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Heterodactyla

Insecta - Diptera

Taxonomy
Heterodactyla is an unranked clade. It is extant.

It was assigned to Brachycera by Shin et al. (2018).

Subtaxa
Asiloidea (syn. Bombylioidea), Bombyliidae, Eremoneura
Sister taxa
Eremoneura

Synonymy list
YearName and author
2018Heterodactyla Shin et al.

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—Ecdysozoa
—Panarthropoda
phylumArthropodaLatreille 1829
RankNameAuthor
superclassHexapodaLatreille 1825
classInsecta
orderDiptera
—NeodipteraMichelsen 1994
suborderBrachyceraZetterstedt 1842
—Heterodactyla
—Heterodactyla

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

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Superfm. Asiloidea Latreille 1802
+
Fm. Apsilocephalidae Nagatomi et al. 1991
+
G. Apsilocephala Kröber 1914
G. †Cascomixticus Poinar and Vega 2021
+
†Cascomixticus tubuliferous Poinar and Vega 2021
G. †Irwinimyia Zhang et al. 2017
+
†Irwinimyia spinosa Zhang et al. 2017
G. †Palaeoapsilocephala Hauser and Greenwalt 2022
+
†Palaeoapsilocephala kishenehnensis Hauser and Greenwalt 2022
Fm. Asilidae Latreille 1802 [robber fly]
+
G. †Araripogon Grimaldi 1990
+
†Araripogon axelrodi Grimaldi 1990
†Araripogon pulchrus Lamas et al. 2021
Subfm. Asilinae Latreille 1802
+
Tr. Apocleini Papavero 1973
+
G. Promachus Loew 1848
Tr. Asilini Latreille 1802
+
G. Asilus Linnaeus 1758
G. †Asilopsis Cockerell 1921
Tr. Machimini Lehr 1996
+
G. Machimus Loew 1849
G. Neolophonotus Engel 1925
+
†Neolophonotus klebsi Meunier 1908
Invalid names: Lophonotus Maquart 1828 [replaced]
G. †Permessus Piton and Théobald 1939
+
†Permessus asiloides Piton and Théobald 1939
Tr. Philonicini Lehr 1996
+
G. Philonicus Loew 1849
G. Proctacanthus Macquart 1838
+
†Proctacanthus fractus Cockerell 1921
G. †Pseudophrisson Dürrenfeldt 1968
+
†Pseudophrisson primus Dürrenfeldt 1968
G. Senoprosopis Macquart 1838
+
†Senoprosopis beckeri Lewis 1972
†Senoprosopis eureka Melander 1949
Subfm. Brachyrhopalinae Hardy 1926
+
G. Holopogon Loew 1847
+
†Holopogon archilestes Hull 1960
†Holopogon pilipes Loew 1850
G. Nicocles Jaennicke 1867
+
†Nicocles miocenicus Cockerell 1909
G. †Burmapogon Dikow and Grimaldi 2014
+
†Burmapogon bruckschi Dikow and Grimaldi 2014
Subfm. Dasypogoninae Macquart 1838
+
Tr. Blepharepiini Papavero 1973
+
G. Blepharepium Rondani 1848
G. Ceraturgus Wiedemann 1824
+
†Ceraturgus praecursor James 1939
Invalid names: Ceraturgopsis Johnson 1903 [synonym]
G. Cophura Osten-Sacken 1887
+
†Cophura antiquella Cockerell 1913
Tr. Dasypogonini Macquart 1838
+
G. Dasypogon Meigen 1803
G. Taracticus Loew 1872
G. Lestomyia Williston 1883
+
†Lestomyia miocenica James 1939
Tr. Megapodini Carrera 1949
+
Subtr. Senobasina Papavero 1975
G. †Palaeomolobra Hull 1962
+
†Palaeomolobra antiqua James 1939
Tr. Saropogonini Hardy 1926
+
G. Saropogon Loew 1847
G. †Stenocinclis Scudder 1878
+
†Stenocinclis anomala Scudder 1878
Invalid names: Damalini [empty]
Subfm. Dioctriinae Enderlein 1936
+
Tr. Dioctriini Enderlein 1936
+
G. Dioctria Meigen 1803
G. †Kishenehnoasilus Dikow 2019
+
†Kishenehnoasilus bhl Dikow 2019
Subfm. Laphriinae Macquart 1838
+
Tr. Atomosiini Lynch Arribalzaga 1882
+
G. †Protoloewinella Schumann 1984
G. Psilocurus Loew 1874
+
†Psilocurus tarsalis Statz 1940
Subfm. Leptogastrinae Schiner 1862 [grass fly]
+
G. †Cretagaster Dikow and Grimaldi 2014
+
†Cretagaster raritanensis Dikow and Grimaldi 2014
Tr. Leptogastrini Schiner 1862
+
G. Leptogaster Meigen 1803
G. Schildia Aldrich 1923
+
†Schildia martini Dikow and Bayless 2009
Subfm. Ommatiinae Hardy 1927
+
G. Ommatius Wiedemann 1821
+
†Ommatius fimbriatus Scarbrough and Poinar 1992
†Ommatius subtus Scarbrough and Poinar 1992
Subfm. Stenopogoninae Hull 1962
+
Tr. Enigmomorphini Hull 1962
+
G. Microstylum Macquart 1838
Fm. Evocoidae Yeates et al. 2006
+
G. †Burmapsilocephala Gaimari and Mostovski 2000
+
†Burmapsilocephala cockerelli Gaimari and Mostovski 2000
†Burmapsilocephala evocoa Grimaldi 2016
G. †Myanmarpsilocephala Zhang et al. 2017
+
†Myanmarpsilocephala grimaldii Zhang et al. 2017
