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Artiodactyla (even-toed ungulate)
Taxonomy
Artiodactyla was named by Owen (1848) [Owen does not give a rank in his original description of Artiodactyla.]. It is extant.
It was reranked as the suborder Artiodactyla by Cope (1889), Flower and Lydekker (1891), Hay (1902), Andrews (1906), Lambe (1908); it was synonymized subjectively with Cetartiodactyla by Waddell et al. (1999), Ursing et al. (2000), Boisserie et al. (2005), Boisserie et al. (2011); it was reranked as the unranked clade Artiodactyla by Gadow (1898), McKenna (1975), Hooker (1986), Prothero and Liter (2008), Prothero (2008), Spaulding et al. (2009), Morgan et al. (2009), Uhen et al. (2011), Rose et al. (2020), Carbot-Chanona et al. (2023).
It was assigned to Diplarthra by Cope (1889); to Ungulata by Flower and Lydekker (1891), Gadow (1898), Hay (1902), Lambe (1908); to Paraxonia by Gregory (1910); to Ungulata by Thorpe (1937); to Eparctocyona by McKenna (1975); to Eutheria by Hooker (1986); to Eutheria by Carroll (1988), Nowak (1991); to Ungulata by Owen (1848), Flower (1883), Andrews (1906), Archibald (1998); to Cetartiodactyla by Montgelard et al. (1997), Thewissen et al. (2001); to Artiodactylamorpha by Spaulding et al. (2009); to Mammalia by Uhen et al. (2011); to Eutheria by Vianey-Liaud et al. (2014); to Euungulata by Asher and Helgen (2010), Rose et al. (2020); and to Mammalia by Leidy (1873), Ferrusquia-Villafranca (1969), Hamilton (1978), Thurmond and Jones (1981), Stuenes (1989), Shoshani et al. (1989), Lucas et al. (1998), Weston (2000), Boisserie et al. (2003), Wilson and Reeder (2005), Boisserie (2005), Ferrusquía-Villafranca (2006), Hulbert and Whitmore (2006), Prothero and Liter (2008), Prothero (2008), Morgan et al. (2009), Lander and Lindsay (2011), Koufos (2020), Carbot-Chanona et al. (2023).
It was reranked as the suborder Artiodactyla by Cope (1889), Flower and Lydekker (1891), Hay (1902), Andrews (1906), Lambe (1908); it was synonymized subjectively with Cetartiodactyla by Waddell et al. (1999), Ursing et al. (2000), Boisserie et al. (2005), Boisserie et al. (2011); it was reranked as the unranked clade Artiodactyla by Gadow (1898), McKenna (1975), Hooker (1986), Prothero and Liter (2008), Prothero (2008), Spaulding et al. (2009), Morgan et al. (2009), Uhen et al. (2011), Rose et al. (2020), Carbot-Chanona et al. (2023).
It was assigned to Diplarthra by Cope (1889); to Ungulata by Flower and Lydekker (1891), Gadow (1898), Hay (1902), Lambe (1908); to Paraxonia by Gregory (1910); to Ungulata by Thorpe (1937); to Eparctocyona by McKenna (1975); to Eutheria by Hooker (1986); to Eutheria by Carroll (1988), Nowak (1991); to Ungulata by Owen (1848), Flower (1883), Andrews (1906), Archibald (1998); to Cetartiodactyla by Montgelard et al. (1997), Thewissen et al. (2001); to Artiodactylamorpha by Spaulding et al. (2009); to Mammalia by Uhen et al. (2011); to Eutheria by Vianey-Liaud et al. (2014); to Euungulata by Asher and Helgen (2010), Rose et al. (2020); and to Mammalia by Leidy (1873), Ferrusquia-Villafranca (1969), Hamilton (1978), Thurmond and Jones (1981), Stuenes (1989), Shoshani et al. (1989), Lucas et al. (1998), Weston (2000), Boisserie et al. (2003), Wilson and Reeder (2005), Boisserie (2005), Ferrusquía-Villafranca (2006), Hulbert and Whitmore (2006), Prothero and Liter (2008), Prothero (2008), Morgan et al. (2009), Lander and Lindsay (2011), Koufos (2020), Carbot-Chanona et al. (2023).
Subtaxa
Antiacodontidae, Antiacodontinae, Bunomerycidae, Cainotherioidea, Cetruminantia, Choeropotamoidea, Climacoceratidae, Diacodexeidae, Dichobunoidea, Entelodontoidea, Eolantianus, Erlianhyus, Eurytheriidae, Haplobunodontidae, Hsanotherium, Leptochoerinae, Merycoidodontoidea, Myanmarius, Neoselenodontia, Nonruminantia, Omnivora, Raphenacodus, Ruminantia (syn. Ruminantes), Ruminantiamorpha, Selenodontia, Suiformes (syn. Palaeodonta, Tesserachenae), Suoidea (syn. Suinamorpha), Tapirulidae, Tragulohyus, Trigonolestidae, Tylopoda (syn. Camelidamorpha, Hydrophorae), Whippomorpha (syn. Cetancodontamorpha, Cetancodonta)
Entered
by J. Alroy on 2003-01-23; modified by M. Uhen on 2013-10-27
Synonyms
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Synonymy list
| Year | Name and author |
|---|---|
| 1848 | Artiodactyla Owen p. 131 |
| 1873 | Artiodactyla Leidy p. 317 |
| 1883 | Artiodactyla Flower p. 184 |
| 1889 | Artiodactyla Cope p. 876 |
| 1891 | Artiodactyla Flower and Lydekker p. 89 |
| 1898 | Artiodactyla Gadow p. 48 |
| 1902 | Artiodactyla Hay p. 647 |
| 1906 | Artiodactyla Andrews p. 178 |
| 1908 | Artiodactyla Lambe |
| 1910 | Artiodactyla Gregory p. 466 |
| 1937 | Artiodactyla Thorpe p. 31 |
| 1969 | Artiodactyla Ferrusquia-Villafranca |
| 1975 | Artiodactyla McKenna p. 43 |
| 1978 | Artiodactyla Hamilton p. 168 |
| 1981 | Artiodactyla Thurmond and Jones p. 187 |
| 1986 | Artiodactyla Hooker p. 376 |
| 1988 | Artiodactyla Carroll |
| 1989 | Artiodactyla Shoshani et al. p. 436 |
| 1989 | Artiodactyla Stuenes p. 243 |
| 1991 | Artiodactyla Nowak |
| 1997 | Cetartiodactyla Montgelard et al. |
| 1997 | Artiodactyla Montgelard et al. p. 555 |
| 1997 | Cetartiodactyla Montgelard et al. p. 556 |
| 1998 | Artiodactyla Archibald |
| 1998 | Artiodactyla Lucas et al. p. 426 |
| 1999 | Cetartiodactyla Waddell et al. p. 3 |
| 2000 | Artiodactyla Weston p. 177 |
| 2001 | Cetartiodactyla Madsen et al. p. 610 |
| 2001 | Artiodactyla Thewissen et al. p. suppl. |
| 2001 | Cetartiodactyla Thewissen et al. p. suppl. |
| 2001 | Cetartiodactyla Waddell et al. p. 148 |
| 2003 | Artiodactyla Boisserie et al. p. 16 |
