PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Condylarthra

Mammalia - Condylarthra

Taxonomy

Condylarthra was named by Cope (1881). It is not extant.

It was reranked as the unranked clade Condylarthra by Cope (1889); it was reranked as the suborder Condylarthra by Flower and Lydekker (1891), Gadow (1898), Hay (1902), Gregory (1910) and McKenna (1956); it was reranked as the suborder Condylarthri by Jaekel (1911).

It was assigned to Taxeopoda by Cope (1889); to Ungulata by Flower and Lydekker (1891), Gadow (1898) and Hay (1902); to Protungulata by Gregory (1910); to Ungulati by Jaekel (1911); to Mammalia by McKenna (1956); to Phenacodonta by McKenna (1975); to Eutheria by Hooker (1986); to Eutheria by Sloan and Van Valen (1965), Kielan-Jaworowska et al. (1979) and Carroll (1988); and to Epitheria by Kielan-Jaworowska et al. (2004).

Subtaxa

Desmatoclaenus, Didolodontoidea, Hyopsodontidae (syn. Mioclaenidae, Limnotheriidae), Meniscotheridae, Paroxyclaenidae (syn. Kochictidae), Pleuraspidotheriidae

Synonymy list

Year	Name and author
1881	Condylarthra Cope
1889	Condylarthra Cope p. 876
1891	Condylarthra Flower and Lydekker p. 90
1898	Condylarthra Gadow p. 45
1902	Condylarthra Hay p. 601
1910	Condylarthra Gregory p. 466
1911	Condylarthri Jaekel p. 238
1956	Condylarthra McKenna p. 737
1965	Condylarthra Sloan and Van Valen p. 226
1975	Condylarthra McKenna p. 43
1979	Condylarthra Kielan-Jaworowska et al. p. 223
1986	Condylarthra Hooker p. 339
1988	Condylarthra Carroll
2004	Condylarthra Kielan-Jaworowska et al. pp. 495, 513

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Deuterostomia
phylum	Chordata	Haeckel 1874
subphylum	Vertebrata
superclass	Gnathostomata
—	Osteichthyes	()
subclass	Sarcopterygii	()
subclass	Dipnotetrapodomorpha	(Nelson 2006)
subclass	Tetrapodomorpha	()
—	Tetrapoda
—	Reptiliomorpha
—	Anthracosauria
subclass	Amphibiosauria	Kuhn 1967
—	Cotylosauria	()

Rank	Name	Author
—	Amniota
subclass	Synapsida
—	Therapsida	()
infraorder	Cynodontia	()
—	Mammaliamorpha
—	Mammaliaformes
class	Mammalia
—	Cladotheria
—	Zatheria
subclass	Tribosphenida	()
subclass	Theria
—	Eutheria	()
—	Placentalia
—	Epitheria
order	Condylarthra	Cope 1881
order	Condylarthra	Cope 1881

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Or. †Condylarthra Cope 1881

show all | hide all

G. †Desmatoclaenus Gazin 1941

†Desmatoclaenus dianae Van Valen 1978

†Desmatoclaenus hermaeus Gazin 1941

Invalid names: Desmatoclaenus paracreodus Gazin 1941 [synonym]

†Desmatoclaenus mearae Van Valen 1978

†Desmatoclaenus protogonioides Cope 1882

Fm. †Hyopsodontidae Trouessart 1879 [condylarth]

G. †Asiohyopsodus Tong and Wang 2006

†Asiohyopsodus confuciusi Tong and Wang 2006

G. †Bubogonia Johnston and Fox 1984

†Bubogonia bombadili Van Valen 1978

†Bubogonia saskia Johnston and Fox 1984

G. †Chacomylus Williamson and Weil 2011

†Chacomylus sladei Williamson and Weil 2011

G. †Choeroclaenus Simpson 1937

†Choeroclaenus turgidunculus Cope 1888

Subfm. †Hyopsodontinae Trouessart 1879

G. †Decoredon Xu 1977

G. †Obtususdon Xu 1977

†Obtususdon hanhuaensis Xu 1977

G. †Hyopsodus Leidy 1870

†Hyopsodus despiciens Matthew 1909

†Hyopsodus fangxianensis Huang 1995

†Hyopsodus fastigatus Russell and Wickenden 1933

†Hyopsodus huashigouensis Tong 1989

†Hyopsodus itinerans Godinot 1978

†Hyopsodus latidens Denison 1937

†Hyopsodus lepidus Matthew 1909

†Hyopsodus loomisi McKenna 1960

†Hyopsodus lysitensis Gingerich 1974

†Hyopsodus markmani Abel and Cook 1925

†Hyopsodus marshi Osborn 1902

†Hyopsodus mentalis Cope 1875

Invalid names: Hyopsodus lemoinianus Cope 1882 [synonym]

