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Arthropoda (arthropod)

Taxonomy

Arthropoda was named by Latreille (1829) [also attributed to Siebold and Stannius 1845]. It is extant.

It was assigned to Metazoa by Shoshani et al. (1989), Sepkoski (2002); to Animalia by Zhang and Shu (2005); to Ecdysozoa by Edgecombe et al. (2011); and to Panarthropoda by Zrzavy et al. (1998), Nielsen (2012).

Subtaxa

Actaeus, Agonata, Almenia, Aptera, Arachnoidea, Arachnomorpha, Artiopoda, Blattina splendens, Bradoriida (syn. Bradoriid, Archaeocopida), Branchiata, Bundenbachiellus, Burgessia, Clypecaris, Cryptocaris, Dalmaniopsis, Deuteropoda, Devonopilio, Devonopilio hutchinsoni, Dicerocaris, Dinocarida, Dinopilio parvus, Dongshanocaris, Douglasocaris, Emeiella, Ercaia, Esmeraldacaris, Euthycarcinoidea, Forfexicaris, Gaoqiaioella, Geracus, Geracus tubifer, Gigantopleurida, Goticaris, Graeophonus, Guangweicarididae, Habelia, Hadranax, Hanburia, Henningsmoenicaris, Huangshandongia, Jianfengia, Jiucunella, Kablikia, Kampecaris obanensis, Keurbos, Kleptothule, Kounicia, Liantuoia, Libellula carbonaria, Longquania, Marrellomorpha, Marrelomorpha, Megadictyon, Meristosoma, Merostomoidea, Monoculus, Motina, Myriapoda, Naptile, Notchia, Occacaris, Odaraiida, Orstenocarida, Palaeophonus arctus, Panlongia, Paratrilobita, Parioscorpio, Parvancorinomorpha, Parvibellus, Pauloterminus, Pedestria, Pisinnocaris, Platicytes, Plenocaris, Pleurolycosa, Pleurolycosa prolifera, Poliochera pustulatus, Priscansermarinus, Proricaris, Proschizoramia, Protocaris, Pseudoiulia, Pycnogonida (syn. Arachnopoda), Pygmaclypeatus, Radiodonta, Rhombicalvaria, Rhynimonstrum, Schizodiscus, Scorpionida, Serracaris, Shankouia, Siberion, Skaracarida, Synistata, Tanglangia, Tuzoida, Unogata, Utahcaris, Youti, Zagoracaris

Synonyms

Euarthropoda is a phylum.

It was synonymized subjectively with Arthropoda by Zrzavy et al. (1998).
Monoculus was named by Linnaeus (1758). It is extant.

It was considered a nomen oblitum by Fox (1951).

It was assigned to Insecta by Geoffroy (1762); to Pedestria by Scopoli (1763); to Aptera by Linnaeus (1758), Linnaeus (1761), Müller (1764), Linnaeus (1767); and to Synistata by Fabricius (1775).
Schizoramia was named by Bergström (1976).

It was synonymized subjectively with Arthropoda by Kiessling (2009).
Palaeophonus arctus was named by Matthew (1895). It is an impression. Its type locality is Fern Ledges, Plant bed 2, Carleton, Saint John, which is in a Westphalian A estuary/bay shale in the Lancaster Formation of Canada.

It was considered a nomen dubium by Dunlop and Garwood (2023).
Libellula carbonaria was named by Scudder (1876). It is an impression. Its type locality is Cossett's Pit, Sydney coalfield, Cape Breton, which is in a Westphalian C/Westphalian D terrestrial shale in Canada.

It was recombined as Graeophonus carbonarius by Scudder (1890), Petrunkevitch (1913), Petrunkevitch (1953), Dunlop et al. (2013); it was considered a nomen dubium by Scudder (1885), Dunlop (2018).
Blattina splendens was named by Göppert (1865). It is an impression. Its type locality is Ottendorf, which is in an Asselian fluvial-lacustrine horizon in the Otovice Formation of the Czech Republic.

It was considered a nomen dubium by Handlirsch (1906), Štamberg and Zajíc (2008).
Pleurolycosa was named by Fritsch (1904).

It was considered a nomen vanum by Dunlop et al. (2013), Selden (2021).
Arthrolycosa prolifera was named by Fritsch (1899). Its type specimen is CGH Me 77, an exoskeleton, and it is a compression fossil. Its type locality is Nýřany (Insect Collection), which is in a Moscovian lacustrine siliciclastic in the Kladno Formation of the Czech Republic.

