †Eurydictyella Carpenter 1986

†Eurydictyella richteri Guthörl 1934

Invalid names: Eurydictya Guthörl 1934 [replaced]

G. †Eurythmopteryx Handlirsch 1906

†Eurythmopteryx antiqua Handlirsch 1906

Subor. †Frankenholziina Guthörl 1962

Fm. †Hanidae Kukalová-Peck 1975

G. †Forcynthia Sinitshenkova 2004

†Forcynthia cynthiae Sinitshenkova 2004

G. †Hana Kukalová-Peck 1975

†Hana filia Kukalová-Peck 1975

†Hana lineata Kukalová-Peck 1975

G. †Gegenemene Handlirsch 1906

†Gegenemene sinuosa Kliver 1883

G. †Haplophlebium Scudder 1867

†Haplophlebium barnesii Scudder 1867

Invalid names: Dictyoneura haplophlebia Goldenberg 1877 [objective synonym]

G. †Heolus Handlirsch 1906

†Heolus providentiae Handlirsch 1906

Fm. †Homoiopteridae Handlirsch 1906

G. †Adolarryia Kukalová-Peck and Richardson 1983

†Adolarryia bairdi Kukalová-Peck and Richardson 1983

G. †Ametretus Handlirsch 1911

†Ametretus laevis Handlirsch 1911

G. †Amousus Handlirsch 1911

†Amousus mazonus Handlirsch 1911

G. †Anglopterum Prokop et al. 2006

†Anglopterum magnificum Prokop et al. 2006

G. †Boltonia Pruvost 1919

†Boltonia robusta Pruvost 1919

Invalid names: Boltopruvostia Strand 1929 [objective synonym]

G. †Homoioptera Brongniart 1894

†Homoioptera gigantea Agnus 1902

Invalid names: Archaeoptilus gaullei Meunier 1910 [objective synonym]

†Homoioptera kortumi Brauckmann and Herd 2002

†Homoioptera latipenne Handlirsch 1904

†Homoioptera vorhallensis Brauckmann and Koch 1982

†Homoioptera woodwardi Brongniart 1890

Invalid names: Anthracentomon Handlirsch 1904 [synonym], Homoeophlebia Handlirsch 1906 [synonym]

G. †Larryia Kukalová-Peck and Richardson 1983

†Larryia osterbergi Kukalová-Peck and Richardson 1983

G. †Mazonopterum Kukalová-Peck and Richardson 1983

†Mazonopterum cooperi Prokop et al. 2018

†Mazonopterum wolfforum Kukalová-Peck and Richardson 1983

G. †Mazothairos Kukalová-Peck and Richardson 1983

†Mazothairos enormis Kukalová-Peck and Richardson 1983

G. †Ostrava Kukalová 1960

†Ostrava nigra Kukalová 1960

G. †Paraostrava Prokop and Nel 2004

†Paraostrava stanislavi Prokop and Nel 2004

G. †Parathesoneura Sinitshenkova 1977

†Parathesoneura anfractuosa Sinitshenkova 1977

†Parathesoneura carpenteri Sinitshenkova 1977

G. †Pharciphyzelus Beckemeyer and Engel 2011

†Pharciphyzelus lacefieldi Beckemeyer and Engel 2011

G. †Scepasma Handlirsch 1911

†Scepasma gigas Handlirsch 1911

Invalid names: Roechlingia hitleri Guthörl 1934 [synonym]

†Scepasma mediomatricorum Brauckmann and Becker 1992

Invalid names: Roechlingia Guthörl 1934 [synonym]

G. †Thesoneura Carpenter 1943

†Thesoneura americana Carpenter 1943

G. †Turneropterum Kukalová-Peck and Richardson 1983

†Turneropterum turneri Kukalová-Peck and Richardson 1983

Invalid names: Roechlingiidae Guthörl 1934 [synonym], Scepasmidae Haupt 1949 [synonym], Thesoneuridae Carpenter 1943 [synonym]

Fm. †Homothetidae Scudder 1885

G. †Homothetus Scudder 1867

†Homothetus fossilis Scudder 1867

G. †Idoptilus Wootton 1972

†Idoptilus onisciformis Wootton 1972

†Idoptilus peachii Woodward 1887

G. †Kansasia Tillyard 1937

†Kansasia pulchra Tillyard 1937

G. †Mammia Handlirsch 1906

†Mammia alutacea Handlirsch 1906

G. †Mecynoptera Handlirsch 1904

†Mecynoptera splendida Handlirsch 1904

G. †Mecynostomites Handlirsch 1919

†Mecynostomites brongniarti Handlirsch 1919

Fm. †Megaptilidae Handlirsch 1906

G. †Lithoptilus Lameere 1917

†Lithoptilus boulei Meunier 1908

Invalid names: Anaxion Handlirsch 1919 [objective synonym]