†Hyopsodus minor Loomis 1905

†Hyopsodus minusculus Leidy 1873

†Hyopsodus miticulus Cope 1874

†Hyopsodus paulus Leidy 1870

Invalid names: Hyopsodus distans Marsh 1875 [synonym], Hyopsodus vicarius Cope 1873 [synonym], Stenacodon rarus Marsh 1872 [synonym]

†Hyopsodus pauxillus Gingerich 1994

†Hyopsodus powellianus Cope 1884

Invalid names: Hyopsodus browni Loomis 1905 [synonym], Hyopsodus jacksoni Loomis 1905 [synonym], Hyopsodus lawsoni Loomis 1905 [synonym]

†Hyopsodus pygmaeus Cope 1872

†Hyopsodus sholemi Krishtalka 1979

†Hyopsodus simplex Loomis 1905

†Hyopsodus tonksi Eaton 1982

†Hyopsodus uintensis Osborn 1902

†Hyopsodus walcottianus Matthew 1915

†Hyopsodus wardi Hooker 1979

†Hyopsodus wortmani Osborn 1902

Invalid names: Lemuravus Marsh 1875 [synonym], Stenacodon Marsh 1872 [synonym]

G. †Lessnessina Hooker 1979

†Lessnessina khushuensis Hooker and Dashzeveg 2003

†Lessnessina packmani Hooker 1979

†Lessnessina praecipuus Russell 1987

G. †Litaletes Simpson 1935

†Litaletes disjunctus Simpson 1935

†Litaletes gazinensis Robison 1986

†Litaletes gazini Robison 1986

†Litaletes ondolinde Van Valen 1978

G. †Midiagnus Russell 1987

†Midiagnus gracilis Kondrashov 2004

G. †Mioclaenus Cope 1881

†Mioclaenus baldwini Cope 1882

†Mioclaenus turgidus Cope 1881

Invalid names: Mioclaenus lydekkerianus Cope 1888 [synonym], Mioclaenus zittelianus Cope 1888 [synonym]

G. †Molinodus Muizon and Marshall 1987

†Molinodus suarezi Muizon and Marshall 1987

G. †Promioclaenus Trouessart 1904

†Promioclaenus acolytus Cope 1882

Invalid names: Ellipsodon aquilonius Simpson 1935 [synonym], Mioclaenus minimus Cope 1882 [synonym]

†Promioclaenus lemuroides Matthew 1897

Invalid names: Ellipsodon shepherdi Gazin 1939 [synonym], Jepsenia mantiensis Gazin 1939 [synonym]

†Promioclaenus pipiringosi Gazin 1956

†Promioclaenus thnetus Scott et al. 2013

†Promioclaenus walshi Lofgren et al. 2014

Invalid names: Jepsenia Gazin 1939 [synonym]

G. †Protoselene Matthew 1897

†Protoselene ashtoni Lofgren et al. 2014

†Protoselene griphus Gazin 1939

†Protoselene novissimus Gazin 1956

†Protoselene opisthacus Cope 1882

Invalid names: Dracoclaenus Gazin 1939 [synonym]

Subfm. †Tricuspiodontinae Simpson 1929

G. †Paratricuspiodon Russell 1964

†Paratricuspiodon krumbiegeli Russell 1964

G. †Tricuspiodon Lemoine 1885

†Tricuspiodon magistrae Russell 1964

†Tricuspiodon rutimeyer Lemoine 1891

†Tricuspiodon sobrinus Russell 1980

G. †Valenia de Muizon and Cifelli 2000

†Valenia wilsoni Van Valen 1978

G. †Yuodon Chow et al. 1973

†Yuodon protoselenoides Chow et al. 1973

Invalid names: Limnotheriidae Marsh 1875 [synonym], Mioclaenidae Osborn and Earle 1895 [synonym], Pleuraspidotheriinae Zittel 1892 [empty]

Fm. †Paroxyclaenidae Weitzel

G. †Kopidodon Weitzel 1933

Subfm. †Merialinae Russell and Godinot 1988

G. †Euhookeria Russell and Godinot 1988

†Euhookeria hopwoodi Cray 1973

G. †Merialus Russell and Godinot 1988

†Merialus martinae Russell and Godinot 1988

Subfm. †Paroxyclaeninae Weitzel 1933

G. †Paravulpavoides Harrison 2009

G. †Paroxyclaenus Teilhard de Chardin 1922

†Paroxyclaenus lemuroides Teilhard de Chardin 1922

G. †Pugiodens Matthes 1952

†Pugiodens mirus Matthes 1952

Invalid names: Russellites Van Valen 1965 [synonym]

G. †Spaniella Crusafont-Pairo and Russell 1968

G. †Vulpavoides Matthes 1952

†Vulpavoides cooperi Hooker 1986

†Vulpavoides germanica Matthes 1952

†Vulpavoides simplicidens Van Valen 1965

†Vulpavoides vanvaleni Russell and Godinot 1988

Invalid names: Kochictidae Kretzoi 1943 [synonym]