It was recombined as Pleurolycosa prolifera by Fritsch (1904), Štamberg and Zajíc (2008); it was considered a nomen vanum by Petrunkevitch (1953), Dunlop et al. (2013), Selden (2021).
Graeophonus was named by Scudder (1890).

It was considered a nomen dubium by Dunlop (2018).

It was assigned to Paracharontidae by Dunlop et al. (2013), Engel and Grimaldi (2014); and to Paleoamblypygi by Garwood et al. (2017).
Poliochera pustulatus was named by Laurentiaux-Vieira and Laurentiaux (1963). It is a compression fossil. Its type locality is Chao-Ko-Chwang Mine, Kaiping, Tangshan, which is in a Westphalian D terrestrial siliciclastic in the Benxi Formation of China.

It was considered a nomen dubium by Selden (1992), Dunlop et al. (2013).
Dinopilio parvus was named by Petrunkevitch (1953). Its type specimen is BMNH In. 37101 and is an impression. Its type locality is Surface dump at Chislet Colliery, which is in a Westphalian B terrestrial siliciclastic in the United Kingdom.

It was considered a nomen dubium by Selden (2021).
Kounicia was named by Frič (1901). Its type is Kounicia bioculata.

It was considered a nomen dubium by Handlirsch (1907).
Geracus was named by Matthew (1897).

It was considered a nomen dubium by Handlirsch (1906).

It was assigned to Insecta by Matthew (1897).
Geracus tubifer was named by Matthew (1897). Its type specimen is NMBG 3012 and is an impression. Its type locality is Fern Ledges, Plant bed 2, Carleton, Saint John, which is in a Westphalian A estuary/bay shale in the Lancaster Formation of Canada.

It was considered a nomen dubium by Handlirsch (1906).
Synistata was named by Fabricius (1775). It is extant.

It was considered an invalid subgroup of Arthropoda by Clapham (2023).

It was assigned to Insecta by Fabricius (1775), Fabricius (1798).
Agonata was named by Fabricius (1775). It is extant.

It was considered an invalid subgroup of Arthropoda by Clapham (2023).

It was assigned to Insecta by Fabricius (1775).
Unogata was named by Fabricius (1775). It is extant.

It was considered an invalid subgroup of Arthropoda by Clapham (2023).

It was assigned to Insecta by Fabricius (1775).
Pedestria was named by Scopoli (1763). It is extant.

It was considered an invalid subgroup of Arthropoda by Clapham (2023).

It was assigned to Insecta by Scopoli (1763).
Aptera was named by Linnaeus (1758). It is extant.

It was considered an invalid subgroup of Arthropoda by Clapham (2023).

It was assigned to Insecta by Linnaeus (1758), Linnaeus (1761), Müller (1764), Schaeffer (1766), Linnaeus (1767), Strøm (1768), Drury (1770), Müller (1776).
Kampecaris obanensis was named by Peach (1899). Its type specimen is Institute of Geological Sciences, Edinburgh 10385 and is an impression. Its type locality is Kerrera, near Oban, which is in a Ludfordian terrestrial shale in the United Kingdom.

It was considered a nomen dubium by Almond (1985).
Devonopilio was named by Tihelka et al. (2020).

It was considered a nomen dubium by Pérez‐González and Schultz (2021).

It was assigned to Opiliones by Tihelka et al. (2020).
Devonopilio hutchinsoni was named by Tihelka et al. (2020). Its type specimen is BRSGM slide Db7365 and is a 3D body fossil. Its type locality is Rhynie cherts (Tait collection), which is in a Pragian pond chert in the Dryden Flags Formation of the United Kingdom.

It was considered a nomen dubium by Pérez‐González and Schultz (2021).
Zagoracaris was named by von Roy (2006). Its type is Zagoracaris fezouatensis.

It was considered a nomen nudum by von Roy (2006).