| 2004 | Cetartiodactyla Uhen p. 11 |
| 2005 | Artiodactyla Boisserie p. 12 |
| 2005 | Artiodactyla Wilson and Reeder |
| 2006 | Artiodactyla Ferrusquía-Villafranca p. 993 |
| 2006 | Artiodactyla Hulbert and Whitmore p. 6 |
| 2008 | Cetartiodactyla Bisconti p. 162 |
| 2008 | Artiodactyla Prothero |
| 2008 | Artiodactyla Prothero and Liter |
| 2009 | Artiodactyla Morgan et al. |
| 2009 | Artiodactyla Spaulding et al. |
| 2010 | Artiodactyla Asher and Helgen p. 3 |
| 2010 | Cetartiodactyla Weston and Boisserie p. 853 |
| 2011 | Artiodactyla Lander and Lindsay p. 221 |
| 2011 | Artiodactyla Uhen et al. p. 957 |
| 2014 | Artiodactyla Vianey-Liaud et al. p. 606 |
| 2020 | Cetartiodactyla Bisconti and Bosselaers p. 8 |
| 2020 | Artiodactyla Koufos |
| 2020 | Artiodactyla Rose et al. p. 101 figs. Fig. 81 |
| 2023 | Artiodactyla Carbot-Chanona et al. p. 4 |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Unr. Artiodactyla Owen 1848 [even-toed ungulate]
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Superfm. †Cainotherioidea Camp and VanDerHoof 1940
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Fm. †Cainotheriidae Cope 1881
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Subfm. †Cainotheriinae Camp and VanderHoof 1940
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G. †Caenomeryx Hürzeler 1936
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†Caenomeryx filholi Lydekker 1885
†Caenomeryx procommunis Filhol 1877
G. †Cainotherium Bravard 1828
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†Cainotherium bavaricum Berger 1959
†Cainotherium commune Bravard 1835
†Cainotherium huerzeleri Heizmann 1983
†Cainotherium laticurvatum Geoffroy 1833
+
Invalid names: Caenotherium commune Bravard 1835 [synonym], Caenotherium concinnum Meyer 1843 [synonym], Caenotherium cournonense Blainville 1849 [synonym], Caenotherium elegans Pomel 1847 [synonym], Caenotherium geoffroyi Pomel 1851 [synonym], Caenotherium leptognathum Leizer and Parieu 1838 [synonym], Caenotherium leptorhynchum Pomel 1846 [synonym], Caenotherium medium Blainville 1849 [synonym], Caenotherium metopias Pomel 1851 [synonym], Caenotherium minimum Bravard 1835 [synonym], Caenotherium nouleti Lartet 1851 [synonym]
†Cainotherium lintillae Baudelot and Crouzel 1974
†Cainotherium miocaenicum Crusafont-Pairo 1955
†Cainotherium renggeri Meyer 1837
Invalid names: Caenotherium Agassiz 1846 [objective synonym], Crinotherium Filhol 1882 [synonym]
G. †Procaenotherium Hürzeler 1936
Subfm. †Oxacroninae Hurzeler 1936
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Fm. †Robiacinidae Weppe et al. 2019
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G. †Robiacina Sudre 1969
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†Robiacina lavergnensis Sudre 1977
†Robiacina minuta Sudre 1969
†Robiacina quercyi Sudre 1977
†Robiacina weidmanni Sudre 1978
Superfm. †Choeropotamoidea
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Fm. †Choeropotamidae Owen 1845
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G. †Amphirhagatherium Deperet 1908
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†Amphirhagatherium edwardsi Hooker and Thomas 2001
†Amphirhagatherium fronstettense Kowalevsky 1874
†Amphirhagatherium louisi Sudre 1978
†Amphirhagatherium neumarkensis Erfurt and Haubold 1989
†Amphirhagatherium ruetimeyeri Pavlov 1900
†Amphirhagatherium weigelti Heller 1934
G. †Choeropotamus Desmarest 1822
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†Choeropotamus affinis Gervais 1852
†Choeropotamus depereti Stehlin 1908
†Choeropotamus parisiensis Cuvier 1822
†Choeropotamus sudrei Casanovas-Cladellas 1975
G. †Haplobunodon Deperet 1908
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†Haplobunodon lydekkeri Stehlin 1908
†Haplobunodon meridonale Sudre 1997
†Haplobunodon muelleri Rutimeyer 1862
†Haplobunodon solodurense Stehlin 1908
†Haplobunodon venatorum Hooker 1986
G. †Lophiobunodon Deperet 1908
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†Lophiobunodon hookeri Godinot et al. 2018
†Lophiobunodon minervoisensis Deperet 1908
†Lophiobunodon rhodanicum Deperet 1908
G. †Masilabune Tobien 1980
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†Masillabune franzeni Erfurt and Haubold 1989
†Masillabune martini Tobien 1980
G. †Rhagatherium Pictet 1857
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†Rhagatherium kowalevskyi Stehlin 1908
†Rhagatherium valdense Pictet 1857
Fm. †Climacoceratidae Hamilton 1978
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G. †Climacoceras MacInnes 1936
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†Climacoceras africanus MacInnes 1936
†Climacoceras gentryi Hamilton 1978
G. †Prolibytherium Arambourg 1961
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†Prolibytherium fusus Danowitz et al. 2016
†Prolibytherium magnieri Arambourg 1961
Invalid names: Climacoceratinae Hamilton 1978 [empty], Sperrgebietomerycinae Morales et al. 1999 [empty]
Fm. †Diacodexeidae Krishtalka and Stucky 1985
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G. †Bunophorus Sinclair 1914
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†Bunophorus etsagicus Cope 1882
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Invalid names: Eohippus borealis Granger 1908 [synonym], Eohippus resartus Granger 1908 [synonym], Hyracotherium cristatum Wortman 1896 [synonym]
†Bunophorus grangeri Sinclair 1914
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Invalid names: Wasatchia dorseyana Sinclair 1914 [synonym], Wasatchia lysitensis Sinclair 1914 [synonym]
†Bunophorus macropternus Cope 1882
