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Actinopteri

Osteichthyes

Taxonomy
Actinopteri was named by Cope (1871). It is extant.

It was reranked as the superorder Actinopterygia by Cope (1887); it was reranked as the superorder Actinopteri by Hay (1902); it was replaced with Actinopterygii by Goodrich (1930); it was reranked as the unranked clade Actinopteri by Lauder and Liem (1983) and Patterson (1983); it was reranked as the class Actinopteri by Betancur et al. (2013); it was reranked as the infraclass Actinopteri by Grande et al. (2002), Grande and Hilton (2006), van der Laan (2019).

It was assigned to Teleostomi by Cope (1887); to Teleostomi by Hay (1902); to Pisces by Cope (1871), Jordan (1963); to Actinopterygii by Lauder and Liem (1983), Patterson (1983), Cartanyà (1999); to Actinopterygii by Grande et al. (2002), Grande and Hilton (2006); to Actinopterygii by Betancur et al. (2013); and to Actinopterygii by van der Laan (2019).

Synonyms
  • Palaeophoxinus was named by Coates (1988). It is not extant.

    It was considered a nomen nudum by Clark (1988).
Synonymy list
YearName and author
1871Actinopteri Cope p. 449
1887Actinopterygia Cope p. 1017
1902Actinopteri Hay p. 364
1963Actinopteri Jordan p. 611
1983Actinopteri Lauder and Liem p. 103
1983Actinopteri Patterson
1988Palaeophoxinus Coates
1999Actinopteri Cartanyà p. 540
2002Actinopteri Grande et al.
2006Actinopteri Grande and Hilton p. 5
2013Actinopteri Betancur et al.
2019Actinopteri van der Laan p. 51

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
RankNameAuthor
superclassGnathostomata
classOsteichthyes
subclassActinopterygii()
infraclassActinopteri()
infraclassActinopteri()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Infracl. Actinopteri Cope 1871
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Or. †Aeduelliformes Heyler 1969
+
Fm. †Aeduellidae Romer 1945
+
G. †Aeduella Westoll 1937
+
†Aeduella blainvillei Agassiz 1833
+
Invalid names: Amblypterus angustus Agassiz 1833 [synonym], Palaeoniscus landrioti Sauvage 1891 [synonym], Palaeoniscus lepidurus Agassiz 1834 [synonym], Palaeoniscus voltzii Agassiz 1833 [synonym], Palaeothrissum inaequilobum Blainville 1818 [synonym], Palaeothrissum parvum Blainville 1818 [synonym]
G. †Amelangia Štamberg and Werneburg 2023
+
†Amelangia ornata Štamberg and Werneburg 2023
G. †Bourbonnella Heyler 1967
+
†Bourbonnella fourrieri Poplin 2001
†Bourbonnella guilloti Heyler 1967
†Bourbonnella hirsuta Štamberg 2007
†Bourbonnella jocelynae Mickle 2011
†Bourbonnella sottyi Anonymous 1972
G. †Neslovicella Štamberg 2007
+
†Neslovicella elongata Štamberg 2010
†Neslovicella rzehaki Štamberg 2007
G. †Platysella Heyler and Poplin 1983
+
†Platysella descusi Heyler and Poplin 1983
†Platysella lallyi Heyler and Poplin 1983
†Platysella poplinae Heyler 2002
G. †Puertollanichthys Forey and Young 1985
+
†Puertollanichthys ritchiei Forey and Young 1985
G. †Shafeevus Yankevich 1998
+
†Shafeevus sulcatus Yankevich 1998
G. †Spinarichthys Štamberg 1986
+
†Spinarichthys dispersus Fritsch 1895
+
Fm. †Amphicentridae Young 1866
+
G. †Adroichthys Gardiner 1969
+
†Adroichthys tuberculatus Gardiner 1969
G. †Amphicentrum Young 1866
+
†Amphicentrum crassum Traquair 1890
+
Invalid names: Cheirodus granulosum Traquair 1879 [synonym]
†Amphicentrum jurgenai Zidek 1992
†Amphicentrum orbiculare Newberry and Worthen 1870
†Amphicentrum striatum Hancock and Atthey 1872
+
Invalid names: Cheirodus striatus Traquair 1879 [objective synonym]
G. †Cheirodopsis Traquair 1881
+
†Cheirodopsis geikiei Traquair 1881
G. †Chirodus M'Coy 1848
+
†Chirodus acutus Newberry 1856
Invalid names: Chirodus pesranae M'Coy 1848 [nomen nudum]
G. †Eurynothus Agassiz 1843
G. †Proteurynotus Moy-Thomas and Bradley Dyne 1938
+
†Proteurynotus traquairi Moy-Thomas and Bradley Dyne 1938
G. †Tompoichthys Obruchev 1964
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†Tompoichthys abramovi Obruchev 1964
Fm. †Styracopteridae Moy-Thomas 1937
+
G. †Benedenius Traquair 1877
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†Benedenius deneensis van Beneden 1871
†Benedenius soreili Fraipont 1890
Invalid names: Benedenius deneensis van Beneden 1871 [valid], Benedenius soreili Fraipont 1890 [valid]
G. †Benedenius Traquair 1877
+
†Benedenius deneensis van Beneden 1871
†Benedenius soreili Fraipont 1890
Invalid names: Benedenius deneensis van Beneden 1871 [valid], Benedenius soreili Fraipont 1890 [valid]
G. †Chichia Liu and Wang 1978
+
†Chichia gracilis Liu and Wang 1978
G. †Fouldenia White 1927
+
†Fouldenia ischiptera Traquair 1881
†Fouldenia striatulus Schultze et al. 2021
G. †Styracopterus Traquair 1890
+