G. †Megaptilus Brongniart 1885

†Megaptilus blanchardi Brongniart 1885

Invalid names: Anaxionidae Handlirsch 1919 [synonym], Lithoptilidae Handlirsch 1922 [synonym]

G. †Monsteropterum Kukalová-Peck 1972

†Monsteropterum moravicum Kukalová-Peck 1972

G. †Palaiotaptus Handlirsch 1906

†Palaiotaptus mazonus Handlirsch 1906

G. †Palapteris Guthörl 1940

†Palapteris stenodictyus Guthörl 1940

G. †Paraboltonia Handlirsch 1922

†Paraboltonia macroptera Bolton 1916

G. †Paramecynostoma Handlirsch 1919

†Paramecynostoma dohrnianum Handlirsch 1919

G. †Paramegaptilus Handlirsch 1906

†Paramegaptilus scudderi Brongniart 1894

G. †Platephemera Scudder 1867

†Platephemera antiqua Scudder 1867

G. †Propalingenia Handlirsch 1906

†Propalingenia feistmanteli Frič 1880

G. †Pseudomecynostoma Handlirsch 1919

†Pseudomecynostoma dubium Handlirsch 1919

Fm. †Psychroptilidae Riek 1976

G. †Psychroptilus Riek 1976

†Psychroptilus burrettae Riek 1976

G. †Pteronidia Bolton 1912

†Pteronidia plicatula Bolton 1912

G. †Pyebrodia Brauckmann and Herd 2002

†Pyebrodia martinsnetoi Brauckmann and Herd 2002

Invalid names: Breyeria brevis Brauckmann and Herd 2002 [synonym]

G. †Rochdalia Woodward 1913

†Rochdalia parkeri Woodward 1913

G. †Saarlandia Guthörl 1930

†Saarlandia flexisubcostata Guthörl 1930

G. †Sabitaptus Pruvost 1930

†Sabitaptus lagagei Pruvost 1930

G. †Schedoneura Carpenter 1963

†Schedoneura amii Copeland 1957

G. †Severinopsis Kukalová 1958

†Severinopsis vetusta Kukalová 1958

Superfm. †Spilapteroidea Brongniart 1894

Fm. †Fouqueidae Handlirsch 1906

G. †Fouquea Brongniart 1894

†Fouquea lacroixi Brongniart 1894

Invalid names: Fouquea comstocki Lameere 1917 [synonym]

†Fouquea needhami Lameere 1917

†Fouquea sauvagei Brongniart 1894

†Fouquea superba Meunier 1909

Invalid names: Archaecompsoneura Meunier 1909 [synonym], Oustaletia Brongniart 1885 [nomen nudum]

G. †Neofouquea Carpenter 1967

†Neofouquea suzanneae Carpenter 1967

Fm. †Mecynostomatidae Kukalová 1969

G. †Mecynostomata Metcalf 1952

†Mecynostomata dohrni Brongniart 1894

Invalid names: Mecynostoma Brongniart 1894 [replaced]

Fm. †Spilapteridae Brongniart 1894

G. †Abaptilon Zalessky 1946

†Abaptilon sibiricum Zalessky 1946

G. †Baeoneura Sinitshenkova 1977

†Baeoneura desperata Sinitshenkova 2020

†Baeoneura obscura Sinitshenkova 1977

G. †Becquerelia Brongniart 1894

†Becquerelia elegans Brongniart 1894

†Becquerelia superba Brongniart 1894

†Becquerelia tincta Brongniart 1894

Invalid names: Pseudobecquerelia Handlirsch 1919 [synonym]

G. †Bizarrea Prokop et al. 2015

†Bizarrea obscura Prokop et al. 2015

G. †Delitzschala Brauckmann and Schneider 1996

†Delitzschala bitterfeldensis Brauckmann and Schneider 1996

G. †Dunbaria Tillyard 1924

†Dunbaria borealis Sharov and Sinitshenkova 1977

†Dunbaria fasciipennis Tillyard 1924

†Dunbaria mira Zalessky 1946

Invalid names: Doropteron Zalessky 1946 [synonym]