G. †Kuhwahldyia Solórzano Kraemer and Cumming 2019
+
†Kuhwahldyia indefinita Solórzano Kraemer and Cumming 2019
G. †Kumaromyia Grimaldi and Hauser 2011
+
†Kumaromyia burmitica Grimaldi and Hauser 2011
G. †Myanmyia Grimaldi 2011
+
†Myanmyia asteiformia Grimaldi 2011
Fm. Mydidae Latreille 1806 [mydas fly]
+
G. †Cretomydas Wilkommen 2007
+
†Cretomydas santanensis Wilkommen 2007
G. Mydas Fabricius 1794
+
†Mydas miocenicus Cockerell 1913
Fm. †Protapioceridae Ren 1998
+
G. †Protapiocera Ren 1998
+
†Protapiocera convergens Zhang et al. 2007
G. †Pseudapiocera Zhang 2015
+
Fm. Scenopinidae Fallen 1817 [window fly]
+
Subfm. Proratinae Theodor 1983
+
G. †Burmaprorates Jouault and Nel 2020
+
†Burmaprorates alagracilis Jouault and Nel 2020
G. †Proratites Grimaldi and Cumming 1999
+
†Proratites simplex Grimaldi and Cumming 1999
Subfm. Scenopininae Fallen 1817
+
Tr. Metatrichini Winterton and Ware 2015
+
G. Brevitrichia Hardy 1994
G. †Eocenotrichia Garrouste et al. 2016
G. Metatrichia Coquillett 1900
Fm. Therevidae Newman 1834 [stiletto fly]
+
G. †Arctogephyra Hauser 2007
+
†Arctogephyra agilis Meunier 1908
G. †Cretothereva Carmo et al. 2021
+
†Cretothereva antiqua Carmo et al. 2021
G. †Kroeberiella Hauser 2007
+
†Kroeberiella pinguis Loew 1850
G. †Palaeopherocera Hauser and Irwin 2005
+
†Palaeopherocera scudderi Cockerell 1909
Subfm. Phycusinae Lyneborg 1976
+
G. †Glaesorthactia Hennig 1967
+
G. Rueppellia Wiedemann 1830
G. Psilocephala Zetterstedt 1838
+
†Psilocephala electrella Cockerell 1920
Invalid names: Paraclia Enderlein 1936 [synonym]
Subfm. Therevinae Newman 1834
+
G. †Ambradolon Metz and Irwin 2000
+
†Ambradolon grimaldii Metz and Irwin 2000
G. Nebritus Coquillett 1894
G. Thereva Latreille 1796
+
†Thereva carbonum von Heyden 1856
†Thereva marcelini Theobald 1937
Invalid names: Thereua bosniaskii Handlirsch 1907 [nomen nudum]
Subfm. Xestomyzinae Lyneborg 1976
+
G. †Peratrimera Hauser and Irwin 2005
+
†Peratrimera mexicana Hauser and Irwin 2005
Invalid names: Bombylioidea Latreille 1802 [synonym]
Fm. Bombyliidae Latreille 1802 [bee fly]
+
Subfm. Anthracinae Latreille 1804
+
Tr. Anthracini Latreille 1804
+
G. Anthrax Scopoli 1763
+
†Anthrax succini Greenwalt and Evenhuis 2015
G. Spogostylum Macquart 1840
Tr. Exoprosopini Becker 1913
+
G. Exoprosopa Macquart 1840
+
†Exoprosopa romani Theobald 1937
Tr. Villini Hull 1973
+
G. †Eoanomala Greenwalt and Evenhuis 2015
+
†Eoanomala melas Greenwalt and Evenhuis 2015
G. Hemipenthes Loew 1869
+
†Hemipenthes dominicana Evenhuis 2013
†Hemipenthes gabbroensis Handlirsch 1907
†Hemipenthes provincialis Handlirsch 1907
Invalid names: Hemipenthes tertiaria Handlirsch 1907 [nomen nudum], Hemipenthes tertiarius Handlirsch 1907 [nomen nudum]
G. †Verrallites Cockerell 1913
+
†Verrallites cladurus Cockerell 1913
G. Villa Lioy 1864
+
†Villa setosa Cockerell 1916
Invalid names: Protepacmus Cockerell 1916 [synonym]
Tr. Xeramoebini Hull 1973
+
G. Petrorossia Bezzi 1908
+
†Petrorossia gracilis Giebel 1862
Invalid names: Anthrax dentoni Lewis 1969 [nomen dubium]
Subfm. Bombyliinae Latreille 1802
+
Tr. Bombyliini Latreille 1802
+
G. Bombylius Linnaeus 1758
+
Subg. Bombylius (Bombylius) Linnaeus 1758
Invalid names: Bombylius fossilis Handlirsch 1907 [nomen nudum], Bombylius tertiarius Handlirsch 1907 [nomen nudum]
Tr. Conophorini Becker 1913
+
G. †Eoconophorina Maheu and Nel 2020
+
†Eoconophorina delfineae Maheu and Nel 2020
Tr. Dischistini Hull 1973
+
G. Dischistus Loew 1855
+
†Dischistus tertiarius Theobald 1937
Tr. Eclimini Hall 1969
+
G. †Alepidophora Cockerell 1909
+
†Alepidophora cockerelli Melander 1949
†Alepidophora maxima Lewis 1972 [bee fly]
†Alepidophora minor Melander 1949
†Alepidophora pealei Cockerell 1909
G. †Cretabombylia Ye et al. 2019
+
†Cretabombylia spinifera Ye et al. 2019
Subfm. Crocidiinae Hull 1973
+
G. Desmatomyia Williston 1895
+
†Desmatomyia scopulicornis Cockerell 1917
Invalid names: Acreotrichites Cockerell 1917 [synonym]
Subfm. Cythereinae Becker 1913
+
G. †Amictites Hennig 1966
+
†Amictites regiomontanus Hennig 1966
G. †Glaesamictus Hennig 1966
+
†Glaesamictus hafniensis Hennig 1966
G. †Palaeoamictus Meunier 1916