†Styracopterus fulcratus Traquair 1881
Invalid names: Chichiidae Liu and Wang 1978 [synonym]
Fm. †Teleopterinidae Berg 1936
+
G. †Eurylepis Newberry 1856
+
†Eurylepis granulata Newberry 1856
†Eurylepis tuberculata Newberry 1856
G. †Teleopterina Berg 1936
G. †Anaglyphus Rzehak 1881
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Invalid names: Anaglyphus insignis Rzehak 1881 [nomen nudum]
G. †Angatubichthys De Figueriredo and Carvalho 2004
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†Angatubichthys mendesi De Figueriredo and Carvalho 2004
G. †Bethesdaichthys Bender 2001
+
G. †Blourugia Bender 2006
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†Blourugia seeleyi Woodward 1893
Fm. †Dwykiidae Gardiner 1969
+
G. †Dwykia Gardiner 1969
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†Dwykia analensis Gardiner 1969
G. †Frederichthys Coates 1993
+
Fm. †Kenderlichthyidae Kazantseva-Selezneva 1977
+
G. †Kenderlichthys Kazantseva-Selezneva 1977
+
†Kenderlichthys paluster Kazantseva-Selezneva 1980
†Kenderlichthys tuberculatus Kazantseva-Selezneva 1977
G. †Kompasia Bender 2002
+
†Kompasia delaharpei Bender 2002
G. †Lanarkichthys Elliott 2018
+
†Lanarkichthys gardineri Elliott 2018
G. †Limnomis Daeschler 2000
+
†Limnomis delaneyi Daeschler 2000
G. †Melanecta Coates 1998
+
†Melanecta anneae Coates 1998
G. †Moythomasia Gross 1950
+
†Moythomasia antiqua Gross 1953
†Moythomasia devonica Clarke 1885
†Moythomasia durgaringa Gardiner and Bartram 1977
†Moythomasia laevigata Gross 1953
†Moythomasia lebedevi Plax et al. 2025
†Moythomasia lineata Choo 2015
†Moythomasia nitida Gross 1953
†Moythomasia perforata Gross 1942
†Moythomasia striata Gross 1953
Invalid names: Aldingeria Gross 1942 [replaced]
Fm. †Nematoptychiidae Kazantseva-Selezneva 1977
+
G. †Nematoptychius Traquair 1875
+
†Nematoptychius greenocki Traquair 1866
Fm. †Palaeobergiidae Kazantseva 1968
+
G. †Palaeobergia Berg 1958
+
Or. †Paphosisciformes Grogan and Lund 2015
+
Fm. †Paphosiscidae Grogan and Lund 2015
+
G. †Paphosiscus Grogan and Lund 2015
+
†Paphosiscus circulocaudus Grogan and Lund 2015
†Paphosiscus scalmocristus Grogan and Lund 2015
G. †Pseudohaplolepis Elliott 2018
+
Or. †Scanilepidiformes Sytchevskaya 1999
+
G. †Beishanichthys Xu and Gao 2011
+
†Beishanichthys brevicaudalis Xu and Gao 2011
Fm. †Evenkiidae Selezneva 1985
+
G. †Evenkia Berg 1941
+
†Evenkia eunotoptera Berg 1941
G. †Oshiaichthys Schultze et al. 2021
+
†Oshiaichthys ferganica Schultze et al. 2021
+
Invalid names: Oshia ferganica Sytchevskaya 1999 [replaced]
Invalid names: Oshia Sytchevskaya 1999 [replaced]
G. †Toyemia Minikh 1990
+
†Toyemia blumentalis Minikh 1995
†Toyemia tverdochlebovi Minikh 1990
+
G. †Paratarrasius Lund and Melton Jr. 1982
+
†Paratarrasius hibbardi Lund and Melton Jr. 1982
Invalid names: Apholidotos Lund 1991 [nomen dubium]
Unr. Teleosteomorpha Arratia et al. 2004
+
+
+
Invalid names: Batrachoidida [empty]
+
Or. Beryciformes Patterson 1964
+
G. †Africentrum White and Moy-Thomas 1941
Fm. Berycidae Lowe 1843
Subor. Berycoidei Patterson 1964
G. †Berycomorus Arambourg 1967
G. †Brazosiella Frizzell and Dante 1965
G. †Cryptoberyx Gaudant 1978
Subor. †Dinopterygoidei Patterson 1964
G. †Holocentrites Conrad 1941
Superfm. †Holocentroidea Richardson 1846
Subor. †Holocentroidei Johnson and Patterson 1993
G. Kryptophanaron Silvester and Fowler 1927
G. †Omosoma Costa 1857
G. †Pattersonoberyx Gayet 1980
G. Plectrypops Gill 1863
G. †Plesioberyx Gayet 1980
G. Poromitra Goode and Bean 1883
G. †Pseudholocentrum Arambourg 1967
Fm. †Pseudomonocentrididae González-Rodríguez et al. 2013
G. Sargocentron Fowler 1904
Subor. †Trachichthyoidei Parr 1933
Invalid names: Anoplogastridae [empty], Beryx dalmaticus Steindachner 1863 [nomen dubium], Beryx subovatus Bassani 1882 [nomen dubium], Polymixioidei Patterson 1964 [empty]
+
Fm. Trachichthyidae Bleeker 1859
+
Invalid names: Hoplopterygidae [synonym]
G. †Caprosimilis Bienkowska-Wasiluk and Bonde 2015
+
†Caprosimilis carpathicus Bienkowska-Wasiluk and Bonde 2015
Or. Cataphracti Jordan 1963
+
+
Invalid names: Abyssocottidae [empty], Ascelichthyidae [empty], Aspidophoroididae [empty], Blepsiidae [empty], Comephoridae [empty], Cottocomerphoridae [empty], Hemitripteridae [empty], Icelidae [empty], Neophrynichthyidae [empty], Synchridae [empty]
+
Invalid names: Liparopidae [empty]
+
Invalid names: Anoplopomidae [empty], Erilepidae [empty], Oxylebiidae [empty]
Invalid names: Platycephaliformes Jordan 1963 [empty], Trigliformes [empty]
G. †Choichix Cantalice et al. 2021
+
†Choichix alvaradoi Cantalice et al. 2021
G. †Gigapteryx Forey et al. 2003
+
†Gigapteryx tethyestris Forey et al. 2003