G. †Epitethe Handlirsch 1906

†Epitethe meunieri Brongniart 1894

G. †Epithele Handlirsch 1906

G. †Homaloneura Brongniart 1885

†Homaloneura berenice Brauckmann and Gröning 1998

†Homaloneura bonnieri Brongniart 1894

†Homaloneura brauckmanni Sinitshenkova 2020

†Homaloneura bucklandi Brongniart 1894

†Homaloneura dabasinskasi Carpenter 1964

†Homaloneura elegans Brongniart 1885

†Homaloneura joannae Brongniart 1894

†Homaloneura kiliani Brauckmann et al. 2009

†Homaloneura lehmani Kukalová 1969

†Homaloneura ligeia Brauckmann 1986

†Homaloneura ornata Brongniart 1894

†Homaloneura punctata Brongniart 1894

†Homaloneura redbirdi Brauckmann et al. 1993

†Homaloneura ribeiroi Teixeira 1946

Invalid names: Homaloneurina Handlirsch 1906 [synonym], Homaloneurites Handlirsch 1906 [synonym], Sheltoweeptera Brauckmann et al. 1993 [synonym]

G. †Lamproptilia Brongniart 1885

†Lamproptilia grandeuryi Brongniart 1885

†Lamproptilia tenuitegminata Bolton 1911

G. †Mcluckiepteron Richardson 1956

†Mcluckiepteron luciae Richardson 1956

G. †Neuburgia Martynov 1930

†Neuburgia altaica Martynov 1930

G. †Palaeoptilus Brongniart 1894

†Palaeoptilus brullei Brongniart 1894

G. †Paradunbaria Sharov and Sinitshenkova 1977

†Paradunbaria pectinata Sharov and Sinitshenkova 1977

G. †Permiakovia Martynov 1940

†Permiakovia quinquefasciata Martynov 1940

G. †Saxonyptilus Rosová et al. 2022

†Saxonyptilus sowiaki Rosová et al. 2022

G. †Sinodunbaria Li et al. 2013

†Sinodunbaria jarmilae Li et al. 2013

G. †Spilaptera Brongniart 1885

†Spilaptera americana Carpenter and Richardson 1971

†Spilaptera libelluloides Brongniart 1894

†Spilaptera packardi Brongniart 1885

†Spilaptera peckae Rosová et al. 2022

†Spilaptera splendens Prokop et al. 2014

†Spilaptera tanaica Sharov and Sinitshenkova 1977

†Spilaptera venusta Brongniart 1894

G. †Spiloptilus Handlirsch 1906

†Spiloptilus ramondi Brongniart 1894

G. †Tectoptilus Kukalová 1969

†Tectoptilus grehanti Brongniart 1894

G. †Tytthospilaptera Liu et al. 2015

†Tytthospilaptera wangae Liu et al. 2015

G. †Vorkutoneura Sinitshenkova 1977

†Vorkutoneura variabilis Sinitshenkova 1977

Invalid names: Doropteridae Zalessky 1946 [synonym], Dunbariidae Handlirsch 1937 [synonym], Lamproptilia stirrupi Brongniart 1894 [nomen dubium], Lamproptilidae Handlirsch 1906 [synonym], Neuburgiidae Rohdendorf 1961 [synonym]

Fm. †Straeleniellidae Laurentiaux-Vieira and Laurentiaux 1986

G. †Straeleniella Laurentiaux-Vieira and Laurentiaux 1986

†Straeleniella namurensis Laurentiaux-Vieira and Laurentiaux 1986

G. †Synaptoneura Pruvost 1919

†Synaptoneura champyi Pruvost 1919

G. †Titanophasma Brongniart 1882

†Titanophasma fayoli Brongniart 1882

G. †Turnbullia Richardson 1956

†Turnbullia priscillae Richardson 1956

Invalid names: Dictyoneurida Handlirsch 1906 [synonym], Hemiodonata Zalessky 1946 [synonym], Heolidae Handlirsch 1906 [invalid subgroup], Kansasiidae Demoulin 1954 [invalid subgroup], Mecynopteridae Handlirsch 1904 [invalid subgroup], Mongolianidae Özdikmen 2008 [empty], Neopalaeodictyoptera [empty], Neosecoptera Riek 1976 [invalid subgroup], Palaeodictyoptera amissa Goldenberg 1877 [nomen dubium], Palaeodictyoptera macrophlebia Handlirsch 1906 [nomen dubium], Protohemiptera Handlirsch 1906 [synonym], Pteronidiidae Bolton 1912 [invalid subgroup], Saarlandiidae Guthörl 1930 [invalid subgroup], Spilaptera sutcliffei Bolton 1916 [nomen dubium], Spilapterites Handlirsch 1922 [nomen dubium]

Diagnosis

No diagnoses are available