+
†Palaeoamictus spinosus Meunier 1916
G. †Praecytherea Théobald 1937
+
†Praecytherea sardii Théobald 1937
G. †Endymiomyia Grimaldi 2016
+
†Endymiomyia quadra Grimaldi 2016
Subfm. Lomatiinae Schiner 1868
+
G. †Alomatia Cockerell 1914
+
†Alomatia fusca Cockerell 1914
Tr. Lomatiini Schiner 1868
+
G. Comptosia Macquart 1840
+
†Comptosia miranda Cockerell 1909
†Comptosia pria Wedmann and Yeates 2008
Invalid names: Megacosmus Cockerell 1909 [synonym]
G. Ylasoia Speiser 1920
+
†Ylasoia secunda Cockerell 1911
G. †Paleolomatia Nel 2008
+
Subfm. Mariobezziinae Becker 1913
+
G. †Paracorsomyza Hennig 1966
+
G. †Nealimyia Grimaldi 2016
+
†Nealimyia evenhuisi Grimaldi 2016
G. †Nidergasia Jouault and Nel 2020
+
†Nidergasia neraudeaui Jouault and Nel 2020
G. †Pachysystropus Cockerell 1909
+
†Pachysystropus condemnatus Cockerell 1910
†Pachysystropus rohweri Cockerell 1909
G. †Palaeogeron Meunier 1914
+
†Palaeogeron vetustus Meunier 1914
G. †Paleocytherea Ngô-Muller and Nel 2020
+
†Paleocytherea pouilloni Ngô-Muller and Nel 2020
G. †Peculibombylia Ye et al. 2019
+
†Peculibombylia relictivena Ye et al. 2019
Subfm. Phthiriinae Becker 1913
+
G. †Elektrophthiria Nel 2006
+
Tr. Phthiriini Becker 1913
+
G. Phthiria Meigen 1803
+
†Phthiria fossa Lewis 1975
†Phthiria oligocenica Timon-David 1944
Tr. Poecilognathini Evenhuis 1990
+
G. Poecilognathus Jaennicke 1867
+
†Poecilognathus stigmalis Cockerell 1914
Invalid names: Geronites Cockerell 1914 [synonym]
G. Tmemophlebia Evenhuis 1986
+
†Tmemophlebia carolinae Evenhuis and Greenwalt 2022
G. †Pioneeria Grimaldi 2016
+
†Pioneeria bombylia Grimaldi 2016
G. †Procrocidium Grimaldi 2016
+
†Procrocidium minutum Grimaldi 2016
G. †Pseudorhagio Zhang et al. 2016
+
†Pseudorhagio zhangi Zhang et al. 2016
G. †Similipioneeria Ye et al. 2019
+
†Similipioneeria mirantenna Ye et al. 2019
G. †Tithonomyia Evenhuis 1984
+
†Tithonomyia atra Melander 1949
Subfm. Tomomyzinae Becker 1913
+
G. Amphicosmus Coquillett 1891
+
Subg. Amphicosmus (Amphicosmus) Coquillett 1891
+
G. Paracosmus Osten Sacken 1877
+
Subg. Paracosmus (Paracosmus) Osten Sacken 1877
+
Invalid names: Lithocosmus Cockerell 1909 [synonym], Protolomatia Cockerell 1914 [synonym], Protophthiria Cockerell 1914 [synonym]
Subfm. Toxophorinae Schiner 1868
+
Tr. Gerontini Hesse 1938
+
G. Geron Meigen 1820
+
†Geron figuratus Theobald 1937
†Geron oligocaenicus Timon-David 1944
†Geron platysoma Cockerell 1915
G. †Paradolichomyia Nel and De Ploeg 2004
+
†Paradolichomyia eocenica Nel and De Ploeg 2004
Tr. Systropodini Brauer 1880
+
G. Dolichomyia Wiedemann 1830
+
†Dolichomyia tertiaria Cockerell 1917
†Dolichomyia testea Melander 1949
G. †Melanderella Cockerell 1909
+
†Melanderella glossalis Cockerell 1909
G. Systropus Wiedemann 1820
+
Subg. Systropus (Systropus) Wiedemann 1820
†Systropus acourti Cockerell 1921
Subfm. Usiinae Becker 1913
+
Tr. Apolysini Evenhuis 1990
+
G. Apolysis Loew 1860
+
†Apolysis magister Melander 1946
Tr. Usiini Becker 1913
+
G. Usia Latreille 1802
+
Subg. Usia (Usia) Latreille 1802
Invalid names: Platypyginae Verrall 1909 [empty]
Unr. Eremoneura Grimaldi and Cumming 1999
+
Superfm. Apystomyoidea Melander 1950
+
Fm. Apystomyiidae Melander 1950
+
G. †Hilarimorphites Grimaldi and Cumming 1999
+
†Hilarimorphites burmanica Grimaldi and Cumming 2011
†Hilarimorphites cummingi Grimaldi 2016
†Hilarimorphites longimedia Grimaldi and Cumming 1999
†Hilarimorphites setosa Grimaldi and Cumming 1999
†Hilarimorphites superba Grimaldi and Cumming 1999
†Hilarimorphites yeatesi Grimaldi and Cumming 1999
Fm. †Chimeromyiidae Grimaldi and Cumming 2009
+
G. †Chimeromyia Grimaldi and Cumming 1999
+
†Chimeromyia acuta Grimaldi and Cumming 1999
†Chimeromyia alava Arillo and Grimaldi 2009
†Chimeromyia burmitica Grimaldi and Cumming 2009
†Chimeromyia intriguea Grimaldi and Cumming 1999
†Chimeromyia mediobscura Grimaldi and Cumming 2009
†Chimeromyia pilitibia Grimaldi and Cumming 2009
†Chimeromyia reducta Grimaldi and Cumming 1999
†Chimeromyia vulloi Jouault et al. 2020
G. †Chimeromyina Arillo and Grimaldi 2009
+
†Chimeromyina concilia Arillo and Grimaldi 2009
Subor. Cyclorrhapha Brauer 1863