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Invalid names: Hysterocarpidae [empty]
Unr. Holocentrimorphaceae Betancur et al. 2013
+
Or. Holocentriformes Betancur et al. 2013
+
Fm. †Caproberycidae Patterson 1967
Fm. Holocentridae Richardson 1846
Fm. †Tenuicentridae Gayet 1983
+
Invalid names: Helostomidae [empty], Luciocephalidae [empty], Ophicephalidae [empty], Polyacanthidae [empty]
G. †Pepemkay Alvarado-Ortega and Than-Marchese 2013
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†Pepemkay maya Alvarado-Ortega and Than-Marchese 2013
+
Unr. †Gobiida Lin et al. 2015
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†Gobiida bicornuta Lin et al. 2015
†Gobiida brioche Lin et al. 2015
G. †Kelemejtubus Cantalice and Alvarado-Ortega 2017
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†Kelemejtubus castroi Cantalice and Alvarado-Ortega 2017
Invalid names: Ophidiida [empty], Pelagia Miya et al. 2013 [empty]
Unr. Percomorphaceae Betancur et al. 2013
+
+
Or. Batrachoidiformes [toadfish]
+
Fm. Centropomidae Poey 1868
Fm. Lactariidae Boulenger 1904
Fm. Menidae Gill 1885 [moonfish]
Or. Pleuronectiformes [flatfish]
Fm. Polynemidae Rafinesque 1815
Fm. †Pygaeidae Jordan 1905
Fm. Sphyraenidae Rafinesque 1815 [barracuda]
Fm. Toxotidae Bleeker 1859
+
Invalid names: Anabantomorphariae [empty], Carangimorphariae [empty], Ovalentariae [empty]
Unr. Eupercaria Betancur 2013
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Fm. †Asianthidae Sytchevskaya and Prokofiev 2003
Fm. †Caucasichthyidae Bannikov et al. 2011
Fm. Emmelichthyidae Poey 1867
Fm. †Eotrigonodontidae White 1935
Fm. †Exelliidae Blot 1969
+
Invalid names: Semiophoridae [synonym]
Or. Lophiiformes [anglerfish]
Fm. Malacanthidae Poey 1861
Fm. Monodactylidae Jordan and Evermann 1898
Fm. Moronidae [temperate bass]
Fm. Parascorpididae Smith 1949
Or. Perciformes Bleeker 1859 [perch-like fish]
Fm. †Pietschellidae Carnevale and Bannikov 2015
Or. Priacanthiformes Bleeker 1871
Fm. †Quasimullidae Bannikov 1999
Fm. †Repropcidae Bannikov 1991
Fm. Scatophagidae Gill 1883
Fm. Siganidae Richardson 1837
Fm. Sillaginidae Richardson 1846
Or. Tetraodontiformes Berg 1940
Invalid names: Callanthiidae [empty], Centrogenyidae [empty], Dinolestidae [empty], Dinopercidae Heemstra and Hecht 1986 [empty], Pomacanthidae [empty]
+
Or. Ophidiiformes Berg 1937
Unr. Ovalentaria Smith and Near 2012
+
Superor. Atherinomorphae Greenwood et al. 1966
Superor. Cichlomorphae
G. †Cylindracanthus Leidy 1856
+
Invalid names: Coelorhynchus Agassiz 1844 [replaced]
Fm. Embiotocidae Agassiz 1853
Fm. Grammatidae Jordan 1887
Superor. Mugilomorphae
Fm. Pomacentridae Bonaparte 1831
Fm. Pseudochromidae Mueller and Troschel 1849
Invalid names: Congrogadidae [empty], Moroniformes [empty], Opisthognathidae [empty], Plesiopidae [empty], Polycentridae [empty], Scombrolabraciformes [empty]
+
Fm. Acropomatidae Gill 1891
Fm. Sciaenidae Cuvier 1828
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Invalid names: Scienoides [synonym]
Invalid names: Caesionidae [empty], Cirrhitiformes [empty], Lateolabracidae [empty]
Invalid names: Batrachoidimorpharia [empty], Gobiomorpharia [empty], Ophidiimorpharia [empty], Scombrimorpharia [empty]
+
Invalid names: Hemibranchii [empty], Lophobranchi Cuvier [empty]
Invalid names: Aulostomi Cuvier [empty], Cetomimiformes [empty], Euacanthopterygii Johnson and Patterson 1993 [empty], Hypostomides [empty], Xenoberyces [empty]
Unr. †Aspidorhynchei Nelson et al. 2016
+
Or. †Aspidorhynchiformes Bleeker 1859
+
Fm. †Aspidorhynchidae Bleeker 1859
+
G. †Aspidorhynchus Agassiz 1833
G. †Belonostomus Agassiz 1834
+
Invalid names: Diphyodus Lambe 1902 [synonym]
G. †Richmondichthys Bartholomai 2004
G. †Vinctifer
G. †Atacamichthys Arratia and Schultze 1987
+
†Atacamichthys greeni Arratia and Schultze 1987
G. †Pseudovinctifer Arratia 2015
+
†Pseudovinctifer chilensis Arratia 2015
Or. †Pachycormiformes Berg 1940
+
G. †Domeykos
+
Invalid names: Megalopterus Kner 1866 [replaced]
G. †Hengnania
G. †Ichthyokentema Davies 1887
+
G. †Koonwarria Waldman 1971
+
†Koonwarria manifrons Waldman 1971
G. †Oreochima
Fm. †Pachycormidae Woodward 1895
+
Subfm. †Asthenocorminae Cooper et al. 2022
G. †Eugnathides Gregory 1923
G. †Euthynotus Agassiz 1843
+
Invalid names: Parathrissops Sauvage 1891 [synonym]
Subfm. †Hypsocorminae Vetter 1881
G. †Sauropsis Agassiz 1832
Invalid names: Protosphyraenidae Cope 1892 [synonym]
G. †Prymnetes
G. †Seefeldia
G. †Wadeichthys Waldman 1971
+
†Wadeichthys oxyops Waldman 1971
Or. †Prohaleciteiformes Arratia 2017
+
Fm. †Prohaleciteidae Arratia 2017
+
Fm. †Barschichthyidae Arratia and Schultze 2024
+
G. †Barschichthys Arratia and Schultze 2024
+