†Bunophorus pattersoni Krishtalka and Stucky 1986
†Bunophorus robustus Sinclair 1914
†Bunophorus sinclairi Guthrie 1966
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†Bunophorus sinclairi robinsoni Stucky and Krishtalka 1990
†Bunophorus sinclairi sinclairi Guthrie 1966
Invalid names: Bunophorus gazini Guthrie 1971 [synonym]
G. †Diacodexis Cope 1882
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†Diacodexis antunesi Estravís and Russell 1989
†Diacodexis corsaensis Checa 2004
†Diacodexis gazini Godinot 1978
†Diacodexis gigasei Smith 1996
†Diacodexis gracilis Krishtalka and Stucky 1985
†Diacodexis ilicis Gingerich 1989
†Diacodexis indicus Bajpai et al. 2005
†Diacodexis kelleyi Krishtalka and Stucky 1985
†Diacodexis metsiacus Cope 1882
†Diacodexis minutus Krishtalka and Stucky 1985
†Diacodexis morrisi Hooker 2010
†Diacodexis parvus Kumar et al. 2010
†Diacodexis primus Krishtalka and Stucky 1985
†Diacodexis secans Cope 1881
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Invalid names: Diacodexis laticuneus Cope 1882 [synonym], Diacodexis olseni Sinclair 1914 [synonym], Mioclaenus brachystomus Cope 1882 [synonym]
†Diacodexis varleti Sudre 1983
Invalid names: Trigonolestes Cope 1894 [synonym], Wasatchia Sinclair 1914 [synonym]
Invalid names: Bunophorinae Stucky and Krishtalka 1990 [empty]
Superfm. †Dichobunoidea Gill 1872
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G. †Acoessus Cope 1881
Fm. †Cebochoeridae Lydekker
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G. †Acotherulum Gervais 1850
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†Acotherulum campichii Pictet 1855
†Acotherulum pumilum Stehlin 1908
†Acotherulum quercyi Stehlin 1908
†Acotherulum saturninum Gervais 1850
G. †Cebochoerus Gervais 1852
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†Cebochoerus anceps Gervais 1852
†Cebochoerus fontensis Sudre 1978
†Cebochoerus helveticus Pictet and Humbert 1869
†Cebochoerus lacustris Gervais 1856
†Cebochoerus robiacensis Deperet 1917
G. †Choeromorus Gervais 1848
G. †Gervachoerus Sudre 1978
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†Gervachoerus dawsoni Sudre 1978
†Gervachoerus jaegeri Sudre 1978
†Gervachoerus minor Gervais 1876
†Gervachoerus suillus Gervais 1859
Fm. †Dichobunidae Gill 1872
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G. †Chorlakkia Gingerich et al. 1979
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†Chorlakkia hassani Gingerich et al. 1979
†Chorlakkia valerii Vislobokova 2004
Subfm. †Dichobuninae Turner 1849
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G. †Aumelasia Sudre 1980
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†Aumelasia gabineaudi Sudre 1980
†Aumelasia maniai Erfurt and Haubold 1989
†Aumelasia menieli Sudre 1983
†Aumelasia sudrei Godinot et al. 2018
G. †Dichobune Cuvier 1822
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†Dichobune cervinum Owen 1825
†Dichobune fraasi Schlosser 1902
†Dichobune jehennei Brunet and Sudre 1980
†Dichobune leporina Cuvier 1822
†Dichobune robertiana Gervais 1848
†Dichobune sigei Sudre 1978
G. †Meniscodon Rütimeyer 1888
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†Meniscodon europaeum Rutimeyer 1888
†Meniscodon picteti Rutimeyer 1891
G. †Messelobunodon Franzen 1980
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†Messelobunodon ceciliensis Franzen and Krumbiegel 1980
†Messelobunodon schaeferi Franzen 1980
G. †Metriotherium Filhol 1882
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†Metriotherium mirabile Filhol 1882
†Metriotherium paulum Stehlin 1906
†Metriotherium sarelense Astruc et al. 2003
G. †Protodichobune Lemoine 1891
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†Protodichobune hellmundi Godinot et al. 2018
†Protodichobune oweni Lemoine 1878
Invalid names: Dichobunini Turner 1849 [empty]
Subfm. †Eurodexeinae Erfurt and Sudre 1996
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G. †Eurodexis Erfurt and Sudre 1996
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†Eurodexis ceciliensis Franzen and Krumbiegel 1980
†Eurodexis russelli Sudre and Erfurt 1996
G. †Eygalayodon Sudre and Marandat 1993
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†Eygalayodon isavenaensi Checa 2004
†Eygalayodon montenati Sudre and Marandat 1993
Subfm. †Hyperdichobuninae Gentry and Hooker 1988
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G. †Hyperdichobune Stehlin 1910
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†Hyperdichobune hammeli Sudre 1978
†Hyperdichobune langi Rutimeyer 1891
†Hyperdichobune nobilis Stehlin 1906
†Hyperdichobune spectabilis Stehlin 1910
†Hyperdichobune spinifera Stehlin 1906
G. †Mouillacitherium Filhol 1882
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†Mouillacitherium carteri Rutimeyer 1891
†Mouillacitherium cartieri Rutimeyer 1891
†Mouillacitherium elegans Filhol 1882
†Mouillacitherium schlosseri Sudre 1978
Subfm. †Lantianiinae Metais et al. 2004
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G. †Elaschitotherium Metais et al. 2004
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†Elaschitotherium crepaturus Metais et al. 2008
†Elaschitotherium qii Metais et al. 2004
G. †Mioachoerus
Invalid names: Diacodexinae Gazin 1955 [empty]
Fm. †Helohyidae Marsh 1877
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G. †Achaenodon Cope 1873
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†Achaenodon fremdi Lucas et al. 2004
†Achaenodon insolens Cope 1873
†Achaenodon robustus Osborn 1883
†Achaenodon uintense Osborn 1895
Invalid names: Protelotherium Osborn 1895 [synonym]
G. †Gobiohyus Matthew and Granger 1925