Fm. †Pleuraspidotheriidae Zittel 1892

G. †Hilalia Maas et al. 2001

†Hilalia robusta Maas et al. 2001

†Hilalia saribeya Maas et al. 2001

†Hilalia selanneae Maas et al. 2001

†Hilalia sezerorum Maas et al. 2001

G. †Orthaspidotherium Lemoine 1885

†Orthaspidotherium edwardsi Lemoine 1885

G. †Parabunodon Ducrocq and Sen 1991

†Parabunodon anatolicum Ducrocq and Sen 1991

G. †Pleuraspidotherium Lemoine 1878

†Pleuraspidotherium aumonieri Russell 1964

Invalid names: Didolodontoidea Cifelli 1983 [empty], Meniscotheridae Cope 1882 [empty]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

phosphaticsubp

Environment:

terrestrialsubc

Locomotion:

actively mobilec

Life habit:

scansorialsubc

Diet:

insectivoresubc

Reproduction:

viviparoussubc

Created:

2005-09-06 20:43:42

Modified:

2005-09-06 22:44:32

Source: subc = subclass, c = class, subp = subphylum

References: Carroll 1988, Ji et al. 2002, Lillegraven 1979, Hendy et al. 2009

Age range: base of the Torrejonian to the top of the Duchesnean or 63.80000 to 37.00000 Ma

Collections (958 total)