†Barschichthys ruedersdorfensis Arratia and Schultze 2024
Unr. Lampripterygii Betancur-R et al. 2017
+
Fm. †Aipichthyidae Patterson 1964
+
G. †Aipichthys Steindachner 1860
+
†Aipichthys minor Pictet 1850
+
Invalid names: Platax brevis Davis 1887 [synonym]
†Aipichthys nuchalis Dixon 1850
†Aipichthys oblongus Gayet 1980
†Aipichthys pretiosus Steindachner 1860
†Aipichthys velifer Woodward 1901
G. †Paraipichthys Gaudant 1978
Fm. †Aipichthyoididae Gayet 1980
+
G. †Aipichthyoides Gayet 1980
+
G. †Zoqueichthys Alvarado-Ortega and Than-Marchese 2012
+
†Zoqueichthys carolinae Alvarado-Ortega and Than-Marchese 2012
+
Fm. †Lampridae
+
G. Lampris Retzius 1799
+
Invalid names: Dasceles Jordan 1925 [synonym]
+
G. †Babelichthys Davesne 2017
G. Lophotus Giorna 1809
G. †Oligolophotes Bannikov 1999
Fm. †Palaeocentrotidae Bonde 1966
+
G. †Natgeosocus Bannikov 2014
G. †Palaeocentrotus Kühne 1941
Fm. †Veliferidae Bleeker 1859
+
G. Velifer Temminck and Schlegel 1850
G. †Veronavelifer Bannikov 1990
G. †Wettonius Carnevale and Bannikov 2018
Invalid names: Radiicephalidae [empty]
Fm. †Pharmacichthyidae Patterson 1964
+
G. †Pharmacichthys Woodward 1942
+
†Pharmacichthys venenifer Woodward 1942
Fm. †Pycnosteroididae Patterson 1964
+
G. †Magrebichthys Murray and Wilson 2014
+
†Magrebichthys nelsoni Murray and Wilson 2014
G. †Pycnosteroides Woodward 1942
+
+
Invalid names: Aipichthys formosus Hay 1903 [synonym]
Fm. †Marcopoloichthyidae Tintori et al. 2008
+
G. †Marcopoloichthys Tintori et al. 2008
+
†Marcopoloichthys andreetti Tintori et al. 2008
†Marcopoloichthys ani Tintori et al. 2008
†Marcopoloichthys faccii Gortani 1907
†Marcopoloichthys furreri Arratia 2022
†Marcopoloichthys mirigioliensis Arratia et al. 2024
Fm. †Palaeolabridae Estes 1969
+
G. †Palaeolabrus Casier 1967
+
Unr. Paracanthopterygii Greenwood et al. 1966
+
+
G. †Macruridarum Koken
+
†Macruridarum acuminatus Weiler 1943
†Macruridarum ovalis Weiler 1943
Fm. †Mcconichthyidae Grande 1988
+
G. †Mcconichthys Grande 1988
Invalid names: Eretmophoridae [empty], Macrouroididae [empty], Ophidioidea [empty]
Or. Gadiformes Goodrich 1909
+
Fm. Bathygadidae Jordan and Evermann 1898
+
G. Bathygadus Guenther 1878
G. Gadomus Regan 1903
G. †Paleobathygadus David 1956
G. †Probathygadus David 1956
G. Brosme Rafinesque 1815
Fm. †Euclichthyidae Cohen 1984
+
G. †Archaemacruroides Stinton 1965
G. Euclichthys McCulloch 1926
Subor. Gadoidei
+
Fm. Gadidae Rafinesque 1810
Fm. Lotidae Bonaparte 1842
Fm. Merlucciidae Gill 1884
Fm. Gaidropsaridae Jordan and Evermann 1898
+
G. †Enchelyopus Agassiz 1835
G. †Onogadus Buen 1934
G. Lota Oken 1817
Fm. Macrouridae Jordan and Evermann 1898 [grenadiers]
+
G. †Amblygoniolepidus David 1956
G. †Bolbocara Jordan 1927
G. †Calilepidus David 1956
G. Coelorhynchus Giorna 1809
G. Coelorinchus Giorna 1809
G. Coryphaenoides Gunnerus 1765
G. †Homeocoryphaenoides David 1956
G. †Homeomacrurus David 1956
G. †Homeonezumia David 1956
G. Hymenocephalus Giglioli 1884
G. Lepidorhynchus Richardson 1846
G. †Leptacantholepidus David 1956
Subfm. †Macrourinae Bonaparte 1832
G. †Macrourus Bloch 1786
G. Nezumia Jordan and Starks 1904
G. †Oxygoniolepidus David 1956
G. †Promacrurus David 1956
G. †Pyknolepidus David 1956
G. †Razelainia Carnevale 2007
G. Trachyrincus Giorna 1809
Invalid names: Trachyrincinae Günther 1870 [empty]
+
G. †Macruroides Smith and Radcliffe 1912
Fm. †Macruronidae Regan 1903
+
G. Macruronus Guenther 1873
G. †Maorigadus Schwarzhans 1980
+
†Maorigadus anarchicus Schwarzhans 1980
+
Invalid names: Maorigadus delicatulus Schwarzhans 1980 [synonym]
†Maorigadus ukrainicus Schwarzhans and Bratischko 2011
+
Fm. Melanonidae Goode and Bean 1896
G. †Pollachius Nilsson 1832
G. †Pseudoraniceps Fedotov 1974
+
G. Raniceps Oken 1817
G. †Rankinian David 1946
G. Squalogadus Gilbert and Hubbs 1916
G. Trachyrhynchus Bonaparte 1841
+
Invalid names: Merluccidae [empty], Muraenolepididae [empty]
Or. Percopsiformes Berg 1940
+
Fm. Percopsidae Regan 1911
+
G. †Amphiplaga Cope 1877
G. †Erismatopterus Cope 1871
G. †Lateopisciculus Murray and Wilson 1996
G. †Libotonius Wilson 1977
G. †Massamorichthys Murray 1996
G. Percopsis Agassiz 1849
G. †Percopsiformorum Berg 1940
+
†Percopsiformorum enigmaticus Schwarzhans 2003
†Percopsiformorum resonus Schwarzhans 2004
Invalid names: Amblyopsidae [empty], Aphredoderidae [empty]
Fm. †Protosyngnathidae Boulenger 1902
+
G. †Protosyngnathus von der Marck 1878
Or. †Sphenocephaliformes Rosen and Patterson 1969
+
Fm. †Sphenocephalidae Patterson 1964
+
G. †Sphenocephalus Agassiz 1839
G. †Xenyllion Wilson and Murray 1996
+
Invalid names: Stylephoridae [empty]
Fm. †Trebicianiidae Sorbini and Bannikov 1996
+