Synonymy list

Year	Name and author
1758	Aptera Linnaeus p. 341
1758	Monoculus Linnaeus p. 634
1761	Aptera Linnaeus p. 472
1761	Monoculus Linnaeus p. 497
1762	Monoculus Geoffroy p. 651
1763	Pedestria Scopoli p. 378
1763	Monoculus Scopoli p. 413
1764	Aptera Müller p. 89
1764	Monoculus Müller p. 95
1766	Aptera Schaeffer
1767	Aptera Linnaeus p. 1012
1767	Monoculus Linnaeus p. 1057
1768	Aptera Strøm p. 359
1770	Aptera Drury
1775	Monoculus Fabricius p. 294
1775	Synistata Fabricius p. 294
1775	Agonata Fabricius p. 399
1775	Unogata Fabricius p. 420
1776	Aptera Müller p. 183
1798	Synistata Fabricius p. 197
1829	Arthropoda Latreille
1865	Blattina splendens Göppert p. 276 figs. Pl 64, fig 11
1876	Libellula carbonaria Scudder p. 89 fig. 2
1890	Graeophonus Scudder p. 453
1890	Graeophonus carbonarius Scudder p. 454 figs. Pl 40, figs 2,3,6
1895	Palaeophonus arctus Matthew p. 98 figs. Pl 1, figs 7a,b
1897	Geracus Matthew p. 55
1897	Geracus tubifer Matthew p. 55 fig. 3
1899	Arthrolycosa prolifera Fritsch p. 61 figs. Pl 153, figs 1-3; text-fig 366
1899	Kampecaris obanensis Peach p. 122 figs. Pl 4, fig 2
1901	Kounicia Frič p. 175
1904	Pleurolycosa Fritsch p. 23
1904	Pleurolycosa prolifera Fritsch p. 23 fig. 29
1913	Graeophonus carbonarius Petrunkevitch p. 67 figs. Pl 5, figs 27-29; text-figs 29-31
1953	Graeophonus carbonarius Petrunkevitch p. 99
1953	Dinopilio parvus Petrunkevitch p. 110 fig. 197
1963	Poliochera pustulatus Laurentiaux-Vieira and Laurentiaux p. 25 figs. Pl 3, fig 3; text-fig 3
1976	Schizoramia Bergström
1989	Arthropoda Shoshani et al. p. 435
1998	Arthropoda Zrzavy et al.
2002	Arthropoda Sepkoski
2005	Arthropoda Zhang and Shu p. 187
2006	Zagoracaris von Roy
2008	Pleurolycosa prolifera Štamberg and Zajíc p. 57 fig. 36
2011	Arthropoda Edgecombe et al. p. 153 figs. Fig. 1
2012	Arthropoda Nielsen p. 240 figs. Fig. 42.1
2013	Graeophonus Dunlop et al. p. 190
2013	Graeophonus carbonarius Dunlop et al. p. 190
2014	Graeophonus Engel and Grimaldi p. 3
2017	Graeophonus Garwood et al. p. 12
2020	Devonopilio Tihelka et al. p. 314
2020	Devonopilio hutchinsoni Tihelka et al. p. 315 fig. 2

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia

Rank	Name	Author
—	Ecdysozoa	Aguinaldo et al. 1997
—	Panarthropoda
phylum	Arthropoda	Latreille 1829
phylum	Arthropoda	Latreille 1829

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Ph. Arthropoda Latreille 1829 [arthropod]

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G. †Actaeus

G. †Almenia Hou and Bergström 1997

†Almenia spinosa Hou and Bergström 1997

Or. †Bradoriida Raymond 1935

Fm. †Beyrichonidae Ulrich and Bassler 1931

G. †Aluta Mathew 1896

G. †Beyrichona Mathew 1886

Invalid names: Escasona Mathew 1902 [synonym]

G. †Eremos Moberg and Segerberg 1906

G. †Sellula Wiman 1903

Fm. †Bradoriidae Mathew 1902

G. †Bradoria Mathew 1899

†Bradoria scrutator Mathew 1899

Invalid names: Bradorona Mathew 1903 [synonym]

G. †Walcottella Ulrich and Bassler 1931

Fm. †Cambriidae Li 1975

G. †Petrianna Siveter 1986

Fm. †Comptalutidae Opik 1968

G. †Alutella

†Alutella elongta Peng et al. 2010

G. †Haoia Shu 1990

Fm. †Hipponicharionidae Sylvester-Bradley 1961

G. †Albrunnicola Martinsson 1979

†Albrunnicola bengtsoni Hinz-Schallreuter 1993

G. †Bicarinellata Stigall 2008

†Bicarinellata evansi Rode et al. 2003

Invalid names: Bicarinella Rode et al. 2003 [replaced]