Time interval	Ma	Country or state	Original ID and collection number
Pu3	66.0 - 63.3	USA (New Mexico)	Tiznatzinia sp., Valenia sp., Chacomylus sladei (26575)
Puercan	66.0 - 63.3	USA (Utah)	Desmatoclaenus paracreodus, Promioclaenus wilsoni (14736) Desmatoclaenus sp. (14739 224186) Ellipsodon sp. b, Desmatoclaenus hermaeus, Desmatoclaenus paracreodus (14790) Litaletes gazinensis (14692) Promioclaenus sp., Desmatoclaenus paracreodus (14670)
Puercan	66.0 - 63.3	USA (Colorado)	Mioclaenidae indet., Desmatoclaenus protogonioides (14678 14792)
Puercan	66.0 - 63.3	USA (New Mexico)	Bomburia prisca (193949 193950 193951 193952) Bomburia priscus, Bubogonia bombadili, Choeroclaenus turgidunculus, Platymastus palantir, Tiznatzinia vanderhoofi, Promioclaenus wilsoni (14760) Bubogonia bombadili, Ellipsodon grangeri, Tiznatzinia vanderhoofi, Bomburia prisa, Tiznatzinia turgidunculus, Promioclaenus wilsoni (14662) Choeroclaenus turgidunculus (14794) Choeroclaenus turgidunculus, Desmatoclaenus dianae, Desmatoclaenus protogonioides (14703) Desmatoclaenus sp., Desmatoclaenus sp. (14795) Ellipsodon priscus (193953) Promioclaenus wilsoni (14657)
Puercan	66.0 - 63.3	Canada (Saskatchewan)	Condylarthra indet. (14737) Desmatoclaenus sp. (14691) Hyopsodontidae indet. (14756) Litaletes sp., Bubogonia saskia (14768)
Puercan	66.0 - 63.3	USA (Montana)	Condylarthra indet. (108198) Mioclaenidae indet. (14809)
Puercan	66.0 - 63.3	USA (Wyoming)	Condylarthra indet. (14722) Promioclaenus wilsoni (14706 14729) Protoselene sp. (14720)
Early/Lower Paleocene	66.0 - 61.7	China (Guangdong)	Yuodon protoselenoides, Palasiodon siurenensis (176607) Yuodon protoselenoides, Promioclaenus siurenensis (140280)
Danian	66.0 - 61.6	USA (North Dakota)	Condylarthra indet. (14962)
Danian	66.0 - 61.6	Belgium (Hainaut)	Pleuraspidotheriidae indet. (132949)
Danian	66.0 - 61.6	Bolivia (Mizque)	Molinodus suarezi, Pucanodus gagnieri, Andinodus boliviensis, Simoclaenus sylvaticus (138709) Molinodus suarezi, Tiuclaenus minutus, Kollpania tiupampina, Tiuclaenus cotasi, Tiuclaenus robustus, Pucanodus gagnieri, Simoclaenus sylvaticus (13501) Tiuclaenus cotasi (138710)
Puercan - Torrejonian	66.0 - 60.9	USA (New Mexico)	Ellipsodon grangeri (193958) Ellipsodon yotankae (193959 193960 193961 193962 193963 193964 193965 193966)
To2	63.8 - 60.9	Canada (Alberta)	Promioclaenus acolytus (14894)
Torrejonian	63.8 - 60.9	USA (New Mexico)	Condylarthra indet., Promioclaenus sp., Mioclaenus turgidus, Protoselene opisthacus (14812) Ellipsodon grangeri (14834) Ellipsodon grangeri, Mioclaenus turgidus (14811) Ellipsodon grangeri, Mioclaenus turgidus, Promioclaenus acolytus, Promioclaenus lemuroides, Protoselene opisthacus (14832 14873) Ellipsodon inaequidens (193954 193955 193956 193957) Ellipsodon inaequidens, Mioclaenus turgidus, Promioclaenus acolytus, Promioclaenus lemuroides, Protoselene opisthacus (14824) Mioclaenus sp. (14869 14870) Mioclaenus sp., Ellipsodon grangeri, Promioclaenus acolytus (14867) Mioclaenus sp., Ellipsodon yotankae, Mioclaenus turgidus, Promioclaenus acolytus, Promioclaenus lemuroides, Protoselene opisthacus, Ellipsodon sp. (14816) Mioclaenus sp., Hyopsodontidae indet., Promioclaenus acolytus, Promioclaenus lemuroides, Protoselene opisthacus (14868) Mioclaenus turgidus (14819 14862 14888) Mioclaenus turgidus, Promioclaenus acolytus, Promioclaenus lemuroides, Protoselene opisthacus (14833 14835 14865 14893) Mioclaenus turgidus, Promioclaenus acolytus, Protoselene opisthacus (14815) Mioclaenus turgidus, Protoselene griphus (14859) Promioclaenus acolytus (14892) Protoselene opisthacus (14825) Protoselene sp. (14813) Tricentes inaequidens (14836)
Torrejonian	63.8 - 60.9	USA (Montana)	Ellipsodon aquilonius (14913 14915) Litaletes disjunctus, Ellipsodon aquilonius (14920) Litaletes disjunctus, Promioclaenus aquilonius (14906) Litaletes mantiensis (14885)