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†Gobiohyus orientalis Matthew and Granger 1925
†Gobiohyus pressidens Matthew and Granger 1925
†Gobiohyus reshetovi Vislobokova 2004
†Gobiohyus robustus Matthew and Granger 1925
†Gobiohyus yuanchuensis Young 1937
G. †Helohyus Marsh 1872
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Invalid names: Ithygrammodon cameloides Osborn et al. 1878 [nomen dubium], Lophiohyus Sinclair 1914 [synonym], Thinotherium Marsh 1872 [synonym], Thinotherium Cope 1870 [synonym]
G. †Ithygrammodon Osborn et al. 1878
G. †Lophiohys
G. †Pakkokuhyus Holroyd and Ciochon 1995
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†Pakkokuhyus lahirii Pilgrim 1928
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Invalid names: Siamotherium pondaungensis Ducroq and Soe 2000 [synonym]
Invalid names: Achaenodontidae [synonym]
Fm. †Homacodontidae Marsh 1849
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G. †Antiacodon Marsh 1872
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†Antiacodon crassus Cope 1875
†Antiacodon furcatus Cope 1873
†Antiacodon pygmaeus Cope 1872
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†Antiacodon pygmaeus huerfanensis Robinson 1966
Invalid names: Nanomeryx caudatus Marsh 1894 [synonym]
†Antiacodon vanvaleni Guthrie 1971
†Antiacodon venustus Marsh 1872
Invalid names: Nanomeryx Marsh 1894 [synonym], Sarcolemur bicuspis Loomis 1907 [nomen nudum]
G. †Bunomeryx Wortman 1898
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G. †Homacodon Marsh 1872
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†Homacodon priscus Marsh 1894
†Homacodon pucillus Marsh 1894
†Homacodon vagans Marsh 1872
G. †Hylomeryx Peterson 1919
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†Hylomeryx annectens Peterson 1919
†Hylomeryx quadricuspis Peterson 1919
Invalid names: Sphenomeryx Peterson 1919 [synonym]
G. †Pentacemylus Peterson 1932
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†Pentacemylus leotensis Gazin 1955
†Pentacemylus progressus Peterson 1932
Invalid names: Bunomerycini Gentry and Hooker 1988 [empty], Homacodontini Marsh 1894 [empty]
Fm. †Leptochoeridae Marsh 1894
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G. †Leptochoerus Leidy 1856
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†Leptochoerus emilyae Edwards 1976
†Leptochoerus spectabilis Leidy 1856
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Invalid names: Laopithecus robustus Marsh 1875 [synonym], Leptochoerus gracilis Marsh 1894 [synonym]
†Leptochoerus supremus Macdonald 1955
Invalid names: Laopithecus Marsh 1875 [synonym]
G. †Stibarus Cope 1873
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†Stibarus montanus Matthew 1903
†Stibarus yoderensis Macdonald 1955
Invalid names: Menotherium Cope 1874 [synonym], Nanochoerus Macdonald 1955 [synonym]
Invalid names: Diacodexis woltonensis Krishtalka and Stucky 1985 [invalid subgroup]
Fm. †Mixtotheriidae Pearson 1927
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G. †Mixtotherium Filhol 1880
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†Mixtotherium cuspidatum Filhol 1880
†Mixtotherium depressum Filhol 1884
†Mixtotherium gresslyi Rutimeyer 1891
†Mixtotherium infans Stehlin 1910
†Mixtotherium lavergnense Sudre 1977
†Mixtotherium leenhardti Stehlin 1908
†Mixtotherium priscum Stehlin 1908
†Mixtotherium quercyi Filhol 1884
Fm. †Raoellidae Sahni et al. 1981
+
G. †Indohyus Ranga Rao 1971
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†Indohyus indirae Ranga Rao 1971
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Invalid names: Haqueina haquei Dehm and Oettingen-Spielberg 1958 [synonym], Indohyus kalakotensis Ranga Rao 1971 [synonym], Kunmunella rajauriensis Sahni and Khare 1971 [synonym], Kunmunella transversa Kumar and Sahni 1985 [synonym], Metkatius kashmiriensis Kumar and Sahni 1985 [synonym], Raoella dograi Sahni and Khare 1971 [synonym]
†Indohyus major Thewissen et al. 1987
Invalid names: Kunmunella Sahni and Khare 1971 [synonym], Metkatius Kumar and Sahni 1985 [synonym], Raoella Sahni and Khare 1971 [synonym]
G. †Khirtharia Pilgrim 1940
+
†Khirtharia aurea Thewissen et al. 2001
†Khirtharia dayi Pilgrim 1940
†Khirtharia inflatus Ranga Rao 1972
Invalid names: Bunodentus Ranga Rao 1972 [synonym]
G. †Raphenacodus Gabunia 1971
Subor. Ruminantia Scopoli 1777 [ruminant]
+
Fm. †Agriochoeridae Leidy 1869 [agriochoerid]
+
G. †Agriochoerus Leidy 1850
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†Agriochoerus antiquus Leidy 1850
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†Agriochoerus antiquus dakotensis Thorpe 1921
Invalid names: Agriochoerus latifrons Leidy 1869 [synonym]
†Agriochoerus antiquus antiquus Leidy 1850
†Agriochoerus gaudryi Osborn and Wortman 1893
+
Invalid names: Agriomeryx migrans Marsh 1894 [synonym]
†Agriochoerus guyotianus Cope 1878
+
Invalid names: Agriochoerus bullatus Thorpe 1921 [synonym], Agriochoerus trifrons Cope 1884 [synonym], Coloreodon ferox Cope 1879 [synonym], Coloreodon macrocephalus Cope 1879 [synonym], Coloreodon ryderanus Cope 1881 [synonym]
†Agriochoerus major Leidy 1856
†Agriochoerus maximus Douglass 1902
Invalid names: Agriomeryx Marsh 1894 [synonym], Artionyx Osborn and Wortman 1893 [synonym], Coloreodon Cope 1879 [synonym], Eomeryx Marsh 1894 [synonym], Merycopater Cope 1879 [synonym]
G. †Diplobunops Peterson 1919
+
†Diplobunops crassus Scott 1945
†Diplobunops kardoula Emery et al. 2016
†Diplobunops matthewi Peterson 1919
+
Invalid names: Diplobunops leotensis Scott 1945 [synonym], Diplobunops uintensis Peterson 1931 [synonym], Diplobunops ultimus Peterson 1931 [synonym], Protoreodon pearcei Gazin 1956 [synonym]