G. †Cambria Netskaya and Ivanova 1956

G. †Hipponicharion Mathew 1886

†Hipponicharion ichnusum Elicki and Pillola 2004

Subfm. †Hipponicharioninae Sylvester-Bradley 1961

G. †Vojbokalina Khazanovich et al. 1984

†Vojbokalina magnifica Khazanovich et al. 1984

G. †Polyphyma Groom 1902

G. †Pseudobeyrichona Cui 1990

†Pseudobeyrichona longquanxiensis Cui 1987

†Pseudobeyrichona monile Streng and Geyer 2019

G. †Quadricona Topper et al. 2011

†Quadricona madonnae Topper et al. 2011

G. †Voibokalina Khazanovich et al. 1984

G. †Monceretia Vannier and Williams 2005

†Monceretia erisylvia Vannier and Williams 2005

G. †Qingquanella Huo 1985

G. †Sinskolutella Melnikova 1998

G. †Spinospitella Skovsted et al. 2006

†Spinospitella coronata Skovsted et al. 2006

Fm. †Svealutidae Opik 1968

G. †Liangshanella Huo 1956

†Liangshanella lubrica Qian and Zhang 1983

†Liangshanella sayutinae Melnikova 1988

G. †Yakutingella Melnikova 1998

Invalid names: Archaeocopida Sylvester-Bradley 1961 [synonym], Bradoriid Hu 2005 [synonym]

G. †Bundenbachiellus

G. †Burgessia

G. †Clypecaris Hou 1997

G. †Cryptocaris Barrande 1872

†Cryptocaris bohemica Barrande 1872

†Cryptocaris contracta Barrande 1872

†Cryptocaris obsoleta Barrande 1872

†Cryptocaris pulchra Barrande 1872

†Cryptocaris rhomboidea Barrande 1872

†Cryptocaris solida Barrande 1872

†Cryptocaris tardissima Barrande 1872

G. †Dalmaniopsis

G. †Dicerocaris

G. †Dongshanocaris Hou and Bergström 1998

†Dongshanocaris foliiformis Hou and Bergström 1998

G. †Douglasocaris

G. †Emeiella

G. †Ercaia Chen et al. 2001

G. †Esmeraldacaris Waggoner 2003

†Esmeraldacaris richardsonae Waggoner 2003

Subcl. †Euthycarcinoidea Gall and Grauvogel 1964

Or. †Euthycarcinida Gall and Grauvogel 1964

Fm. †Euthycarcinidae Handlirsch 1914

G. †Euthycarcinus Handlirsch 1914

†Euthycarcinus kessleri Handlirsch 1914

Fm. †Kottixerxidae Starobogatov 1988

G. †Heterocrania Hirst and Maulik 1926

†Heterocrania rhyniensis Hirst and Maulik 1926

Invalid names: Crania Hirst and Maulik 1926 [replaced]

G. †Forfexicaris Hou 1999

G. †Gaoqiaioella

G. †Goticaris

Fm. †Guangweicarididae Luo et al. 2007

G. †Guangweicaris Luo et al. 2007

†Guangweicaris spinatus Luo et al. 2007

G. †Habelia

G. †Hanburia Walcott 1916

G. †Henningsmoenicaris

G. †Huangshandongia

G. †Jianfengia

†Jianfengia multisegmentalis Hou 1987

G. †Jiucunella

G. †Kablikia

G. †Keurbos Gabbott et al. 2025

†Keurbos susanae Gabbott et al. 2025

G. †Kleptothule Budd 1995

†Kleptothule rasmusseni Budd 1995

G. †Liantuoia

G. †Longquania Luo et al. 2008

†Longquania bispinosa Luo et al. 2008

Cl. †Marrelomorpha Beurlen 1930

Or. †Acercostraca Lehmann 1955

Fm. †Vachonisiidae Tasch 1969

G. †Vachonisia

Invalid names: Mimetasterida Beurlen 1934 [empty]

G. †Meristosoma Robison and Wiley 1995

†Meristosoma paradoxum Robison and Wiley 1995

Cl. †Merostomoidea

G. †Nathorsthia

Invalid names: Emeraldellida Størmer 1944 [empty], Limulavida Walcott 1912 [empty]

G. †Motina

Cl. Myriapoda Latreille 1802

G. †Anthracerpes Meek and Worthen 1865

†Anthracerpes typus Meek and Worthen 1865

G. †Archicarabides Fritsch 1901

†Archicarabides pater Fritsch 1901

G. †Cambropodus

Cl. Chilopoda Latreille 1817 [centipede]