G. †Trebiciania Sorbini and Bannikov 1996
Or. Zeiformes Regan 1909
+
Subor. †Cyttoidei
+
Fm. †Archaeozeidae Tyler and Santini 2005
Fm. †Cretazeidae Tyler et al. 2000
Fm. Cyttidae Günther 1860
Fm. †Protozeidae Tyler and Santini 2005
G. †Cyttoides Wettstein 1886
G. †Palaeocyttus Gaudant 1978
G. †Zeiformorum Unknown
+
†Zeiformorum janni Schwarzhans 2003
Fm. †Zeniontidae Myers 1960
+
G. Capromimus Gill 1893
G. Zenion Jordan and Evermann 1896
Subor. Zeoidei
+
Fm. Parazenidae McAllister 1968
+
Invalid names: Isozenidae Schwarzhans 1996 [synonym]
Fm. Zeidae Rafinesque 1815
Invalid names: Grammicolepididae [empty], Oreosomatidae [empty]
Unr. Polymixiipterygii Betancur-R et al. 2017
+
Fm. Polymixiidae Gill 1862
+
G. †Berycopsia Radovcic 1975
G. †Berycopsis Agassiz 1850
+
Berycopsis elegans Dixon 1850
†Berycopsis germanus Agassiz 1837
+
Invalid names: Platycormus oblongus von der Marck 1863 [synonym]
†Berycopsis paucoradius Dietze 2009
†Berycopsis pulcher Bannikov and Bacchia 2005
Invalid names: Platycormus von der Marck 1858 [synonym]
G. †Dalmatichthys Radovcic 1975
G. †Homonotichthys Whitley 1933
+
Invalid names: Homonotus Agassiz 1850 [replaced], Stenostoma Agassiz 1850 [replaced]
G. †Omosomopsis Gaudant 1978
G. †Parapolymixia David 1946
+
G. Polymixia Lowe 1836
G. †Pycnosterinx Heckel 1849
+
†Pycnosterinx discoides Heckel 1849
†Pycnosterinx dorsalis Pictet 1850
†Pycnosterinx dubius Davis 1887
†Pycnosterinx elongatus Pictet and Humbert 1866
†Pycnosterinx gracilis Davis 1887
†Pycnosterinx latus Davis 1887
†Pycnosterinx niger Costa 1857
†Pycnosterinx russeggerii Heckel 1849
+
Invalid names: Imogaster auratus Costa 1857 [synonym]
Invalid names: Imogaster Costa 1857 [synonym]
Invalid names: Omosomopsidae Gaudant 1978 [synonym]
Or. Polymixiiformes [beardfish]
+
Fm. †Boreiohydriidae Murray and Cumbaa 2013
+
G. †Boreiohydrias Murray and Cumbaa 2013
G. †Cumbaaichthys Murray 2016 [beardfish]
+
†Cumbaaichthys oxyrhynchus Murray 2016 [beardfish]
Fm. †Pseudoberycidae Berg 1940
+
G. †Pseudoberyx Pictet and Humbert 1866
+
†Pseudoberyx bottae Pictet and Humbert 1866
†Pseudoberyx grandis Davis 1887
†Pseudoberyx syriacus Pictet and Humbert 1866
G. †Ruedersdorfia Arratia and Schultze 2024
+
†Ruedersdorfia berlinensis Arratia and Schultze 2024
Unr. Teleostei Müller 1846
+
+
Invalid names: Ctenoidei Owen 1859 [empty], Cycloidei Owen 1859 [empty]
G. †Acregoliath Richter 1989
+
†Acregoliath rancii Richter 1989
+
Fm. †Araripichthyidae Santos 1983
+
G. †Araripichthys Silva Santos 1985
G. †Asiatolepis Takai 1943
+
†Asiatolepis sankeyushuensis Ma and Sun 1988
Fm. †Chongichthyidae Arratia 1982
+
G. †Chongichthys Arratia 1982
+
†Chongichthys dentatus Arratia 1982
Unr. Clupeocephala Patterson and Rosen 1977
+
G. †Beurlenichthys de Figueiredo and Gallo 2004
Superor. Clupeomorpha Greenwood et al. 1966
+
Or. †Ellimmichthyiformes Grande 1982
G. †Leufuichthys Gallo et al. 2011
G. †Ornategulum Forey 1973
G. †Ranulfoichthys Alvarado-Ortega 2014
G. †Scombroclupea Kner 1863
Unr. Euteleostei Rosen 1973
+
G. †Avitosmerus Fielitz 2002
Fm. †Protostomiidae Arambourg 1943
G. †Santanasalmo Gallo et al. 2009
Invalid names: Lampridiomorpha [empty], Lamprimorpha [empty], Osmeromorpha [empty], Scopelomorpha [empty]
+
G. †Erichalcis Forey 1975
Unr. Neoteleostei Rosen 1973
Unr. Protacanthopterygii Greenwood et al. 1966
Fm. †Xeneichthyidae Arratia and González-Rodríguez 2024
Invalid names: Erichalcidae [empty]
Fm. †Lycoclupeidae Gowda 1967
+
G. †Lycoclupea Gowda 1967
Unr. †Otocephala
+
G. Otariophysi Rosen and Greenwood 1970
+
Superor. Alepocephali
Fm. †Clupavidae Bertin and Arambourg 1958
Superor. Clupei
Fm. †Ganolytidae Jordan 1921
Superor. Ostariophysi Jordan and Evermann 1896
Invalid names: Ostarioclupeomorpha [synonym]
Or. †Crossognathiformes Taverne 1989
+
G. †Bavarichthys Arratia and Tischlinger 2010
+
†Bavarichthys incognitus Arratia and Tischlinger 2010
Subor. †Crossognathoidei Taverne 1989
+
G. †Luxilites
Subor. †Pachyrhizodontoidei Forey 1977
+
G. †Michin Alvarado-Ortega et al. 2008
Fm. †Notelopidae Forey 1977
Fm. †Pachyrhizodontidae Cope 1872
+
Invalid names: Raphiosauridae Fowler 1911 [synonym]
G. †Pachyrhizodus Agassiz 1850
Fm. †Salminopsidae Gayet 1985
+
G. †Salminops Gayet 1985
+
Or. †Dorsetichthyiformes Nelson et al. 2016
+
Fm. †Dorsetichthyidae Nelson et al. 2016
+
G. †Dorsetichthys Arratia 2013
+
Invalid names: Elopocephala [empty]
Unr. Elopomorpha Greenwood et al. 1966
+
Or. Albuliformes Greenwood et al. 1966 [bonefish]
+
Fm. Albulidae Bleeker 1859
+
Invalid names: Pterothrissidae Gill 1893 [synonym]
G. †Baugeichthys Filleul 2000
G. †Genartina Frizzell and Dante 1965
G. †Marathonichthys Bartholomai 2013
Invalid names: Notacanthoidei [empty]