+
Superfm. Lonchopteroidea Curtis 1839
+
G. †Alonchoptera Grimaldi 2018
+
†Alonchoptera lebanica Grimaldi 2018
Fm. Lonchopteridae Curtis 1839
+
G. Lonchoptera Meigen 1803
G. †Lonchopterites Grimaldi and Cumming 1999
+
†Lonchopterites burmensis Grimaldi 2018
†Lonchopterites prisca Grimaldi and Cumming 1999
G. †Lonchopteromorpha Grimaldi and Cumming 1999
+
†Lonchopteromorpha asetocella Grimaldi and Cumming 1999
Superfm. Phoroidea Newman 1834
+
G. †Lebambromyia Grimaldi and Cumming 1999
+
†Lebambromyia acrai Grimaldi and Cumming 1999
†Lebambromyia sacculifera Badano et al. 2021
Fm. Pipunculidae Walker 1834 [big-headed fly]
+
Subfm. Chalarinae Hardy 1965
+
G. †Protoverrallia Aczél 1948
G. Verrallia Mik 1899
Subfm. Nephrocerinae Carpenter and Hull 1939
+
G. Nephrocerus Zetterstedt 1838
G. †Priabona Archibald et al. 2014
Invalid names: Nephrocerini Carpenter and Hull 1939 [invalid subgroup]
Subfm. Pipunculinae Walker 1834
+
Tr. Pipunculini Walker 1834
Tr. Tomosvaryellini Hardy 1943
+
Invalid names: Eudorylini Rafael and De Meyer 1992 [synonym]
Subfm. Protonephrocerinae Aczél 1948
+
G. †Metanephrocerus Aczél 1948
G. Protonephrocerus Collin 1931
Superfm. Platypezoidea Fallen 1815
+
Fm. Ironomyiidae McAlpine and Martin 1966 [ironic fly]
+
Subfm. †Cretonomyiinae Li and Yeates 2018
G. †Proironia Grimaldi 2018
Subfm. †Sinolestinae Hong and Wang 1988
G. †Lithopetia Zhang 1987
+
†Lithopetia hirsuta Zhang 1987
G. †Opetiala Coram et al. 2000
+
†Opetiala shatalkini Coram et al. 2000
Fm. Opetiidae Kessel and Maggioncalda 1968 [flat-footed fly]
+
G. Opetia Meigen 1830
Fm. Phoridae Newman 1834 [scuttle fly]
+
G. Aenigmatias Meinert 1890
+
Invalid names: Protophorites Brues 1939 [synonym], Protoplatyphora Brues 1939 [synonym]
G. Dohrniphora Dahl 1898
G. †Electrophora Brues 1939
Unr. Euphorida Brown 2007
G. †Hypoceridites Brues 1939
G. †Jealia Brown 2009
G. †Limulomyia Brown 1999
G. †Megasella Paetel 1875
G. Metopina Macquart 1835
+
Invalid names: Pseudometopina Disney 1994 [synonym]
G. †Pararhoptrocera Prokop and Nel 2005
G. Paraspiniphora Malloch 1912
Subfm. †Prioriphorinae Mostovski 1996
G. †Prophora Grimaldi 2018
G. †Pseudaenigmatias Brown 2016
Subfm. Sciadocerinae Schmitz 1929
Invalid names: Eophora Handschin 1944 [nomen nudum]
Fm. Platypezidae Fallen 1815 [flat-footed fly]
+
G. Agathomyia Verral 1901
G. †Burmapeza Grimaldi 2018
G. Callomyia Meigen 1804
G. †Calvopeza Grimaldi 2018
G. †Canadopeza Grimaldi 2018
G. †Chandleromyia Grimaldi 2018
G. †Electrosania Grimaldi and Cumming 1999
G. †Lebanopeza Grimaldi 2018
G. Lindneromyia Kessel 1965
G. †Mecistoneuron Melander 1949
G. †Mesopetia Zhang 1987
G. Microsania Zetterstedt 1837
G. †Oppenheimiella Meunier 1893
Subfm. Platypezininae Kessel and Maggioncalda 1968
Invalid names: Clythiidae [synonym]
Unr. Schizophora Becher 1882
+
Unr. Acalyptratae Macquart 1835
+
Fm. †Christelenkidae Roháček 2023
Superfm. Diopsoidea Billberg 1820
Superfm. Nerioidea Westwood 1840
Superfm. Opomyzoidea
Superfm. Sphaeroceroidea Macquart 1835
Invalid names: Anthomyza rhenana Statz 1940 [nomen dubium], Ephydra oligocena Cockerell 1915 [nomen dubium], Ephydra sepulta Cockerell 1915 [nomen dubium], Leria insatiabilis Meunier 1910 [nomen dubium], Leria insaturabilis Meunier 1910 [nomen dubium], Leriella Meunier 1908 [nomen dubium], Leriella crassifemorata Meunier 1908 [nomen dubium]
Fm. Acartophthalmidae Czerny 1928
+
G. Acartophthalmus Czerny 1902
Fm. Agromyzidae Fallén 1823 [leaf-miner fly]
+
Subfm. Agromyzinae Fallén 1823
Subfm. Ophiomyiinae Xuan et al. 2023
Subfm. Phytomyzinae Fallén 1823
Fm. Asteiidae Rondani 1856
+
G. Asteia Meigen 1830
G. †Succinasteia Hennig 1969
Fm. Aulacigastridae Duda 1924
+
G. †Protaulacigaster Hennig 1965
Unr. Calyptratae Robineau-Desvoidy 1830
+
Fm. Anthomyiidae Robineau-Desvoidy 1830
Fm. Fanniidae Schnabl 1911
Superfm. Hippoboscoidea Samouelle 1819
Fm. Muscidae Latreille 1802 [house fly]
Superfm. Oestroidea Leach 1815
Fm. Scathophagidae Robineau-Desvoidy 1830 [dung fly]
Invalid names: Muscoidea Latreille 1802 [empty]
Fm. Carnidae Newman 1834 [bird fly]
+
G. Meoneura Rondani 1856
G. †Meoneurites Hennig 1965