G. †Nisaukankoil Malkani 2019

†Nisaukankoil beakeri Malkani 2019

Unr. Notostigmophora

Or. Scutigeromorpha

Fm. †Crussolidae Shear et al. 1998

G. †Latzelia Scudder 1890

Fm. Scutigeridae Gervais 1837

G. †Phailawaghkankoil Malkani 2019

†Phailawaghkankoil derabugti Malkani 2019

Unr. Pleurostigmophora

Or. Lithobiomorpha

Fm. Henicopidae Pocock 1901

Fm. Lithobiidae

G. †Lithopendra Haug et al. 2023

†Lithopendra anjafliessae Haug et al. 2023

Unr. Phylactrometria

Or. †Devonobiomorpha Shear and Bonamo 1988

Unr. Epimorpha Haase 1880

G. †Rhyniognatha Tillyard 1928

†Rhyniognatha hirsti Tillyard 1928

Invalid names: Anamorpha Haase 1880 [empty], Succinipatopsidae Poinar 2000 [invalid subgroup], Succinipatopsis Poinar 2000 [nomen dubium], Succinipatopsis balticus Poinar 2000 [nomen dubium]

Unr. Dignatha Tiegs 1947

Subcl. Diplopoda Gervais 1844 [millipede]

Subcl. †Arthropleuridea Waterlot 1934

Or. †Arthropleurida Waterlot 1934

Subcl. Chilognatha Latreille 1802

Infracl. Helminthomorpha Pocock 1887

Infracl. Pentazonia Brandt 1833

Or. Chordeumatida Cook 1895

Fm. Craspedosomatidae Cook 1895

Subor. Heterochordeumatidea Shear 2000

G. †Decorotergum Jell 1983

†Decorotergum warrenae Jell 1983

Fm. Iulidae Leach 1814

G. Iulus Linnaeus 1758

Subcl. Penicillata Latreille 1825

Or. Polyxenida Lucas 1840

Fm. †Proiulidae Fritsch 1899

G. †Tomiulus Martynov 1936

Cl. Pauropoda

Or. Tetramerocerata

Fm. Pauropodidae Lubbock 1867

G. †Eileticus Scudder 1882

†Eileticus aequalis Scudder 1890

†Eileticus anthracinus Scudder 1882

G. †Eojulus Kušta 1886

†Eojulus fragilis Kušta 1886

G. †Hemiphoberia Fritsch 1899

†Hemiphoberia alternans Fritsch 1899

G. †Heterovorhoeffia Fritsch 1899

†Heterovorhoeffia crassa Fritsch 1899

G. †Juloites Haupt 1956

†Juloites quadripustulatus Haupt 1956

G. †Kralupia Fritsch 1907

†Kralupia carbonaria Fritsch 1907

G. †Palenarthrus Scudder 1890

†Palenarthrus impressus Scudder 1890

G. †Sandtneria Fritsch 1899

†Sandtneria gemmata Fritsch 1899

Cl. Symphyla Ryder 1880

Or. Scolopendrellida Hoffman 1969

Fm. Scolopendrellidae Newport 1845

G. Scolopendrella Gervais 1836

G. Symphylella Silvestri 1902

Fm. Scutigerellidae Bagnall 1913

G. Hanseniella Bagnall 1913

†Hanseniella baltica Scheller 2004

G. Scutigerella Ryder 1882

†Scutigerella baltica Scheller 2004

†Scutigerella dominicana Poinar and Edwards 1995

Invalid names: Eoscolopendridae Scudder 1890 [invalid subgroup], Gerascutigeridae Scudder 1890 [invalid subgroup], Tiphoscorpionoidea Kjellesvig-Waering 1986 [invalid subgroup]

G. †Naptile

G. †Notchia Lerosey-Aubril 2015

†Notchia weugi Lerosey-Aubril 2015

G. †Occacaris Hou 1999

Or. †Odaraiida Simonetta and Delle Cave 1975

Fm. †Pectocariidae Hou 1999

G. †Pectocaris Hou 1999

†Pectocaris spatiosa Hou 1999

Invalid names: Odaraia [synonym]