†Diplobunops vanhouteni Gazin 1955
G. †Protoreodon Scott and Osborn 1887
+
†Protoreodon minimus Douglass 1902
†Protoreodon minor Scott 1899
†Protoreodon pacificus Golz 1976
†Protoreodon parvus Scott and Osborn 1887
+
Invalid names: Agriotherium paradoxicus Scott 1898 [synonym], Hyomeryx breviceps Marsh 1894 [synonym]
†Protoreodon petersoni Gazin 1955
†Protoreodon pumilus Marsh 1875
+
Invalid names: Mesagriochoerus primus Peterson 1934 [synonym], Protagriochoerus annectens Scott 1899 [synonym], Protoreodon medius Peterson 1919 [synonym], Protoreodon tardus Scott 1945 [synonym]
†Protoreodon transmontanus Stock 1949
†Protoreodon walshi Theodor 1999
Invalid names: Agriotherium Scott 1898 [synonym], Chorotherium Berg 1899 [synonym], Hyomeryx Marsh 1894 [synonym], Mesagriochoerus Peterson 1934 [synonym], Protagriochoerus Scott 1899 [synonym]
Invalid names: Agriochoerinae Cope 1884 [empty], Protoreodontinae Scott 1890 [empty]
Fm. †Bachitheriidae Janis 1987
+
G. †Bachitherium Filhol 1882
+
†Bachitherium curtum Filhol 1877
†Bachitherium guirounetensis Sudre 1995
†Bachitherium insigne Filhol 1882
†Bachitherium lavocati Sudre 1986
†Bachitherium thraciensis Mennecart et al. 2018
†Bachitherium vireti Sudre 1986
G. †Merycotherium Bojanus 1834
G. †Palaeohypsodontus Trofimov 1958
+
†Palaeohypsodontus asiaticus Trofimov 1958
†Palaeohypsodontus zinensis Métais et al. 2003
Unr. Pecora Linnaeus 1758
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Fm. Antilocapridae Gray 1866 [pronghorn]
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Subfm. †Antilocaprinae Gray 1866
+
G. †Capromeryx Matthew 1902
+
Invalid names: Breameryx Furlong Eustace 1946 [synonym], Dorcameryx [synonym]
G. †Ceratomeryx Gazin 1935
G. †Hexobelomeryx Furlong 1941
G. †Osbornoceros Frick 1937
G. †Ottoceros Miller and Downs 1974
G. †Proantilocapra Barbour and Schultz 1934
G. †Sphenophalos Merriam 1909
Tr. †Stockoceratini Frick 1937
G. †Tetrameryx Lull 1921
G. †Hexameryx White 1941
+
G. †Ilingoceros Merriam 1909
+
†Ilingoceros alexandrae Merriam 1909
†Ilingoceros schizoceras Merriam 1911
G. †Merycodus Leidy 1854
+
†Merycodus crucensis Frick 1937
†Merycodus hookwayi Furlong 1934
†Merycodus joraki Frick 1937
†Merycodus major Frick 1937
†Merycodus minimus Frick 1937
†Merycodus minor Frick 1937
†Merycodus necatus Leidy 1854
†Merycodus sabulonis Matthew and Cook 1909
+
Invalid names: Cosoryx (Subparacosoryx) savaronis Frick 1937 [synonym]
†Merycodus tehuanus Cope 1877
†Merycodus trilateralis Cope 1877
Invalid names: Dicrocerus Cope 1875 [synonym], Meryceros Frick 1937 [synonym], Merycodus prodromus Cook 1934 [nomen dubium]
G. †Paracosoryx Frick 1937
+
†Paracosoryx alticornis Frick 1937
†Paracosoryx dawesensis Frick 1937
†Paracosoryx furlongi Frick 1937
†Paracosoryx loxoceros Furlong 1934
†Paracosoryx nevadensis Merriam 1911
†Paracosoryx wilsoni Frick 1937
G. †Plioceros Frick 1937
+
†Plioceros blicki Frick 1937
†Plioceros dehlini Frick 1937
†Plioceros floblairi Frick 1937
G. †Texoceros Frick 1937
+
†Texoceros altidens Matthew 1924
+
Invalid names: Capromeryx texanus Hesse 1935 [synonym], Dorcameryx optimae Reed and Longnecker 1928 [synonym]
†Texoceros edensis Frick 1937
†Texoceros guymonensis Frick 1937
†Texoceros minorei Frick 1937
†Texoceros vaughani Frick 1937
Invalid names: Cosorycinae Frick 1937 [empty], Ramocerotinae Frick 1937 [empty]
Superfm. Bovoidea Gray 1821
+
G. †Amphimoschus Gray 1852
+
†Amphimoschus ponteleviensis Bourgeois 1873
†Amphimoschus xishuiensis Li et al. 2022
Fm. Bovidae Gray 1821 [antelope]
+
G. †Alephis Gromolard 1980
Subfm. Antilopinae Gray 1821
G. †Antilospira Teilhard de Jardin and Young 1931
Subfm. Bovinae Gray 1821
G. †Brabovus Gentry 1987
G. †Cambayella Pilgrim 1939
Subfm. Caprinae Gray 1821
G. †Caprotragoides Thenius 1979
G. †Criotherium Forsyth Major 1891
G. †Deperetia Schaub 1923
G. †Dorcadoryx Teilhard de Jardin and Trassaert 1938
G. †Dorcadoxa Pilgrim 1939
G. †Dystychoceras Kretzoi 1941
G. †Dytikodorcas Bouvrain and de Bonis 2007
G. †Eosyncerus Vekua 1972
G. †Euceratherium Furlong and Sinclair 1904 [shrub-ox]
+
Invalid names: Aftonius Hay 1913 [synonym], Preptoceras Furlong 1905 [synonym]
G. †Gallogoral Guérin 1865
G. †Gangraia Kostopoloulos 2021
G. †Gazellospira Pilgrim and Schaub 1939
G. †Gentrytragus Azanza 1994
G. †Gobiocerus Sokolov 1952
G. †Grevenobos Cregut-Bonnoure and Tsoukala 2017
G. †Hanhaicerus Huang 1985
G. †Helicoportax Pilgrim 1937
G. †Hemibos Falconer 1865
+
Invalid names: Amphibos Falconer and Cautley 1868 [synonym], Peribos Lydekker 1878 [synonym], Probubalus Rütimeyer [objective synonym]
G. †Hemistrepsiceros Pilgrim and Hopwood 1928
G. †Hesperidoceras Crusafont Pairo 1965
G. †Hezhengia Qiu et al. 2000
Subfm. †Hypsodontinae Köhler 1987
G. †Kabulicornis Heintz and Thomas 1981
G. †Kobikeryx Pilgrim 1939
G. †Kubanotragus Gabunia 1973
G. †Leptobos Rutimeyer 1878
G. †Leptotragus Bohlin 1936
G. †Linxiatragus Wang et al. 2023
G. †Liops Gidley 1906
G. †Lyrocerus Teilhard de Jardin and Trassaert 1938
G. †Majoreas Kostopoulos 2004
G. †Megalovis Schaub 1924
G. †Mesembriacerus Bouvrain 1975
G. †Mesembriportax Gentry 1974
G. †Namacerus Morales et al. 2003
G. Nesotragus von Duben 1846
G. †Nisidorcas Bouvrain 1979
Tr. †Oiocerini Pilgrim 1934
G. †Olonbulukia Bohlin 1937
G. †Oreonager Pomel 1895
G. †Orygotherium von Meyer 1838
G. †Pachygazella Teilhard de Jardin and Young 1931
G. †Pachyportax Pilgrim 1937
G. †Pachytragus Schlosser 1904
G. †Palaeoreas Gaudry 1861
G. †Palaeoryx Gaudry 1861
G. †Paraprotoryx Bohlin 1935
G. †Paratragocerus Sokolov 1949
G. †Parurmiatherium Sickenberg 1932