Fm. Chloropidae Rondani 1856
+
Subfm. Chloropinae Rondani 1856
G. Hippelates Loew 1863
G. †Protoscinella Hennig 1965
Invalid names: Agromyza aberrans Meunier 1905 [nomen dubium]
Fm. Clusiidae Frey 1921 [druid fly]
+
Subfm. Clusiodinae Frey 1960
G. †Xenanthomyza Hennig 1967
Invalid names: Clusiinae Frey 1921 [empty]
G. †Cretaphormia McAlpine 1970
+
†Cretaphormia fowleri McAlpine 1970
Superfm. Ephydroidea Zetterstedt 1837
+
Fm. Camillidae Frey 1921
Fm. Campichoetidae Griffiths 1972
Fm. Cryptochetidae Brues and Melander 1932
Fm. Curtonotidae Duda 1934 [Quasimodo fly]
Fm. Drosophilidae Rondani 1856 [fruit fly]
Fm. Ephydridae Zetterstedt 1837 [shore fly]
Invalid names: Diastatidae Frey 1921 [empty]
G. †Hoffeinsia Stuke 2005
+
†Hoffeinsia baltica Stuke 2005
Fm. †Hoffeinsmyiidae Michelsen 2009
+
G. †Hoffeinsmyia Michelsen 2009
Superfm. Lauxanioidea Macquart 1835
+
Fm. Chamaemyiidae Hendel 1910 [aphid fly]
Fm. Lauxaniidae Macquart 1835
Fm. Milichiidae Schiner 1864
+
Subfm. Madizinae Czerny 1909
Subfm. Milichiinae Schiner 1864
G. Pholeomyia Bilimek 1867
Subfm. Phyllomyzinae Brake 2000
Fm. Neurochaetidae McAlpine 1978 [upside-down fly]
+
G. †Anthoclusia Hennig 1965
Fm. Odiniidae Hendel 1918
+
Subfm. Odiniinae
Fm. Opomyzidae Fallen 1810
+
G. Opomyza Fallen 1820
Fm. Periscelididae Oldenberg 1914
+
Subfm. Periscelidinae Oldenberg 1914
G. †Procyamops Hoffeins and Rung 2005
Subfm. Stenomicrinae Papp 1984
G. †Poliomyia Scudder 1878
+
†Poliomyia recta Scudder 1878
Superfm. Sciomyzoidea Fallen 1820
+
Fm. Conopidae Latreille 1802 [thick-headed fly]
Fm. Sciomyzidae Fallen 1820 [marsh fly]
Fm. Sepsidae Walker 1833
+
G. †Protorygma Hennig 1965
G. Sepsis Fallen 1810
G. Themira Robineau-Desvoidy 1830
Fm. Sphaeroceridae Macquart 1835 [lesser dung fly]
+
Subfm. Copromyzinae Stenhammar 1855
Subfm. Limosininae Frey 1921
Subfm. Sphaerocerinae Macquart 1835
Superfm. Tephritoidea Newman 1834
+
Fm. Lonchaeidae Rondani 1856
Fm. Pallopteridae Loew 1862
Fm. Piophilidae Macquart 1835 [cheese skipper]
Fm. Platystomatidae Schiner 1862 [signal fly]
Fm. Pyrgotidae Schiner 1868
Fm. Richardiidae Loew 1868
Fm. Tephritidae Newman 1834 [fruit fly]
Fm. Ulidiidae Macquart 1838
Invalid names: Herina splendida Meunier 1910 [nomen dubium]
Fm. †Yantaromyiidae Barták 2019
+
G. †Yantaromyia Barták 2019
Fm. Syrphidae Latreille 1802 [hover fly]
+
G. †Archisyrphus Hull 1960
+
†Archisyrphus opacus Hull 1960
Subfm. Eristalinae Newman 1834
+
Tr. Brachyopini Williston 1885
+
Invalid names: Chrysogasterini Shannon 1922 [synonym], Myioleptini Shannon 1922 [synonym]
Tr. Eristalini Newman 1834
Tr. Merodontini Edwards 1915
+
Invalid names: Eumerini Smirnov 1924 [synonym]
Tr. Milesiini Rondani 1845
Tr. Rhingiini Meigen 1822
Invalid names: Cheilosinae Williston 1885 [synonym], Milesiinae Rondani 1845 [synonym], Volucellini Newman 1834 [empty]
G. †Liomyodia Evenhuis 1994
+
†Liomyodia chalybea Hull 1945
Invalid names: Doliomyia Hull 1945 [replaced]
Subfm. Microdontinae Rondani 1845
+
Tr. Microdontini Rondani 1845
Subfm. Pipizinae Williston 1885
+
G. †Oligopipiza Nidergas et al. 2018
Tr. Pipizini Williston 1885
G. †Praeptilocephala Evenhuis 1994
+
Invalid names: Ptilocephala Hull 1945 [replaced]
G. †Prosyrphus Grimaldi 2018
+
†Prosyrphus thompsoni Grimaldi 2018
Subfm. Syrphinae Latreille 1802
+
Tr. Bacchini Williston 1885
G. †Eoxanthandrus Ngô-Muller and Nel 2020
Tr. Syrphini Latreille 1802
Invalid names: Chrysotoxinae Newman 1834 [synonym]
Invalid names: Syrphus bremii Heer 1864 [nomen dubium], Syrphus curvipetiolatus Meunier 1904 [nomen dubium], Syrphus schellenbergi Heer 1864 [nomen dubium]
Superfm. Syrphoidea Latreille 1802
+
G. †Aschizomyia Grimaldi 2018
+
†Aschizomyia burmensis Grimaldi 2018
Superfm. Empidoidea Latreille 1804
+
Fm. Atelestidae Hennig 1970
+
Subfm. Atelestinae Hennig 1970
+
G. Alavesia Waters and Arillo 1999
+
Invalid names: Neoalavesia Poinar and Vega 2020 [synonym]
G. †Atelestites Grimaldi and Cumming 1999
G. †Dianafranksia Coram et al. 2000
G. †Kurnubempis Kaddumi 2007
G. †Meghyperites Sinclair and Stark 2024
G. †Promeghyperus Zhang et al. 2024
G. †Trichinites Hennig 1970