G. †Panlongia Liu et al. 2006

†Panlongia spinosa Liu et al. 2006

†Panlongia tetranodusa Liu et al. 2006

Unr. †Parvancorinomorpha Lin et al. 2006

Fm. †Parvancorinidae Glaessner 1979

G. †Primicaris Zhang et al. 2003

†Primicaris larvaformis Zhang et al. 2003

†Primicaris lavaformis Zhang et al. 2003

G. †Skania Walcott 1931

†Skania fragilis Walcott 1931

†Skania sundbergi Lin et al. 2006

G. †Parvibellus Liu et al. 2022

†Parvibellus atavus Liu et al. 2022

G. †Pauloterminus Taylor 2002

†Pauloterminus spinodorsalis Taylor 2002

G. †Pisinnocaris Hou and Bergström 1998

G. †Platicytes Garvey and Hasiotis 2008

†Platicytes lioparadus Garvey and Hasiotis 2008

G. †Plenocaris

G. †Priscansermarinus Collins and Rudkin 1981

†Priscansermarinus barnetti Collins and Rudkin 1981

G. †Proricaris Baily 1870

†Proricaris machenrici Baily 1870

G. †Protocaris

†Protocaris marshi Walcott 1884

G. †Pseudoiulia Hou and Bergström 1998

†Pseudoiulia cambriensis Hou and Bergström 1998

Cl. Pycnogonida Latreille 1810 [sea spider]

G. †Austrodecus

G. †Cambropycnogon Waloszek and Dunlop 2002

†Cambropycnogon klausmuelleri Waloszek and Dunlop 2002

G. †Flagellopantopus Poschmann and Dunlop 2006

†Flagellopantopus blocki Poschmann and Dunlop 2006

G. †Haliestes Siveter et al. 2004

†Haliestes dasos Siveter et al. 2004

Fm. †Palaeoisopodidae Dubinin 1957

G. †Palaeoisopus Broili 1928

†Palaeoisopus problematicus Broili 1928

G. †Palaeomarachne Rudkin et al. 2013

†Palaeomarachne granulata Rudkin et al. 2013

Or. †Palaeopantopoda

Fm. †Palaeopantopodidae Hedgpeth 1955

G. †Palaeopantopus Broili 1929

†Palaeopantopus maucheri Broili 1929

G. †Palaeothea Bergstrom et al. 1980

†Palaeothea devonica Bergstroem et al. 1980

Or. Pantopoda Gerstaecker 1863

Fm. Ammotheidae Dohrn 1881

Invalid names: Ammothellidae Fry 1978 [synonym], Ephyrogymnidae Fry 1978 [synonym], Eurycididae Sars 1891 [synonym], Oorhynchidae Schimkewitsch 1913 [synonym], Paranymphonidae Fry 1978 [synonym], Sericosuridae Fry 1978 [synonym], Tanystylidae Shimkevich 1913 [synonym], Trygaeidae Fry 1978 [synonym]

Fm. Ascorhynchidae Hoek 1881

G. Eurycyde Schiödte 1857

†Eurycyde golem Sabroux et al. 2019

Fm. Callipallenidae Hilton 1942

Invalid names: Cheilapallenidae Fry 1978 [synonym], Clavigeropallenidae Fry 1978 [synonym], Hannonidae Fry 1978 [synonym], Metapallenidae Fry 1978 [synonym], Pallenidae Wilson 1878 [synonym], Queubidae Fry 1978 [synonym], Stylopallenidae Fry 1978 [synonym]

Fm. Colossendeidae Jarzynsky 1870

Subfm. Colossendeinae Jarzynsky 1870

G. †Colossopantopodus Charbonnier et al. 2007

Invalid names: Pasithoidae Sars 1891 [synonym], Rhopalorhynchidae Fry 1978 [synonym]

Fm. Endeidae Norman 1904

G. †Palaeoendeis Charbonnier et al. 2007

†Palaeoendeis elmii Charbonnier et al. 2007

Fm. †Palaeopycnogonididae Sabroux et al. 2023

G. †Palaeopycnogonides Charbonnier et al. 2007

†Palaeopycnogonides gracilis Charbonnier et al. 2007

Fm. Phoxichilidiidae Sars 1891

Invalid names: Anoplodactylidae Fry 1978 [synonym], Phoxiphilyridae Fry 1978 [synonym]

Invalid names: Austrodecidae Stock 1954 [empty], Nymphonidae Wilson 1878 [empty], Pallenopsidae Fry 1978 [empty], Pycnogonidae Wilson 1878 [empty], Rhynchothoracidae Thompson 1909 [empty]

G. †Pentapantopus Kühl et al. 2013

†Pentapantopus vogteli Kühl et al. 2013

Invalid names: Arachnopoda Dana 1853 [synonym], Palaeoisopoda [empty]