Or. Anguilliformes Greenwood et al. 1966 [eel]
+
G. †Abisaadia Belouze et al. 2003
Fm. †Anguillavidae Hay 1903
+
Invalid names: Encheliidae Hay 1903 [synonym], Mylomyridae Berg 1940 [synonym]
Fm. Anguillidae Rafinesque 1810
G. †Bolcanguilla Blot 1981
G. †Brannerion Jordan 1920
G. †Casierius Estes 1969
G. Congermuraena Kaup 1856
Fm. Congridae Kaup 1856 [conger eel]
Superfm. Congroidea
Subor. Congroidei Regan 1909
Fm. †Derrhiidae Jordan 1925
G. †Eomuraena Casier 1967
G. Eomyrophis Whitley 1950
G. †Gazolapodus Blot 1981
G. †Hayenchelys Belouze et al. 2003
G. Hoplunnis Kaup 1860
G. Japonoconger Asano 1958
G. Lemkea Kotthaus 1968
G. Leptocephalus Gronow 1763
Fm. †Libanechelyidae Taverne 2004
G. †Luenchelys Belouze et al. 2003
G. †Micromyrus Casier 1967
G. †Muraenanguilla Schwarzhans 2019
+
Invalid names: Heteromyridae [synonym]
Subor. †Muraenoidei
G. †Palaeomyrus Casier 1967
G. †Proserrivomer Arambourg 1967
G. †Rhynchocymba Jordan and Hubbs
Subor. Saccopharyngoidei Nelson et al. 2016
Fm. †Urenchelyidae Hay 1903
Invalid names: Apodes Latreille 1804 [synonym], Eurypharyngidae [empty], Saccopharyngidae [empty]
Or. Elopiformes Sauvage 1875
+
Subor. Albuloidei
G. †Antofagastaichthys Arratia 1986
G. †Carsothrissops D'Erasmo 1946
G. †Coryphaenopsis Frič 1896
G. †Dactylopogon von der Marck 1868
G. †Davichthys Forey 1973
G. †Echinelops Murray and Hoşgör 2012
G. †Eichstaettia Arratia 1987
Fm. Elopidae Bonaparte 1846
Subor. Elopoidei
G. †Epaelops Alves et al. 2019
G. †Flindersichthys Longman 1932
G. †Lyrolepis Romanowski 1886
Fm. Megalopidae Jordan 1923
+
Invalid names: Sedenhorstiidae Goody 1969 [synonym]
G. †Ostariostoma Schaeffer 1949
Fm. †Phyllodontidae Sauvage 1875
Or. Megalopiformes [tarpon]
+
G. †Broweria de Beaufort 1926
G. †Naiathaelon Poyato-Ariza and Wenz 1993
+
†Naiathaelon okkidion Poyato-Ariza and Wenz 1993
Invalid names: Saccopharygiformes [empty], Saccopharyngiformes [empty]
Unr. Euteleosteii Greenwood et al. 1967
+
G. †Gaudryella Patterson 1970
+
†Gaudryella gaudryi Pictet and Humbert 1866
Invalid names: Ateleopodomorpha [empty], Cyclosqamata [empty], Lampriomorpha [empty], Stenopterygii [empty]
+
G. †Gasterorhamphosus Sorbini 1981
+
Invalid names: Aulorhynchoidea Pietsch 1978 [empty], Gasterosteoidea Pietsch 1978 [empty]
G. Nerophis Rafinesque 1810
G. †Protorhamphosus Daniltshenko 1968
G. †Urosphenopsis Daniltshenko 1968
+
Invalid names: Gobioididae [empty], Periophthalmidae [empty], Psammichthyidae [empty], Trypauchenidae [empty]
Or. †Ichthyodectiformes Bardack and Sprinkle 1969
+
G. †Aidachar Nessov 1981
+
†Aidachar paludalis Nessov 1981
Fm. †Allothrissopidae Patterson and Rosen 1977
+
G. †Allothrissops Nybelin 1964
G. †Occithrissops Schaeffer and Patterson 1984
G. †Thrissops Agassiz 1833
G. †Unamichthys Alvarado-Ortega 2004
Invalid names: Unamichthyidae Alvarado-Ortega 2004 [synonym]
G. †Antarctithrissops Arratia et al. 2004
+
†Antarctithrissops australis Arratia et al. 2004
G. †Ascalabothrissops Arratia 2000
+
†Ascalabothrissops voelkli Arratia 2000
G. †Cavenderichthys Bean 2006
G. †Chuhsiungichthys Yabumoto 1994
+
†Chuhsiungichthys japonicus Yabumoto 1994
†Chuhsiungichthys yanagidai Yabumoto 1994
Fm. †Cladocyclidae Maisey 1991
+
G. †Chiromystus Cope 1885
G. †Cladocyclus Agassiz 1843
G. †Eubiodectes Hay 1903
G. †Cooyoo Lees and Bartholomai 1987
+
†Cooyoo australis Woodward 1894
G. †Garganoichthys Taverne 2009
+
†Garganoichthys decosmoi Taverne 2009
Subor. †Ichthyodectoidei Romer 1966
+
Fm. †Bardackichthyidae Hacker and Shimada 2021
Fm. †Ichthyodectidae Crook 1892
Fm. †Saurodontidae Stewart
G. †Mesoclupea Ping and Yen 1933
+
†Mesoclupea globicephala Ping and Yen 1933
†Mesoclupea showchangensis Ping and Yen 1933
G. †Ogunichthys Alvarado-Ortega and Brito 2009
+
†Ogunichthys triangularis Alvarado-Ortega and Brito 2010
G. †Pachythrissops Woodward 1919
+
†Pachythrissops furcatus Eastman 1914
†Pachythrissops laevis Woodward 1919
†Pachythrissops propterus Wagner 1863
†Pachythrissops vectensis Woodward 1890
Invalid names: Parathrissops Eastman 1914 [replaced]
G. †Proportheus Jaekel 1909
+
†Proportheus kameruni Jaekel 1909
+
Invalid names: Chirocentrites guinensis Weiler 1922 [synonym]
G. †Sultanuvaisia Nessov 1981
+
†Sultanuvaisia antiqua Nessov 1981
G. †Vallecillichthys Blanco and Cavin 2003
+
†Vallecillichthys multivertebratum Blanco and Cavin 2003
Invalid names: Chuhsiungichthiidae Yabumoto 1994 [empty]
G. †Jinanichthys Ma and Sun 1988
+
†Jinanichthys longicephalus Ma and Sun 1988
+
+
Invalid names: Bleekeriidae [empty]
+
Invalid names: Bovictidae [empty]
+
Invalid names: Limnichthyidae [empty], Muglioididae [empty], Parapercidae [empty], Percophididae [empty], Pteropsaridae [empty]