Torrejonian	63.8 - 60.9	USA (Utah)	Desmatoclaenus sp. (14883) Promioclaenus shepherdi (14889) Promioclaenus sp., Ellipsodon grangeri, Ellipsodon sternbergi, Promioclaenus lemuroides, Dracoclaenus griphus (14830)
Torrejonian	63.8 - 60.9	Canada (Alberta)	Condylarthra indet., Promioclaenus sp. (14828) Promioclaenus thnetus (143197 143198 143200 194573)
Torrejonian	63.8 - 60.9	USA (Texas)	Promioclaenus acolytus (14884) Promioclaenus sp. (14887) Protoselene opisthacus (14975) Protoselene sp., Ellipsodon priscus (14829)
Torrejonian	63.8 - 60.9	USA (Wyoming)	Litaletes disjunctus (14900 14903) Litaletes disjunctus, Promioclaenus acolytus (14896 14898 14905) Litaletes disjunctus, Promioclaenus aquilonius (14912) Litaletes sp., Promioclaenus acolytus (14886) Promioclaenus acolytus (14897 14899 14902)
Torrejonian - Tiffanian	63.8 - 56.8	USA (Wyoming)	Mioclaenus turgidus, Promioclaenus lemuroides (14820) Protoselene sp., Promioclaenus acolytus, Promioclaenus lemuroides (14851)
Middle Paleocene	61.7 - 58.7	China (Anhui)	Obtususdon hanhuaensis (132942 132964)
Middle Paleocene	61.7 - 58.7	China (Jiangxi)	Hyopsodontidae indet. (176612)
Ti1	61.7 - 56.8	Canada (Alberta)	Desmatoclaenus sp., Claenodon sp. (14939)
Ti3	61.7 - 56.8	USA (Colorado)	Promioclaenus acolytus (99821 99822)
Ti3	61.7 - 56.8	Canada (Alberta)	Hyopsodontidae indet. (14992)
Tiffanian	61.7 - 56.8	USA (Wyoming)	Desmatoclaenus mearae, Promioclaenus pipiringosi, Protoselene novissimus, Promioclaenus sp. (14989) Litaletes disjunctus, Promioclaenus acolytus (15013) Litaletes disjunctus, Promioclaenus acolytus, Promioclaenus lemuroides (14965) Litaletes disjunctus, Promioclaenus aquilonius (198263) Promioclaenus acolytus (15012 15014 15015 198262 198265) Promioclaenus pipiringosi (14964) Promioclaenus sp. (15067) Promioclaenus sp., Litaletes disjunctus, Promioclaenus acolytus (198264) Protoselene opisthacus, Promioclaenus sp. (15034)
Tiffanian	61.7 - 56.8	USA (Colorado)	Tetraclaenodon minor (14935)
Tiffanian	61.7 - 56.8	USA (California)	Promioclaenus walshi (154724 224203) Protoselene ashtoni (224209)
Tiffanian	61.7 - 56.8	USA (Montana)	Ellipsodon sp. (15050) Haplaletes sp. (15037)
Selandian	61.6 - 59.2	Germany (Saxony-Anhalt)	Paratricuspiodon krumbiegeli (74643)
Thanetian	59.2 - 56.0	Romania	Condylarthra indet. (83307 138239)
Thanetian	59.2 - 56.0	Spain (Cataluña)	Condylarthra indet. (79304 79305 79306) Pleuraspidotherium sp. (79307)
Thanetian	59.2 - 56.0	Morocco (Meknès-Tafilalet)	Condylarthra indet. (88301)
Thanetian	59.2 - 56.0	France (Champagne-Ardenne)	Orthaspidotherium edwardsi, Pleuraspidotherium aumonieri (137125) Orthaspidotherium edwardsi, Tricuspiodon magistrae, Tricuspiodon sobrinus (68428) Tricuspiodon magistrae, Tricuspiodon sobrinus (87943) Tricuspiodon sp., Tricuspiodon rutimeyer, Tricuspiodon sobrinus (87925)
Thanetian - Early/Lower Eocene	59.2 - 48.6	France (Marne)	Orthaspidotherium edwardsi, Pleuraspidotherium aumonieri (133367)
Late/Upper Paleocene	58.7 - 55.8	USA (Wyoming)	Hyopsodontidae indet. (15024)
Late/Upper Paleocene	58.7 - 55.8	France (Oise)	Pleuraspidotheriidae n. gen. (166598)
Clarkforkian	56.2 - 54.9	USA (Colorado)	Hyopsodontidae indet. (15239)
Clarkforkian	56.2 - 54.9	USA (Wyoming)	Hyopsodus sp. (15114 15165)
Casamayoran	56.0 - 48.0	Argentina (Chubut)	Pascualodus patagoniensis (186880)
Ypresian	56.0 - 47.8	France (Provence-Alpes-Côte d'Azur)	Merialus martinae (178050)
Ypresian	56.0 - 47.8	Turkey (Amasya)	Parabunodon anatolicum (84608)
Ypresian	56.0 - 47.8	Morocco	Condylarthra indet. (133216)
Ypresian	56.0 - 47.8	United Kingdom (England)	Hyopsodus wardi (133368) Lessnessina packmani, Hyopsodus wardi (38868)
Ypresian	56.0 - 47.8	France (Marne)	Paroxyclaenidae indet. (35537) Paroxyclaenidae indet., Paroxyclaenus sp. (35533 35535)
Ypresian	56.0 - 47.8	United Kingdom	Hyopsodus wardi (87950)
Ypresian	56.0 - 47.8	France (Oise)	Hyopsodus sp. (68174)
Ypresian - Lutetian	56.0 - 41.3	Algeria (Bechar region)	Condylarthra indet. (77014)