G. †Pygmaclypeatus Zhang et al. 2000

G. †Rhombicalvaria Hou 1987

†Rhombicalvaria acantha Hou 1987

G. †Rhynimonstrum Anderson and Trewin 2003

†Rhynimonstrum dunlopi Anderson and Trewin 2003

G. †Schizodiscus Clarke 1888

Cl. Scorpionida Latreille 1817 [scorpion]

G. †Waeringoscorpio Størmer 1970

Invalid names: Euscorpionina Petrunkevitch 1949 [empty], Neoscorpionina [empty], Palaeoscorpiones Stockwell 1989 [empty], Proscorpionidae Scudder 1885 [empty]

G. †Serracaris

G. †Shankouia Waloszek et al. 2005

†Shankouia zhenghei Chen et al. 2004

G. †Tanglangia Lou and Hu 1999

Or. †Tuzoida Simonetta and Delle Cave 1975

Fm. †Tuzoiidae Raymond 1935

G. †Tuzoia Walcott 1912

†Tuzoia australis Glaessner 1979

†Tuzoia bispinosa Yuan and Zhao 1999

Invalid names: Tuzoia magna Yuan and Zhao 1999 [synonym]

†Tuzoia burgessensis Resser 1929

†Tuzoia canadensis Resser 1929

Invalid names: Tuzoia praemorsa Resser 1929 [synonym]

†Tuzoia dunbari Resser 1929

†Tuzoia guntheri Robison and Richards 1981

†Tuzoia limba Shu 1990

†Tuzoia manchuriensis Resser and Endo 1937

†Tuzoia parva Walcott 1912

†Tuzoia peterseni Robison and Richards 1981

†Tuzoia polleni Resser 1929

Invalid names: Tuzoia getzi Resser 1929 [synonym], Tuzoia nitida Resser and Howell 1938 [synonym], Tuzoia nodosa Resser 1929 [synonym], Tuzoia spinosa Resser 1929 [synonym], Tuzoia vermontensis Resser 1929 [synonym]

†Tuzoia retifera Walcott 1912

†Tuzoia sinensis Pan 1957

G. †Utahcaris Conway Morris and Robison 1988

†Utahcaris orion Conway Morris and Robison 1988

Invalid names: Agonata Fabricius 1775 [invalid subgroup], Aptera Linnaeus 1758 [invalid subgroup], Arachnoidea Guthörl 1934 [empty], Blattina splendens Göppert 1865 [nomen dubium], Branchiata Fitzinger 1843 [empty], Devonopilio Tihelka et al. 2020 [nomen dubium], Devonopilio hutchinsoni Tihelka et al. 2020 [nomen dubium], Dinopilio parvus Petrunkevitch 1953 [nomen dubium], Euarthropoda [synonym], Geracus Matthew 1897 [nomen dubium], Geracus tubifer Matthew 1897 [nomen dubium], Gigantopleurida Guthörl 1934 [empty], Graeophonus Scudder 1890 [nomen dubium], Kampecaris obanensis Peach 1899 [nomen dubium], Kounicia Frič 1901 [nomen dubium], Libellula carbonaria Scudder 1876 [nomen dubium], Monoculus Linnaeus 1758 [nomen oblitum], Orstenocarida Müller and Walossek 1988 [empty], Palaeophonus arctus Matthew 1895 [nomen dubium], Paratrilobita [empty], Pedestria Scopoli 1763 [invalid subgroup], Pleurolycosa Fritsch 1904 [nomen vanum], Pleurolycosa prolifera Fritsch 1899 [nomen vanum], Poliochera pustulatus Laurentiaux-Vieira and Laurentiaux 1963 [nomen dubium], Schizoramia Bergström 1976 [synonym], Skaracarida Müller and Walossek 1985 [empty], Synistata Fabricius 1775 [invalid subgroup], Unogata Fabricius 1775 [invalid subgroup], Zagoracaris von Roy 2006 [nomen nudum]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

No ecological data are available

Age range: base of the Pridoli to the top of the Kungurian or 422.70000 to 274.40000 Ma

Collections (2 total)

Time interval	Ma	Country or state	Original ID and collection number
Pridoli	422.7 - 419.62	China (Xinjiang)	Sciadophyton pristinum (10931)
Sakmarian - Kungurian	293.52 - 274.4	USA (Kansas)	Eukaryota indet. (32659)