G. †Perimia Pilgrim 1939
G. †Platybos Pilgrim 1939
G. †Pliotragus Kretzoi 1941
+
Invalid names: Hesperoceras Villalta and Crusafont Pairó 1956 [synonym]
G. †Praeovibos Staudinger 1908
G. †Proamphibos Pilgrim 1939
G. †Procamptoceras Schaub 1923
G. †Procobus Khomenko 1913
G. †Prodamaliscus Schlosser 1904
G. †Proleptobos Pilgrim 1939
G. †Prosinotragus Bohlin 1935
G. †Prostrepsiceros Forsyth Major 1891
G. †Protoryx Forsyth Major 1891
G. †Protragelaphus Dames 1883
G. †Pseudobos Schlosser 1903
G. †Pseudoeotragus van der Made 1989
G. †Pseudotragus Schlosser 1904
G. †Pultiphagonides Hopwood 1934
G. †Qurliqnoria Bohlin 1937
G. Rhynchotragus Neumann 1905 [bontebok]
G. †Rhynotragus Reck 1925
G. †Ruticeros Pilgrim 1939
G. †Samokeros Solounias 1981
G. †Selenoportax Pilgrim 1937
G. †Shensispira Chiu et al. 1979
G. †Sinoreas Teilhard de Jardin and Trassaert 1938
G. †Sinoryx Teilhard de Jardin and Trassaert 1938
G. †Sinotragus Bohlin 1935
G. †Sivacapra Pilgrim 1939
G. †Sivaceros Pilgrim 1937
G. †Sivaportax Pilgrim 1939
G. †Sivatragus Pilgrim 1939
G. †Sivoryx Pilgrim 1939
G. †Skoufotragus Kostopoulos 2009
G. †Soergelia Schaub 1951
G. †Spirocerus Boule and Teilhard de Jardin 1928
G. †Strepsiportax Pilgrim 1937
G. †Strogulognathus Filhol 1890
G. †Tchaltacerus Bouvrain 1998
G. †Tethytragus Azanza 1994
G. †Toribos
G. †Torticornis Dmitrieva 1977
G. †Tossunnoria Bohlin 1937
G. †Tragocerus Gaudry 1861
G. †Tragoreas Schlosser 1908
G. †Tragospira Kretzoi 1954
G. †Tsaidamotherium Bohlin 1935
G. †Turritragus Abbazzi et al. 2008
Subfm. †Urmiatheriinae Sickenberg 1933
G. †Wellsiana Vrba 1987
Invalid names: Antilopina Bonaparte 1850 [empty], Camelopardalina Gray 1825 [empty], Capra depressa Linnaeus 1758 [nomen dubium], Capra reversa Linnaeus 1758 [nomen dubium], Capridae Gray 1821 [synonym], Caprina Bonaparte 1850 [empty], Hippotraginae Sundevall 1845 [empty], Peleinae Gray 1872 [empty], Tragelaphinae Jerdon 1874 [empty]
Fm. Moschidae Gray 1821 [musk deer]
+
G. †Bedenomeryx Jehenne 1988
Subfm. †Blastomerycinae Frick 1937
G. †Friburgomeryx Becker et al. 2001
G. †Hispanomeryx Morales et al. 1981
G. †Hydropotopsis Jehenne 1991
Subfm. Moschinae Gray 1821
G. †Oriomeryx Ginsburg 1985
G. †Pomelomeryx Ginsburg and Morales 1989
Invalid names: Booidea Gill 1872 [synonym]
G. †Bugtimeryx Ginsburg et al. 2001
Superfm. †Cervoidea
+
G. †Amphiprox Kaup 1839
Fm. Cervidae Gray 1821 [deer]
+
G. †Agalmaceros Hoffstetter 1952
Subfm. Alcinae Jerdon 1874
G. †Anoglochis Croizet and Jobert 1826
G. †Bretzia Fry and Gustafson 1974
Subfm. Capreolinae Brookes 1828
G. †Cervalces Scott 1885 [stag moose]
G. †Cervavitulus Kretzoi 1951
Tr. Cervina Gray 1825
Subfm. Cervinae Goldfuss 1820
G. †Cervodama Pidoplichko and Flerov 1952
G. †Charitoceros Hoffstetter 1963
G. †Eocoileus Webb 2000
G. †Euctenoceros Trouessart 1898
G. †Eumeryx Matthew and Granger 1924
G. †Furcifer Wagner 1844
G. †Lucentia Azanza and Montoya 1995
G. †Metacervulus Teilhard de Jardin and Trassaert 1937
Tr. Moschina Gray 1825
G. †Navahoceros Kurten 1975
G. †Nemomeryx Morea 1981
G. †Palaeoplatyceros Hernandez-Pacheco 1913
G. †Paracervulus Teilhard de Chardin and Trassaert 1937
G. †Praemegaceros Portis 1920
+
Invalid names: Allocaenelaphus Radulesco and Samson 1967 [synonym], Praerangifer Portis 1920 [synonym], Psecupsoceros Radulesco and Samson 1967 [synonym], Tamanalces Vereshchagin 1957 [synonym]
G. †Procervulus Gaudry 1877
G. Pseudaxis Gray 1872
Unr. Telemetacarpalia
Invalid names: Cervulinae [empty], Cervus aguangae Frick 1937 [nomen nudum], Cervus guineensis Linnaeus 1758 [nomen dubium], Plesiometacarpalia [empty], Pliocervinae Symeonidis 1974 [empty], Sangamona Hay 1920 [nomen vanum], Sangamona fugitiva Hay 1920 [nomen vanum], Surameryx Prothero et al. 2014 [nomen dubium], Surameryx acrensis Prothero et al. 2014 [nomen dubium]
Fm. Lagomerycidae Pilgrim 1941
+
Tr. Dicrocerini Simpson 1945
Tr. Lagomerycini Pilgrim 1941
G. †Ligeromeryx Azanza and Ginsburg 1997
G. †Paradicrocerus Gabunia 1959
G. †Teruelia Moya Sola 1987
G. †Dremotherium Geoffroy 1833
+
†Dremotherium cetinensis Ginsburg et al. 1994
†Dremotherium feignouxi Geoffroy 1833
†Dremotherium guthi Jehenne 1987
†Dremotherium quercyi Filhol 1887
Superfm. Giraffomorpha Sánchez et al. 2015
+
Fm. Giraffidae Gray 1821 [giraffe]
+
G. †Alcicephalus Rodler and Wiethofer 1890
G. †Birgerbohlinia Crusafont Pairó and Villalta 1951
G. †Bohlinia Matthew 1929
G. †Bramiscus Ríos et al. 2024
Subfm. †Canthumerycinae Hamilton 1978
G. †Decennatherium Crusafont Pairo 1952
G. †Georgiomeryx Paraskevaidis 1940
Tr. Giraffini
Subfm. †Giraffokerycinae Solounias 2007
G. †Helladotherium Gaudry 1860
G. †Honanotherium Bohlin 1926
Subfm. †Okapiinae Bohlin 1926
Tr. Okapini
G. †Palaeogiraffa Bonis and Bouvrain 2003
Tr. †Palaeotragini Pilgrim 1911
G. †Progiraffa Pilgrim 1908
G. †Propalaeomeryx Lydekker 1883
G. †Samotherium Forsyth Major 1888
G. †Schansitherium Kilgus 1922
Subfm. †Sivatheriinae Zittel 1893
G. †Umbrotherium Hurzeler and Engesser 1976
G. †Vishnutherium Lydekker 1876
Invalid names: Bohlininae Solounias 2007 [empty], Giraffinae [empty], Sivatheriini Zittel 1893 [empty]
Fm. †Palaeomerycidae Lydekker 1883
+
Subfm. †Aletomerycinae Frick 1937
G. †Amphitragulus Pomel 1847
G. †Germanomeryx Rössner 2010
G. †Heterocemas Young 1937
G. †Hoplitomeryx Leinders 1984
Subfm. †Palaeomerycinae Lydekker 1883
G. †Xenokeryx Sánchez et al. 2015
Fm. †Leptomerycidae Zittel 1893
+
G. †Hendryomeryx Black 1978
+
†Hendryomeryx defordi Wilson 1974
+
Invalid names: Hidrosotherium transpecoensis Wilson 1974 [synonym]
†Hendryomeryx wilsoni Black 1978
Invalid names: Hidrosotherium Wilson 1974 [synonym]
G. †Leptomeryx Leidy 1853