Eocene	56.0 - 33.9	Germany (Sachsen-Anhalt)	Vulpavoides germanica (45771)
Graybullian	55.8 - 50.3	USA (Wyoming)	Hyopsodus loomisi (15875) Hyopsodus miticulus, Hyopsodus simplex (15701)
Lostcabinian	55.8 - 50.3	USA (Wyoming)	Hyopsodus miticulus (15997 15999 16003) Hyopsodus paulus (15901 15903) Hyopsodus paulus, Hyopsodus wortmani (16073) Hyopsodus sp., Hyopsodus paulus (16078) Hyopsodus sp., Hyopsodus walcottianus, Hyopsodus wortmani (15915) Hyopsodus walcottianus, Hyopsodus wortmani (15917 15930) Hyopsodus wortmani (15950)
Lysitean	55.8 - 50.3	USA (Wyoming)	Hyopsodus wortmani (15880 15881)
Lysitean	55.8 - 50.3	USA (Colorado)	Hyopsodus wortmani (15654)
Lysitean - Lostcabinian	55.8 - 50.3	USA (Wyoming)	Hyopsodus miticulus (15451)
Wasatchian	55.8 - 50.3	USA (Texas)	Hyopsodus wortmani, Hyopsodus miticulus (15847)
Wasatchian	55.8 - 50.3	Mexico (Baja California)	Hyopsodus sp. (15671)
Wasatchian	55.8 - 50.3	USA (Virginia)	Hyopsodus sp. (15595)
Wasatchian	55.8 - 50.3	USA (North Dakota)	Hyopsodus loomisi, Hyopsodus miticulus (15895)
Wasatchian	55.8 - 50.3	USA (California)	Hyopsodus sp. (15676 15677)
Wasatchian	55.8 - 50.3	USA (New Mexico)	Esthonyx miticulus (15430) Hyopsodus lysitensis, Hyopsodus mentalis (15432 15433 15434 15435 15436 15438) Hyopsodus lysitensis, Hyopsodus mentalis, Hyopsodus minor (15437 15663) Hyopsodus miticulus (15439 15441 15452 15607 15612) Hyopsodus powellianus (15459) Hyopsodus sp. (15446 15448)
Wasatchian	55.8 - 50.3	USA (Colorado)	Hyopsodus latidens (15278 15279 15350 15353 15356 15360 15361 15363 15365 15367 15370 15371 15372 15374 15383) Hyopsodus latidens, Hyopsodus wortmani (15377) Hyopsodus loomisi (15865) Hyopsodus loomisi, Hyopsodus miticulus (15450 15480 15485 15660 15725 15845) Hyopsodus loomisi, Hyopsodus simplex (15887) Hyopsodus miticulus (15284 15287 15290 15295 15306 15385 15389 15393 15859) Hyopsodus powellianus (15289 15311 15318 15319 15326 15327 15329 15331 15333 15338 15341 15391 15398 15408 15409 15410 15412 15420 15422) Hyopsodus powellianus, Hyopsodus miticulus (15302 15310 15325 15388) Hyopsodus powellianus, Hyopsodus wortmani (15225 15334 15414) Hyopsodus sp. (15266 15855 15861 15863 15864 15866 15868 15870) Hyopsodus sp., Hyopsodus latidens (15222 15264 15267 15273 15277 15357) Hyopsodus walcottianus (15649) Hyopsodus walcottianus, Hyopsodus wortmani (15650) Hyopsodus wortmani (15224 15226 15227 15280 15293 15294 15296 15314 15316 15321 15322 15336 15390 15394 15413 15652 15882)
Wasatchian	55.8 - 50.3	USA (Wyoming)	Condylarthra indet. (15886) Condylarthra indet., Hyopsodus mentalis, Hyopsodus powellianus (15894) Hyopsodus aemulor, Hyopsodus loomisi (46800) Hyopsodus browni (15661) Hyopsodus latidens (15573 15577 15682 15695 15696 15697 15698 15699 15702 15704 15805 46717 46720 46730 46731 46732 46733 46737 46738 46743 46749 46753 46756 46757 46759 46760 46761 46832 46834 46842 46843 46845) Hyopsodus latidens, Hyopsodus pauxillus (15705 46739 46758) Hyopsodus latidens/miticulus, Hyopsodus loomisi/minor (15551) Hyopsodus latidens/miticulus, Hyopsodus loomisi/minor, Hyopsodus powellianus (15554) Hyopsodus loomisi (15586 15587 15588 15589 15590 15592 15683 15717 15727 15759 15760 15761 15771 15779 15793 15817 15830 46636 46639 46641 46642 46643 46645 46648 46650 46656 46657 46659 46664 46667 46668 46669 46673 46675 46684 46685 46686 46687 46688 46689 46690 46693 46694 46697 46698 46721 46722 46724 46726 46727 46768 46771 46772 46773 46774 46775 46776 46777 46780 46781 46782 46784 46785 46786 46788 46789 46790 46791 46792 46793 46794 46795 46796 46797 46798 46801 46803 46804 46805 46806 46807 46808 46809 46813 46814 46816 46817 46818 46819 46821 46822 46823 46824 46830 46831 46835 46836 46838 46839 46840 46844 46852) Hyopsodus loomisi, Hyopsodus latidens (15933) Hyopsodus loomisi, Hyopsodus minor, Hyopsodus simplex (15626) Hyopsodus loomisi, Hyopsodus miticulus (15455) Hyopsodus loomisi, Hyopsodus pauxillus (46644 46672 46680) Hyopsodus loomisi, Hyopsodus