+
Invalid names: Cerdalidae [empty], Derepodichthyidae [empty], Lycodapodidae [empty]
Invalid names: Bathymasteriformes [empty], Brotuliformes [empty], Carapiformes [empty], Haplodoci [empty]
+
Fm. †Ascalaboidae Arratia 2016
+
G. †Ascalobos Muenster 1839
Fm. †Leptolepididae Owen 1859
+
G. †Leptolepis Agassiz 1832
+
Invalid names: Ascalabos Münster 1839 [synonym], Megastoma Costa 1850 [synonym], Oxygonius Agassiz 1845 [synonym], Sarginites Costa 1850 [synonym], Tharsis Giebel 1848 [synonym]
G. †Longileptolepis Arratia 2003
+
Invalid names: Paraleptolepis Yabumoto 1994 [replaced]
G. †Nephrotus Meyer 1851
G. †Leucichthyops Cockerell 1919
G. †Longdeichthys Liu 1982
+
G. †Luisiella
+
Invalid names: Lipogenyidae [empty]
+
Invalid names: Osteoglossocephala [empty]
Unr. Osteoglossomorpha Greenwood et al. 1966
+
G. †Chandlerichthys Grande 1986
+
+
Fm. Hiodontidae Cuvier and Valenciennes 1846
Fm. †Jiuquanichthyidae Ma 1993
+
G. †Jiuquanichthys Ma 1993
G. †Joffrichthys Li and Wilson 1996
+
†Joffrichthys symmetropterus Li and Wilson 1996
†Joffrichthys tanyourus Murray et al. 2018
†Joffrichthys triangulpterus Newbrey and Bozek 2000
Fm. †Kipalaichthyidae Taverne 1976
+
G. †Kipalaichthys Casier 1965
Fm. †Kuyangichthyidae Liu et al. 1982
+
G. †Kuyangichthys Liu and Wang 1986
+
Or. Osteoglossiformes Berg 1940
+
G. †Heterosteoglossum Bonde 2008
G. †Lopadichthys Murray et al. 2018
G. †Monopteros Volta 1796
Fm. Notopteridae [knifefish]
G. †Opsithrissops Daniltshenko 1968
Fm. Osteoglossidae Bonaparte 1832 [arowana]
G. †Tongxinichthys Ma 1980
Fm. †Wakinoichthiidae Yabumoto 1994
G. Wilsonichthys Murray et al. 2016
G. †Xixiaichthys Zhang 2004
Invalid names: Mormyridae [empty], Saurocephalidae [empty]
G. †Shuleichthys Murray et al. 2010
+
†Shuleichthys brachypteryx Murray et al. 2010
G. †Paralycoptera Chang and Chou 1977
+
†Paralycoptera changi Ma and Sun 1988
Or. †Pediculati
+
Invalid names: Aceratiidae [empty], Gigactinidae [empty]
Unr. †Percomorphacea Wiley and Johnson 2010
+
G. †Acronuroides Bannikov et al. 2019
+
†Acronuroides eocaenicus Bannikov et al. 2019
G. †Ammutichthys Calzoni et al. 2025
+
†Ammutichthys loricatus Calzoni et al. 2025
G. †Erebusia Calzoni et al. 2025
+
†Erebusia tenebrae Calzoni et al. 2025
+
Invalid names: Coridae [empty], Neolabridae [empty], Sciaenoidea Gill 1872 [empty], Siphonognathidae [empty], Sparisomidae [empty]
+
Fm. †Ankylophoridae Gaudant 1978
+
G. †Ankylophorus Gaudant 1978
G. †Pholidorhynchodon Zambelli 1980
Fm. †Archaeomaenidae Wade 1941
+
G. †Gurvanichthys Jakovlev 1986
G. †Madariscus Wade 1941
Fm. †Eurycormidae Arratia 2017
+
G. †Grimmenichthys Konwert and Hörnig 2018
+
†Grimmenichthys ansorgei Konwert and Hörnig 2018
Fm. †Ichthyokentemidae Griffith and Patterson 1963
+
G. †Elpistoichthys Griffith 1977
Fm. †Ligulellidae Saint-Seine 1955
+
G. †Ligulella
G. †Luxembourgichthys Taverne and Steurbaut 2017
+
Fm. †Oligopleuridae Woodward 1895
+
G. †Oeonoscopus Costa 1864
G. †Oligopleura Thiolliere 1850
G. †Oligopleurus Thiollière 1850
Fm. †Pholidophoridae Woodward 1890
+
G. †Annaichthys Arratia 2013
G. †Archaeomaene Woodward 1895
G. †Ceneichthys Taverne and Capasso 2015
G. †Ceramurus Egerton 1845
G. †Eopholidophorus Zambelli 1989
G. †Knerichthys Arratia 2013
G. †Lombardichthys Arratia 2017
G. †Malingichthys Tintori et al. 2015
G. †Neopholidophoropsis Taverne 1981
G. †Pholidoctenus Zambelli 1978
G. †Pholidolepis Nybelin 1966
G. †Pholidophoretes Griffith 1977
G. †Pholidophoroides Woodward 1941
G. †Pholidophoropsis Nybelin 1966
G. †Pholidophorus Agassiz 1832
+
Invalid names: Microps Agassiz 1833 [synonym], Pholidophorus Zigno 1856 [objective synonym]
G. †Pseudopholidoctenus Arratia and Schultze 2024
G. †Zambellichthys Arratia 2013
Invalid names: Pholidophorinae Zambelli 1986 [empty]
+
G. †Gondwanapleuropholis Brito and Gallo 2002
G. †Pleuropholis Egerton 1858
Invalid names: Catervariolidae [empty]
+
G. †Ctenopomichthys Whitley 1940
G. Eutrigla Fraser-Brunner 1938
+
G. Hoplichthys Cuvier and Valenciennes 1829
G. †Praehoplichthys Schwarzhans 1980
G. Neoplatycephalus Castelnau 1872
G. Notesthes Ogilby 1904
G. Ophiodon Girard 1854
+
Ophiodon elongatus Girard 1854 [lingcod]
†Ophiodon ozymandias Jordan 1921
+
G. Plectrogenium Gilbert 1905
G. Podothecus Gill 1861
Fm. Triglidae [sea robin]
+
G. †Acanthatrigla Gaemers 1984
G. †Aspitrigla Fowler 1925
G. Chelidonichthys Kaup 1873
G. Lepidotrigla Guenther 1860
G. Peristedion Lacépède 1802
G. Prionotus Lacépède 1802
G. Pterygotrigla Waite 1899
G. Trigla Linnaeus 1758
G. †Triglidarum Risso 1826
G. †Trigloides Van Beneden 1871
G. Zaniolepis Girard 1857
+
Zaniolepis latipinnis Girard 1858 [longspine combfish]