+
†Leptomeryx antecedens Cook 1934
†Leptomeryx blacki Stock 1949
†Leptomeryx evansi Leidy 1853
+
Invalid names: Leptomeryx lenis Cook 1934 [synonym], Trimerodus cedrensis Cope 1873 [synonym]
†Leptomeryx mammifer Cope 1885
†Leptomeryx significans Russell 1954
†Leptomeryx speciosus Lambe 1908
†Leptomeryx yoderi Schlaikjer 1935
Invalid names: Trimerodus Cope 1873 [synonym]
G. †Pronodens Koerner 1940
+
†Pronodens silberlingi Koerner 1940
†Pronodens transmontanus Douglass 1903
G. †Pseudoparablastomeryx Frick 1937
+
†Pseudoparablastomeryx francescita Frick 1937
†Pseudoparablastomeryx scotti Frick 1937
G. †Propalaeoryx Stromer 1926
+
†Propalaeoryx austroafricanus Stromer 1926
†Propalaeoryx nyanzae Whitworth 1958
G. †Walangania Whitworth 1958
+
†Walangania africanus Whitworth 1958
†Walangania gracilis Whitworth 1958
Invalid names: Prodremotheridae Guo et al. 1999 [empty]
G. †Prodremotherium Filhol 1877
+
†Prodremotherium beatrix Pilgrim 1912
†Prodremotherium elongatum Filhol 1877
†Prodremotherium flerowi Trofimov 1957
†Prodremotherium trepidum Gabunia 1964
G. †Sanitherium von Meyer 1866
+
†Sanitherium cingulatum Pilgrim 1926
†Sanitherium masticum Paraskevaidis 1940
†Sanitherium nadiurum Wilkinson 1976
Invalid names: Diamantohyus Stromer 1926 [synonym], Xenochoerus Zdarsky 1909 [synonym]
Subfm. †Sivatherinae Zittel 1853
+
G. †Bramatherium Falconer 1845
+
†Bramatherium megacephalum Pilgrim 1911
†Bramatherium perimense Falconer 1845
Infraor. Tragulina Flower 1883
+
Superfm. †Amphimerycoidea
+
Superfm. †Hypertraguloidea Scott 1940
+
Fm. †Hypertragulidae Cope 1879
+
G. †Nanotragulus Lull 1922
Invalid names: Hypertragulinae [empty], Hypisodinae [empty]
Fm. †Praetragulidae Vislobokova 2001
+
G. †Parvitragulus Emry 1978
G. †Praetragulus Vislobokova 1998
G. †Simimeryx Stock 1934
Fm. †Lophiomerycidae Janis 1987
+
G. †Indomeryx Pilgrim 1928
+
G. †Lophiomeryx Pomel 1853
+
†Lophiomeryx angarae Matthew and Granger 1925
†Lophiomeryx benarensis Gabunia 1964
†Lophiomeryx chalaniati Pomel 1853
†Lophiomeryx kargilensis Nanda and Sahni 1990
†Lophiomeryx minor Lydekker 1885
†Lophiomeryx mouchelini Brunet and Sudre 1987
†Lophiomeryx shinaoensis Miao 1982
†Lophiomeryx turgaicus Flerov 1938
†Lophiomeryx zongi Wang et al. 2025
G. †Nalameryx Métais et al. 2009
+
†Nalameryx savagei Nanda and Sahni 1990
†Nalameryx sulaimani Métais et al. 2009
Fm. Tragulidae Milnes-Edwards 1864 [chevrotain]
+
G. †Afrotragulus Sánchez et al. 2010
+
†Afrotragulus moruorotensis Pickford 2001
†Afrotragulus parvus Whitworth 1958
G. †Cryptomeryx Schlosser 1886
+
†Cryptomeryx decedens Stehlin 1910
†Cryptomeryx gaudryi Filhol 1877
†Cryptomeryx major Schlosser 1886
†Cryptomeryx matsoui Sudre 1984
G. †Dorcatherium Kaup 1839
+
†Dorcatherium bulgaricus Bakalov and Nikilov 1962
†Dorcatherium chappuisi Arambourg 1933
†Dorcatherium crassum Lartet 1851
†Dorcatherium guntianum von Meyer 1846
†Dorcatherium iririensis Pickford 2002
†Dorcatherium jourdani Deperet 1887
†Dorcatherium libiensis Hamilton 1973
†Dorcatherium majus Lydekker 1876
+
Invalid names: Merycopotamus nanus Falconer 1868 [objective synonym]
†Dorcatherium minus Lydekker 1876
†Dorcatherium naui Kaup 1833
†Dorcatherium orientale Qui and Gu 1991
†Dorcatherium peneckei Hoffmann 1892
†Dorcatherium pigotti Whitworth 1958
†Dorcatherium puyhauberti Arambourg and Piveteau 1929
†Dorcatherium rodgeri Hofman 1909
†Dorcatherium vindobonense von Meyer 1846
G. Hyemoschus Gray 1845
+
Hyemoschus aquaticus Ogilby 1841 [water chevrotain]
†Hyomoschus crassus Lartet 1837
Invalid names: Hyemoschus aquaticus Ogilby 1841 [valid], Hyomoschus crassus Lartet 1837 [valid]
G. Hyemoschus Gray 1845
+
Hyemoschus aquaticus Ogilby 1841 [water chevrotain]
†Hyomoschus crassus Lartet 1837
Invalid names: Hyemoschus aquaticus Ogilby 1841 [valid], Hyomoschus crassus Lartet 1837 [valid]
G. †Iberomeryx Gabunia 1964
+
†Iberomeryx miaoi Mennecart et al. 2021
†Iberomeryx minor Filhol 1882
+
Invalid names: Iberomeryx matsoui Heissig 1987 [synonym], Iberomeryx minus Sudre and Blondel 1996 [synonym]
†Iberomeryx parvus Gabunia 1964
G. †Krabimeryx Metais et al. 2001
+
†Krabimeryx gracilis Miao 1982
†Krabimeryx primitivus Metais et al. 2001
G. †Siamotragulus Thomas et al. 1990
+
†Siamotragulus bugtiensis Ginsburg 2001
†Siamotragulus haripounchai Mein and Ginsburg 1997
†Siamotragulus sanyathanai Thomas et al. 1990
†Siamotragulus songhorensis Whitworth 1958
G. Tragulus Brisson 1762 [mouse deer]
+
Tragulus fulviventer Gray 1836
Tragulus javanicus Osbeck 1765 [Java mouse-deer]
+
Invalid names: Tragulus fuscatus Blyth 1858 [synonym]
Tragulus kanchil Raffles 1822 [lesser mouse-deer]
Tragulus napu Cuvier 1822 [greater mouse-deer]
Tragulus nigricans Thomas 1892 [Philippine mouse-deer]
†Tragulus sivalensis Lydekker 1882
Tragulus stanleyanus Gray 1836
Invalid names: Meminna Gray 1836 [synonym], Tragulus malaccensis Gray 1843 [nomen nudum], Tragulus mimenoides Hodgson 1844 [nomen nudum]
Invalid names: Geolocinae Schlosse 1886 [empty], Tragulinae Milnes-Edwards 1864 [empty]
Superfm. †Traguloidea
+
Fm. †Archaeomerycidae Simpson 1945
+
G. †Archaeomeryx Matthew and Granger 1925
G. †Irrawadymeryx Métais et al. 2007
G. †Miomeryx Matthew and Granger 1925
G. †Notomeryx Qiu 1978
G. †Paukkaungmeryx Ducrocq et al. 2020
G. †Xinjiangmeryx Zheng 1978
Fm. †Gelocidae Schlosser 1886
+
G. †Gobiomeryx Trofimov 1957
G. †Paragelocus Schlosser 1902
G. †Phaneromeryx Schlosser 1886
Subfm. †Pseudoceratinae Frick 1937
G. †Pseudogelocus Schlosser 1902
G. †Pseudomeryx Trofimov 1957
Invalid names: Anoplotheriadae Gray 1821 [empty], Giraffoidea Gray 1821 [empty], Hispanomerycidae Moya 1986 [empty], Ruminantes Gray 1821 [synonym]