Subfm. Nemedininae Sinclair and Cumming 2006
+
G. †Cretodromia Grimaldi and Cumming 1999
G. Nemedina Chandler 1981
G. †Nemedromia Grimaldi and Cumming 1999
G. †Neoturonius Grimaldi and Cumming 1999
G. †Phaetempis Grimaldi and Cumming 1999
G. †Prolatomyia Grimaldi and Cumming 1999
Fm. Dolichopodidae Latreille 1809 [long-legged fly]
+
Subfm. Achalcinae Grootaert and Meuffels 1997
+
G. Achalcus Loew 1857
G. Anepsiomyia Bezzi 1902
+
†Anepsiomyia planipedia Meunier 1907
Invalid names: Anepsius Loew 1857 [replaced]
G. †Archichrysotus Negrobov 1978
+
†Archichrysotus hennigi Negrobov 1978
†Archichrysotus incompletus Grimaldi and Cumming 1999
†Archichrysotus manitobus Grimaldi and Cumming 1999
†Archichrysotus minor Negrobov 1978
G. †Avenaphora Grimaldi and Cumming 1999
+
†Avenaphora gallica Nel et al. 2016
†Avenaphora hispida Grimaldi and Cumming 1999
G. Condylostylus Bigot 1859
G. †Cretomicrophorus Negrobov 1978
+
†Cretomicrophorus novemundus Grimaldi and Cumming 1999
†Cretomicrophorus piolencensis Nel et al. 2016
Subfm. Diaphorinae Schiner 1864
+
G. Argyra Macquart 1834
+
Invalid names: Porphyrops Meigen 1824 [replaced]
G. Chrysotus Meigen 1824
+
Invalid names: Lyroneurus Loew 1857 [synonym]
G. Diaphorus Meigen 1824
G. †Palaeoargyra Meunier 1895
G. †Prochrysotus Meunier 1907
Subfm. Dolichopodinae Latreille 1809
+
G. Dolichopus Latreille 1796
+
Invalid names: Hygroceleuthus Loew 1857 [synonym]
G. Gymnopternus Loew 1857
G. Hercostomus Loew 1857
G. †Leptodolichopodites Hong 2002
G. †Prohercostomus Grichanov 1997
+
Invalid names: Sympycnites Grimaldi and Cumming 1999 [synonym]
G. †Wangia Hong 2002
Invalid names: Arpactodolichopodites Hong 2002 [nomen nudum], Convexivertex Hong 2002 [nomen nudum], Eoeuryopterites Hong 2002 [nomen nudum], Orbicapitis Hong 2002 [nomen nudum]
G. †Electrochoreutes Badano et al. 2023
+
†Electrochoreutes electroechinus Badano et al. 2023
†Electrochoreutes falculigerus Badano et al. 2023
†Electrochoreutes furcillatus Badano et al. 2023
†Electrochoreutes hamatus Badano et al. 2023
†Electrochoreutes pankowskii Badano et al. 2023
†Electrochoreutes planitibia Badano et al. 2023
†Electrochoreutes trisetigerus Badano et al. 2023
G. †Gujaratmyia Bickel 2022
+
†Gujaratmyia rotunda Bickel 2022
Subfm. Hydrophorinae Lioy 1864
+
G. Thinophilus Wahlberg 1844
Invalid names: Thinophilinae Aldrich 1905 [synonym]
Subfm. Medeterinae Lioy 1864
+
G. Medetera Fischer von Waldheim 1819
G. †Medeterites Grichanov 2010
G. †Palaeosystenus Grichanov et al. 2014
G. †Plesiomedetera Grichanov 2023
G. †Salishomyia Bickel 2019
G. †Systenites Grichanov et al. 2014
G. Systenus Loew 1857
Tr. Thrypticini Negrobov 1986
G. Thrypticus Gerstacker 1864
G. Medeterus Fischer von Waldheim 1819
Subfm. Microphorinae Collin 1960
+
G. †Curvus Kaddumi 2005
G. †Meghyperiella Meunier 1908
G. Microphor Macquart 1827
G. Schistostoma Becker 1902
Invalid names: Microphorini Collin 1960 [empty]
G. †Microphorites Hennig 1971
+
†Microphorites deploegi Nel et al. 2004
†Microphorites erikai Bramuzzo and Nel 2017
†Microphorites extinctus Hennig 1971
†Microphorites magaliae Perrichot and Engel 2014
†Microphorites moravicus Tkoč et al. 2016
†Microphorites oculeus Grimaldi and Cumming 1999
†Microphorites pouilloni Ngô-Muller and Nel 2020
†Microphorites similis Grimaldi and Cumming 1999
†Microphorites utrillensis Arillo et al. 2008
Subfm. Neurigoninae Aldrich 1905
+
G. Neurigona Rondani 1856
G. Oncopygius Loew 1873
Subfm. Parathalassiinae Chvála 1981
+
G. †Electrophorella Cumming and Brooks 2002
G. Microphorella Becker 1909
G. †Retinitus Negrobov 1978
Subfm. Peloropeodinae Robinson 1970
+
G. †Palaeomedeterus Meunier 1895
+
Invalid names: Gheynia Meunier 1899 [synonym], Palaeochrysotus Meunier 1907 [synonym]
G. Peloropeodes Wheeler 1890
G. †Pristinmicrophor Tang et al. 2019
+
†Pristinmicrophor hukawngensis Tang et al. 2019
G. †Prosystenus Negrobov 1976
+
†Prosystenus zherichini Negrobov 1976
G. †Rajpardia Bickel 2022
+
†Rajpardia grimaldii Bickel 2022
Subfm. Rhaphiinae Bigot 1852
+
G. Nematoproctus Loew 1857
G. Rhaphium Meigen 1803
+
Invalid names: Xiphandrium Loew 1857 [synonym]