Invalid names: Anoplopomatoidei [empty], Aploactinidae [empty], Bathylutichthyidae [empty], Bembridae [empty], Caracanthidae [empty], Eschmeyeridae [empty], Gnathanacanthidae [empty], Hexagrammoidei [empty], Pataecidae [empty], Peristediidae [empty], Platycephaloidei [empty]
Fm. †Sorbinicharacidae Taverne 2003
+
G. †Sorbinicharax Taverne 2003
+
+
Invalid names: Alabidae [empty]
Subor. Holostomi
+
Invalid names: Amphipnoidae [empty]
+
Invalid names: Flutidae [empty]
G. †Teleosteorum Unknown
+
†Teleosteorum pujoli Steurbaut 1984
G. †Todiltia Schaeffer and Patterson 1984
+
†Todiltia schoewei Dunkle 1942
G. †Tonghuaichthys Ma and Sun 1988
+
†Tonghuaichthys sanyuanpuensis Ma and Sun 1988
Or. †Tselfatiiformes Nelson 1994
+
G. †Eoplethodus Taverne 2000
+
†Eoplethodus chaneti Taverne 2000
Fm. †Plethodidae Loomis 1900
+
G. †Bananogmius Whitley 1940
+
Invalid names: Ananogmius White and Moy-Thomas 1940 [objective synonym], Anogmius Cope 1877 [replaced]
G. †Dixonanogmius Taverne 2000
G. †Niobrara
G. †Paranogmius Weiler 1935
G. †Pentanogmius Taverne 2000
G. †Plethodus Dixon 1850
G. †Rhamphoichthys El Hossny et al. 2023
G. †Thryptodus Loomis 1900
G. †Tselfatia
G. †Zanclites
Invalid names: Bananogmiidae Applegate 1970 [synonym], Thryptodontidae [synonym], Tselfatiidae Arambourg 1954 [synonym]
Fm. †Protobramidae Le Danois and Le Danois 1964
+
G. †Abisaadichthys Taverne and Gayet 2004
G. †Eusebichthys Taverne and Gayet 2004
G. †Protobrama Woodward 1942
G. †Pseudanogmius Taverne 2002
+
†Pseudanogmius maiseyi Taverne 2002
G. †Ypsiloichthys El Hossny and Cavin 2023
+
†Ypsiloichthys sibelleae El Hossny and Cavin 2023
Invalid names: Amphiuma nordica Brunner 1956 [nomen dubium], Bathyclupeiformes [empty], Catosteomi Boulenger [empty], Chromides [empty], Discocephali [empty], Malacopteri [empty], Opisthomi [empty], Osteossomorpha [empty], Polymyxiomorpha [empty], Stephanoberyciformes [empty], Teleocephala de Pinna 1996 [empty], Xenopterygii [empty]
G. †Westlepis Bender 2004
+
†Westlepis kempeni Bender 2004
G. †Woodichthys Coates 1998
Invalid names: Discordichthyiformes Minikh 1998 [empty], Docopteri Cope 1877 [empty], Inichthyidae Kazantseva-Selezneva 1981 [empty], Lysopteri Cope 1880 [empty], Merospondyli Cope 1887 [empty], Mesichthyes Hay 1902 [empty], Palaeophoxinus Coates 1988 [nomen nudum], Phthinobranchii Hay 1902 [empty], Plectospondyli Cope 1872 [empty]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: phosphaticsubp
Environment: marinesubp
Locomotion: actively mobilec
Life habit: planktonicp
Diet: suspension feederp
Created: 2009-07-20 21:50:54
Modified: 2009-07-20 23:50:54
Source: c = class, subp = subphylum, p = phylum
References: Kiessling 2004, Bambach et al. 2007, Hendy et al. 2009, Carroll 1988

Age range

Maximum range based only on fossils: base of the Eifelian to the top of the Holocene or 393.47000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 387.95 Ma

Collections (3916 total)

Oldest occurrences

Time interval Ma Country or state Original ID and collection number
Eifelian393.47 - 387.95Belarus (Vitebsk) Moythomasia sp. (232958)
Eifelian393.47 - 387.95Belarus Moythomasia sp. (225881)
Middle Devonian393.47 - 382.31Hong Kong Moythomasia sp. (275906)
Givetian387.95 - 382.31USA (Wyoming) Actinopterygia indet. (240984)
Givetian387.95 - 382.31Belarus (Brest) Moythomasia lebedevi (242243 242244)
Givetian387.95 - 382.31Belarus (Gormel) Moythomasia lebedevi (242242)
Givetian - Frasnian387.95 - 372.15USA (Iowa) Moythomasia sp. (226053)
Middle Givetian385.9 - 383.4Belarus Moythomasia lebedevi (225887)
Late/Upper Givetian383.4 - 382.31Belarus Moythomasia sp. (225888)
Late/Upper Givetian383.4 - 382.31Estonia Moythomasia sp. (225778)
Late/Upper Givetian - Early/Lower Frasnian383.4 - 379.0Germany (North Rhine-Westphalia) Moythomasia nitida, Moythomasia lineata (197071)
Early/Lower Frasnian382.31 - 379.0Iran (Esfahan) Moythomasia sp. (90626)
Early/Lower Frasnian382.31 - 379.0Iran Moythomasia devonica (226048)
Early/Lower Frasnian382.31 - 379.0Australia (Western Australia) Moythomasia durgaringa (170988 197263)
Early/Lower Frasnian382.31 - 379.0Latvia (Koknese) Aldingeria perforata (197072)
Frasnian382.31 - 372.15Australia (Western Australia) Moythomasia durgarina (232512) Moythomasia durgaringa (136090 232511 232517)
Frasnian382.31 - 372.15Australia (New South Wales) Moythomasia dungaringa (232510)
Frasnian382.31 - 372.15USA (New York) Moythomasia antiqua (232991) Palaeoniscus devonicus (226073)
Frasnian382.31 - 372.15Germany (Hesse) Moythomasia striata, Moythomasia laevigata (232992)
Frasnian382.31 - 372.15Russian Federation Moythomasia perforata (239134)