powellianus, Hyopsodus latidens (15934) Hyopsodus loomisi, Hyopsodus simplex (15482 15624 15625 15627 15628 15629 15630 15631 15632 15633 15634 15635 15636 15637 15638 15639 15640 15641 15790) Hyopsodus lysitensis (46695 46705 46709 46715 46764 46810 46811 46848) Hyopsodus lysitensis, Hyopsodus mentalis (15925) Hyopsodus lysitensis, Hyopsodus mentalis, Hyopsodus minor (15924) Hyopsodus lysitensis, Hyopsodus minor (46699 46700 46701 46702 46703 46706 46710 46711 46763 46826 46849 46851) Hyopsodus lysitensis, Hyopsodus minor, Hyopsodus powellianus (15927 15928 15939 15965 15966 15967 15971 15973 15974 15977 15978 15984 15986 15987 15988 46692) Hyopsodus lysitensis, Hyopsodus powellianus (15919 15920 15921 15938 15940 15941 15968 15969 15970 15972 15975 15976 15979 15980 15981 15982 15983 15985 46712 46714) Hyopsodus mentalis (16010 16011 16012 16013) Hyopsodus mentalis, Hyopsodus walcottianus, Hyopsodus wortmani (15710) Hyopsodus mentalis, Hyopsodus wortmani (15662 16074) Hyopsodus minor (46847 186932 186933 186934 186935 186937 186938 186944 186945 186946 186949 186950 186951 186952) Hyopsodus minor, Hyopsodus miticulus (15552 15575 15576 15578 15689 46825) Hyopsodus minor, Hyopsodus powellianus (186943) Hyopsodus minor, Hyopsodus powellianus, Hyopsodus wortmani (15964) Hyopsodus minusculus, Hyopsodus paulus (15942) Hyopsodus miticulus (15456 15457 15467 15468 15469 15474 15477 15478 15688 15721 15852 15889 15912 15932 15994 16019 16020 16021 16024 16027 16028 16029 16032 16055) Hyopsodus miticulus, Hyopsodus browni (15597) Hyopsodus miticulus, Hyopsodus simplex (15692) Hyopsodus paulus (15673 118482) Hyopsodus pauxillus (46815) Hyopsodus pauxillus, Hyopsodus powellianus, Hyopsodus simplex (15707) Hyopsodus powellianus (15905 46713 186931) Hyopsodus simplex (15687 15694 15706 46682 46683 46696 46704 46707 46719 46729 46734 46735 46736 46740 46741 46745 46746 46747 46748 46750 46751 46752 46754 46755 46765 46766 46767 46820 46829) Hyopsodus sp. (15458 15491 15493 15495 15503 15504 15505 15507 15509 15510 15511 15512 15513 15514 15515 15516 15517 15518 15519 15520 15522 15523 15524 15531 15532 15533 15534 15536 15540 15541 15542 15543 15544 15545 15547 15548 15549 15550 15553 15567 15568 15579 15658 15722 15723 15729 15731 15754 15756 15788 15841 15843 15846 15884 15891 15892 15893 15902 15923 15935 15936 15937 15943 16006 16007 16039 16040 16041 16049 16050 16051 16062 27268 46691 192045 192046) Hyopsodus sp., Hyopsodus lysitensis, Hyopsodus powellianus (15559) Hyopsodus sp., Hyopsodus mentalis (15996 16014) Hyopsodus sp., Hyopsodus mentalis, Hyopsodus minor (16009) Hyopsodus sp., Hyopsodus pauxillus (15538) Hyopsodus sp., Hyopsodus powellianus (15555 15556 15557 15558 15560 15563 15564 15565 15566) Hyopsodus walcottianus (15922) Hyopsodus walcottianus, Hyopsodus wortmani (15946 15948 15955 15989) Hyopsodus walcottianus, Hyopsodus wortmani, Hyopsodus miticulus (15486) Hyopsodus wortmani (15454 15665 15724 15904 15908 15910 15913 15929 15947 15949 15951 15952 15953 15954 15956 15962 15990 15991 15992 15993 15995 16054 16056 16065 16066 16067 16077) Hyopsodus wortmani, Hyopsodus browni (15479) Hyopsodus wortmani, Hyopsodus miticulus (15714 15839 15842)
Bumbanian	55.8 - 48.6	Mongolia (Southern Mongolia)	Hyopsodus orientalis, Midiagnus gracilis (37206)
Bumbanian	55.8 - 48.6	Mongolia (Omnogov)	Hyopsodus orientalis (133397)
Early/Lower Eocene	55.8 - 48.6	China (Hubei)	Hyopsodus fangxianensis (175179)
Early/Lower Eocene	55.8 - 48.6	China (Shandong)	Asiohyopsodus confuciusi (77929 176605 183188)
Early/Lower Eocene	55.8 - 48.6	France (Var)	Hyopsodus itinerans (35538)
Early/Lower Eocene	55.8 - 48.6	USA (Wyoming)	Hyopsodus loomisi (15679 154251 154253 154254 154255 154257 154258 154262 154265 154270 154271 154276 154283 154286) Hyopsodus lysitensis, Hyopsodus minor (15684 15685) Hyopsodus lysitensis, Hyopsodus minor, Hyopsodus powellianus (15680) Hyopsodus minor, Hyopsodus miticulus (15690 15691 15693) Hyopsodus simplex (15700) Hyopsodus sp. (15681)