Subfm. Sciapodinae Becker 1917
+
G. Amblypsilopus Bigot 1888
G. Mesorhaga Schiner 1868
G. Sciapus Zeller 1842
+
Invalid names: Psilopus Meigen 1824 [replaced]
G. †Wheelerenomyia Meunier 1907
Invalid names: Agonosominae Aldrich 1905 [synonym]
Subfm. Sympycninae Aldrich 1905
+
G. Campsicnemus Haliday 1832
G. Chaetogonopteron de Meijere 1914
G. Neoparentia Robinson 1967
G. Teuchophorus Loew 1857
Invalid names: Campsicneminae Becker 1917 [synonym]
Subfm. Xanthochlorinae Aldrich 1905
+
G. Xanthochlorus Loew 1857
Invalid names: Argyra debellata Meunier 1907 [nomen dubium], Argyra debilis Meunier 1907 [nomen dubium], Argyra deceptoria Meunier 1907 [nomen dubium], Chrysotus setosus Giebel 1856 [nomen dubium], Eodolichopoditinae Hong 2002 [invalid subgroup], Hygroceleuthus titanus Meunier 1907 [nomen dubium], Lyroneurus venustus Meunier 1907 [nomen dubium], Medetera frauenfeldi Giebel 1856 [nomen dubium], Sciara pusilla Meunier 1899 [nomen nudum], Systenus ciliatus Meunier 1907 [nomen dubium], Thinophilus piraticus Meunier 1907 [nomen dubium], Thrypticus sobrius Meunier 1907 [nomen dubium]
G. †Ecommocydromia Schlüter 1978
+
†Ecommocydromia difficilis Schlüter 1978
Fm. Empididae Latreille 1804 [dagger fly]
+
G. Acallomyia Melander 1928
+
†Acallomyia probolaea Melander 1949
G. †Basilorhagio Ren 1995
+
Subfm. Brachystomatinae Melander 1908
+
Tr. Brachystomatini Melander 1908
Tr. Trichopezini Vaillant 1981
G. †Burmitempis Cockerell 1917
+
†Burmitempis halteralis Cockerell 1917
Subfm. Clinocerinae Schiner 1862
+
G. Proclinopyga Melander 1928
Invalid names: Fushunempites Hong 2002 [nomen nudum]
Subfm. Empidinae Latreille 1804
+
Tr. Chelipodini Hendel 1936
Tr. Empidini Latreille 1804
G. †Emplita Grimaldi and Cumming 1999
Tr. Hemerodromiini Schiner 1862
Tr. Hilarini Collin 1961
G. †Turonempis Grimaldi and Cumming 1999
Invalid names: Hemerodromiinae Schiner 1862 [synonym]
G. †Hasmona Giebel 1856
+
†Hasmona leo Giebel 1856
G. †Hilarites Heer 1856
+
†Hilarites bellus Heer 1856
G. †Oustaletimyia Meunier 1893
+
†Oustaletimyia succinorum Meunier 1893
G. †Progloma James 1937
+
†Progloma rohweri James 1937
Subfm. †Protempidinae Ussatchov 1968
+
G. †Homalocnemimus Mostovski 1999
G. †Protempis Ussatchov 1968
G. †Protoreogeton Mostovski 1999
G. †Protoedalea Cockerell 1921
+
†Protoedalea brachystoma Cockerell 1921
G. †Thirza Giebel 1856
+
†Thirza naumanni Giebel 1856
Invalid names: Brachystomatidae Melander 1908 [synonym], Tachydromia stilpon Meunier 1899 [nomen dubium]
Fm. Hybotidae Meigen 1820 [dance fly]
+
Subfm. Bicellariinae Sinclair and Cumming 2006
+
G. Hoplocyrtoma Melander 1928
G. †Grimaldipeza Solórzano-Kraemer et al. 2023
+
†Grimaldipeza coelica Solórzano-Kraemer et al. 2023
Subfm. Hybotinae Meigen 1820
+
G. †Eternia Martins-Neto et al. 1992
G. †Euthyneuriella Meunier 1908
G. Hybos Meigen 1803
G. †Palaeoleptopeza Meunier 1908
G. †Pseudoacarterus Waters 1989
G. †Syneproctus Solórzano-Kraemer et al. 2020
G. †Tremembella Martins-Neto et al. 1992
Invalid names: Hybotini Meigen 1820 [invalid subgroup]
Subfm. Ocydromiinae Schiner 1862
+
G. Leptopeza Macquart 1827
Subfm. Oedaleinae Chvala 1983
+
G. Oedalea Meigen 1820
G. †Palaeoedalea Meunier 1902
G. †Pouillonhybos Ngô-Muller et al. 2020
+
†Pouillonhybos venator Ngô-Muller et al. 2020
Subfm. Tachydromiinae Meigen 1822
+
G. †Archaeodrapetiops Martins-Neto et al. 1992
G. †Archiplatypalpus Kovalev 1974
G. †Cretoplatypalpus Kovalev 1978
G. †Drapetiella Meunier 1908
Tr. Drapetini Collin 1961
G. †Electrocyrtoma Cockerell 1917
G. †Mesoplatypalpus Grimaldi and Cumming 1999
Tr. Symballophthalmini Sinclair and Cumming 2006
Tr. Tachydromiini Meigen 1822
Invalid names: Corynetinae Kertesz 1909 [synonym]
G. †Megeana Meunier 1899
Fm. Ragadidae Sinclair 2016
+
Subfm. Ragadinae Sinclair 2016
+
G. Ragas Walker 1837
G. †Francomyina Jouault et al. 2020
+
†Francomyina incomparabilis Jouault et al. 2020
Fm. †Mysteromyiidae Grimaldi 2016
+
G. †Mysteromyia Grimaldi 2016
+
†Mysteromyia plumosa Grimaldi 2016
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Kiessling 2004, Bush and Bambach 2015

Age range