Early/Lower Eocene	55.8 - 48.6	France (Corbieres)	Lessnessina praecipuus (182668)
Wasatchian - Bridgerian	55.8 - 46.2	USA (Wyoming)	Hyopsodus sp. (16134 16249 16436) Hyopsodus wortmani (15669)
Early/Lower Eocene - Middle Eocene	55.8 - 37.71	Japan	Hyopsodus sp. nov. A (183077) Hyopsodus sp. nov. B (183076)
Bridgerian	50.3 - 46.2	Mexico (Guanajuato)	Hyopsodontidae indet. (16598)
Bridgerian	50.3 - 46.2	USA (Colorado)	Hyopsodus paulus (16240 16241) Hyopsodus sp. (16251) Hyopsodus wortmani (16239 16242)
Bridgerian	50.3 - 46.2	USA (Wyoming)	Hyopsodus browni (16198) Hyopsodus despiciens (16236) Hyopsodus lepidus (16169 16235 16244 16287 16418 16431 16432 16433) Hyopsodus lepidus, Hyopsodus paulus (16180) Hyopsodus marshi (16448) Hyopsodus minusculus (16079 16091 16095 16096 16097 16098 16099 16100 16101 16103 16105 16107 16108 16109 16111 16113 16114 16115 16116 16120 16129 16132 16221 16226 16227 16232 16255 16262 16276 16289 16309 16313 16315 16316 16320 16321 16325 16327 16330 16335 16343 16393 16441 16504) Hyopsodus minusculus, Hyopsodus paulus (16201 16219 16246 16254 16284 16295 16397 16408 16439 16451 16500) Hyopsodus minusculus, Hyopsodus paulus, Hyopsodus walcottianus (16207) Hyopsodus minusculus, Hyopsodus wortmani (16415) Hyopsodus paulus (16081 16137 16138 16142 16144 16145 16146 16147 16148 16149 16151 16162 16170 16183 16184 16185 16188 16189 16203 16205 16208 16216 16223 16225 16228 16243 16291 16402 16440 16443 16444 16445 16449 16484 16486 16488 16492 16494 16499 16506 16647) Hyopsodus paulus, Hyopsodus tonksi (16139) Hyopsodus sp. (15837 16084 16209 16280 16296 16372 16438 16496 16502 27269 129895) Hyopsodus sp., Hyopsodus paulus (16388) Hyopsodus sp., Hyopsodus sp. (16204) Hyopsodus tonksi (16224 16437) Hyopsodus wortmani (16247 16248)
Bridgerian	50.3 - 46.2	USA (Nevada)	Hyopsodus paulus (16218)
Bridgerian	50.3 - 46.2	USA (Utah)	Hyopsodus minusculus, Hyopsodus paulus (16396)
Bridgerian	50.3 - 46.2	USA (Texas)	Hyopsodus uintensis (16195)
MP 13 - MP 14	48.6 - 40.4	France (Quercy)	Paroxyclaenus lemuroides (36875)
Lutetian	47.8 - 41.3	France (Alsace Region)	Vulpavoides vanvaleni (31609)
Lutetian	47.8 - 41.3	Morocco	Hyopsodontidae indet. (91339)
Lutetian	47.8 - 41.3	Turkey (Anatolia)	Hilalia saribeya (183977) Hilalia saribeya, Hilalia selanneae, Hilalia robusta, Hilalia sezerorum (43624)
Lutetian	47.8 - 41.3	Germany (Sachsen-Anhalt)	Pugiodens mirus (13322 45739) Vulpavoides germanicus (45000)
Lutetian	47.8 - 41.3	Senegal (Thies region)	Condylarthra indet. (36616)
Middle Eocene	47.8 - 37.71	China (Xinjiang)	Hyopsodus huashigouensis (176332)
Early/Lower Uintan	46.2 - 39.7	USA (Colorado)	Hyopsodus despiciens (16623)
Early/Lower Uintan	46.2 - 39.7	USA (Texas)	Hyopsodus paulus (16586) Hyopsodus paulus, Hyopsodus uintensis (16651)
Early/Lower Uintan	46.2 - 39.7	USA (Wyoming)	Hyopsodus lepidus (93087)
Late/Upper Uintan	46.2 - 39.7	USA (Texas)	Hyopsodus paulus, Hyopsodus uintensis (16614) Hyopsodus uintensis (16542)
Late/Upper Uintan	46.2 - 39.7	Canada (Saskatchewan)	Hyopsodus fastigatus (16626)
Uintan	46.2 - 39.7	USA (Texas)	Hyopsodus paulus (16587 16588) Hyopsodus uintensis (16624)
Uintan	46.2 - 39.7	USA (Utah)	Hyopsodus uintensis (16589 16654) Hyposodontidae indet. (213066)
Uintan	46.2 - 39.7	USA (Montana)	Hyopsodus sp. (222922)
Uintan	46.2 - 39.7	USA (Wyoming)	Hyopsodus paulus (16563 16576 16577) Hyopsodus sp. (16521 16641 16648 202166) Hyopsodus uintensis (16513 16514 16515 16516 16517 16518 16630 16642 16643 16645)
Bartonian	41.3 - 38.0	United Kingdom (England)	Vulpavoides cooperi (39014)
Bartonian	41.3 - 38.0	Switzerland (Baselland)	Vulpavoides simplicidens (182806)
Duchesnean	39.7 - 37.0	USA (Montana)	Hyopsodus sp. (16854)
Late/Upper Eocene	37.2 - 33.9	United Kingdom (England)	Dyspterna hopwoodi (40030)
MP 18 - MP 19	37.2 - 33.9	France	Paroxyclaenus lemuroides (184091)