Maximum range based only on fossils: base of the Late/Upper Eifelian to the top of the Holocene or 389.60000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 387.7 Ma

Collections (371 total)

Oldest occurrences

Time interval Ma Country or state Original ID and collection number
Late/Upper Eifelian389.6 - 387.7Germany Schizophora blankenheimensis (235174)
Bajocian - Bathonian170.3 - 166.1Mongolia (Bayan-Khongor) Protoreogeton admirabilis (114615)
Bajocian - Bathonian170.3 - 166.1Russian Federation (Krasnoyarsk) Palaeoplatypygus zaitzevi (118010)
Callovian - Oxfordian166.1 - 157.3Kazakhstan (Chimkent) Homalocnemimus abjugatus (105970) Homalocnemimus sp. (105971) Homalocnemimus sp., Homalocnemimus abjugatus (105426) Protempis antennata, Homalocnemimus abjugatus (106101)
Early/Lower Berriasian145.0 - 140.2United Kingdom (England) Dianafranksia fisheri (125985)
Late/Upper Berriasian145.0 - 140.2United Kingdom (England) Opetiala shatalkini (103833)
Middle Berriasian145.0 - 140.2United Kingdom (England) Empididae indet. (104008)
Early/Lower Barremian130.0 - 125.45Lebanon Atelestites senectus, Phaetempis lebanensis, Avenaphora hispida, Chimeromyia acuta, Chimeromyia intriguea, Alonchoptera lebanica, Lonchopteromorpha asetocella, Lonchopterites prisca, Archisciada lebanensis (123017) Chimeromyia sp., Chimeromyia pilitibia, Chimeromyia mediobscura, Lebanopeza azari (122471) Microphorites similis, Microphorites oculeus, Chimeromyia reducta, Lebambromyia acrai, Platypezidae indet. (122359) Phaetempis lebanensis (123001) Trichinites cretaceus, Microphorites extinctus, Cyclorrhapha indet. (124276)
Early/Lower Barremian130.0 - 125.45Spain (Cataluña) Sciomyzidae larva sp. 3 (118325)
Late/Upper Barremian130.0 - 125.45China (Hebei) Basilorhagio venustus (107001)
Late/Upper Barremian130.0 - 125.45China (Liaoning) Protapiocera convergens (119055) Protempis minuta, Protapiocera ischyra, Protapiocera megista (138436)
Aptian125.0 - 113.0China (Shandong) Mesopetia tuanwangensis (125196) Palaeopetia laiyangensis, Lithopetia hirsuta (106313) Pseudapiocera shandongensis (161254) Sinolesta lata (154887)
Aptian125.0 - 113.0Russian Federation (Buryatia) Apioceridae indet. (118216) Empididae indet., Protapiocera sp. (113349) Hermaeomyia bisaetigera (125792) Hermaeomyia saeticaudata (122551) Hermaeomyia sp. (122548) Maritulus sospes (125797) Maritulus sospes, Hermaeomyia unisaetigera, Hermaeomyia bellula (125791) Parnasos dissimilis, Parnasos pessimus, Proplatypeza caelebs, Proplatypeza amabilis, Proplatypeza parva, Hermaeomyia baisica, Hermaeomyia unisaetigera, Hermaeomyia parva, Hermaeomyia armata, Hermaeomyia bellula (119498) Parnasos firmipes, Hermaeomyia armata (113353) Parnasos pessimus, Hermaeomyia armata (106103) Parnasos pessimus, Parnasos firmipes, Maritulus sospes, Proplatypeza amabilis, Proplatypeza parva, Hermaeomyia sp., Hermaeomyia armata, Hermaeomyia bellula (122549) Parnasos pessimus, Proplatypeza amabilis, Proplatypeza parva, Hermaeomyia unisaetigera (119441) Proplatypeza amabilis, Hermaeomyia bisaetigera (125413) Protapiocera sp. (105969) Sciadoceridae indet. (125795)
Aptian125.0 - 113.0Mongolia Proplatypeza bontsaganica, Proplatypeza secunda, Ironomyiidae indet., Hermaeomyia opinata, Palaeopetia tertia, Palaeopetia volacris, Hermaeomyia asaetigera, Palaeopetia mina, Palaeopetia gemina, Palaeopetia longisaetigera (103657)
Aptian125.0 - 113.0Mongolia (Govi-Altai) Bombyliidae indet., Empididae indet., Hybotidae indet. (122332)
Late/Upper Aptian122.46 - 113.0Brazil (Ceará) Araripogon pulchrus (149767) Asiloidea indet., Cretomydas santanensis, Asilidae indet. (122557) Cretomydas santanensis (215116) Cretothereva antiqua (125511) Therevidae indet., Araripogon sp. 2?, Araripogon axelrodi (122352)
Late/Upper Aptian122.46 - 113.0Australia (Victoria) Acalyptratae indet. (77192)
Albian113.0 - 100.5Jordan Kurnubempis palistinicus, Curvus khuludi (124847)
Early/Lower Albian112.03 - 109.0Spain Grimaldipeza sp. 1, Grimaldipeza coelica (109438)
Late/Upper Albian105.3 - 99.6Spain (Álava) Alavesia subiasi, Chimeromyina concilia, Chimeromyia alava, Euliphora grimaldii (108769)