Unr. Ruminantiamorpha Spaulding et al. 2009
+
Fm. †Protoceratidae Marsh 1891 [protoceratid]
+
G. †Leptoreodon Wortman 1898
+
†Leptoreodon edwardsi Stock 1936
†Leptoreodon golzi Ludtke and Prothero 2004
†Leptoreodon leptolophus Golz 1976
†Leptoreodon major Golz 1976
†Leptoreodon marshi Wortman 1898
+
Invalid names: Camelomeryx longiceps Scott 1898 [synonym], Merycodesmus gracilis Scott 1898 [synonym]
†Leptoreodon pusillus Golz 1976
†Leptoreodon stocki Kelly 1990
Invalid names: Camelomeryx Scott 1898 [synonym], Leptoreodon (Hesperomeryx) Stock 1936 [invalid subgroup], Merycodesmus Scott 1898 [synonym]
G. †Leptotragulus Scott and Osborn 1887
+
†Leptotragulus clarki Gazin 1955
†Leptotragulus medius Peterson 1919
†Leptotragulus proavus Scott and Osborn 1887
+
Invalid names: Parameryx laevis Marsh 1894 [synonym], Parameryx sulcatus Marsh 1894 [synonym]
†Leptotragulus ultimus Schlaikjer 1935
Invalid names: Parameryx Marsh 1894 [synonym]
G. †Poabromylus Peterson 1932
+
†Poabromylus golzi Black 1978
†Poabromylus kayi Peterson 1932
†Poabromylus robustus Stock 1949
Subfm. †Protoceratinae
+
G. †Paratoceras Frick 1937
+
†Paratoceras coatesi Rincón et al. 2015
†Paratoceras macadamsi Frick 1937
†Paratoceras orarius Rincón et al. 2015
†Paratoceras tedfordi Webb et al. 2003
†Paratoceras wardi Patton and Taylor 1973
G. †Protoceras Marsh 1891
+
†Protoceras celer Marsh 1891
†Protoceras comptus Marsh 1894
†Protoceras nasutus Marsh 1897
†Protoceras neatodelpha Patton and Taylor 1973
†Protoceras skinneri Patton and Taylor 1973
G. †Pseudoprotoceras Cook 1934
+
†Pseudoprotoceras longinaris Cook 1934
†Pseudoprotoceras minor Wilson 1974
†Pseudoprotoceras taylori Emry and Storer 1981
Subfm. †Synthetoceratinae Frick 1937
+
G. †Synthetoceras Stirton 1932
+
Subg. †Synthetoceras (Prosynthetoceras) Frick 1937
†Synthetoceras davisorum Hulbert and Whitmore 2006
†Synthetoceras tricornatus Stirton 1932
G. †Trigenicus Douglass 1903
+
Invalid names: Leptotragulinae [invalid subgroup], Syndyoceratinae [empty]
Subor. Suiformes Jaeckel 1911
+
Invalid names: Ancodonta Matthew [invalid subgroup], Anthracotherioidea Gill 1872 [empty], Hyodonta Matthew 1929 [invalid subgroup], Palaeodonta Matthew [synonym], Tesserachenae Gray 1821 [synonym]
Superfm. Suoidea Gray 1821
+
G. †Palaeochoerus Pomel 1847
+
†Palaeochoerus gergovianus De Blainville 1846
†Palaeochoerus minus Gope Posse 1972
†Palaeochoerus platyops Cope 1881
†Palaeochoerus pusilus Ginsburg 1974
†Palaeochoerus typus Pomel 1847
G. †Schizochoerus Crusafont-Pairo and Lavocat 1954
+
†Schizochoerus anatoliensis van der Made 1997
†Schizochoerus arambourgi Ozansoy 1965
†Schizochoerus sinapensis van der Made 1997
†Schizochoerus vallesiensis Crusafont and Lavocat 1954
Fm. Suidae Gray 1821 [pig]
+
Subfm. †Cainochoerinae Pickford 1995
+
G. †Albanohyus Ginsburg 1974
+
†Albanohyus castellensis Golpe Posse 1977
†Albanohyus pygmaeus Deperet 1892
G. †Cainochoerus Pickford 1988
G. †Dinochoerus Peterson 1905
G. †Hemichoerus Filhol 1882
G. †Hippopotamodon Lydekker 1877
+
†Hippopotamodon erymanthius Roth and Wagner 1854
Invalid names: Dicoryphochoerus Pilgrim 1926 [synonym], Limnostonyx Ginsburg 1988 [synonym]
G. †Hyosus Pilgrim 1926
Subfm. †Hyotheriinae Cope 1888
+
G. †Chicochoerus Orliac et al. 2006
G. †Chleuastochoerus Pearson 1928
G. †Cleuastochoerus Pearson 1928
G. †Hyotherium von Meyer 1834
+
†Hyotherium lacaeillei Orliac et al. 2006
†Hyotherium major Pomel 1847
†Hyotherium meissneri von Meyer 1850
†Hyotherium platyops Cope 1881
†Hyotherium shanwangense Liping et al. 2002
†Hyotherium soemmeringi von Meyer 1834
Invalid names: Hypotherium Ginsburg et al. 1991 [synonym]
Subfm. †Kubanochoerinae Gabunia 1958
+
G. †Kenyasus Pickford 1986
+
†Kenyasus kirimunensis Pickford 2007
†Kenyasus namaquensis Pickford and Senut 1997
†Kenyasus rusingensis Pickford 1986
G. †Megalochoerus Picford 1993
+
†Megalochoerus humongous Pickford 1993
†Megalochoerus khinzikebirus Wilkinson 1976
G. †Kubanochoerus Gabunia 1955
+
†Kubanochoerus gigas Pearson 1928
†Kubanochoerus khinzikebirus Wilkinson 1976
†Kubanochoerus marymuunguae Made 1996
†Kubanochoerus massai Arambourg 1961
Subfm. †Listriodontinae Simpson 1945
+
G. †Bunolistriodon Arambourg 1933
+
†Bunolistriodon minheensis Qiu et al. 1981
Invalid names: Eurolistriodon Pickford and Moya Sola 1995 [synonym]
G. †Listriodon von Meyer 1846
+
†Listriodon akatidogus Wilkinson 1976
†Listriodon akatikubas Wilkinson 1976
†Listriodon bartulensis Pickford 2001
†Listriodon bonaerensis Ameghino 1904
†Listriodon chapalmalensis Ameghino 1908
†Listriodon dukkar van der Made et al. 2022
†Listriodon jeanneli Arambourg 1943
†Listriodon juba Ginsburg 1977
†Listriodon lishanensis Lee and Wu 1978
†Listriodon lockharti Pomel 1848
†Listriodon pentapotamiae Falconer 1870
†Listriodon splendens von Meyer 1846
†Listriodon tarijensis Ameghino 1904
G. †Lopholistriodon Pickford and Wilkinson 1975
+
†Lopholistriodon kidogosana Pickford and Wilkinson 1975
†Lopholistriodon moruoroti Wilkinson 1976
†Lopholistriodon pickfordi Van der Made 1996
G. †Lophochoerus Pilgrim 1926
G. †Mabokopithecus von Koenigswald 1969
G. †Microstonyx Pilgrim 1926
+
†Microstonyx binxianensis Liu et al. 1978
†Microstonyx erymanthius Roth and Wagner 1854
†Microstonyx major Gervais 1848
G. †Paradoxodonides Strand 1943
G. †Promesochoerus Leakey 1965
Subfm. Suinae Gray 1821
+
Tr. †Hippohyini Thenius 1970
+
G. †Hippohyus Falconer and Cautley 1847
G. †Sinohyus von Koenigswald 1963
G. †Sivahyus Pilgrim 1926
G. Hylochoerus Thomas 1904 [giant forest hog]
+
†Hylochoerus euilus Hopwood 1926
Hylochoerus meinertzhageni Thomas 1904 [giant forest hog]
G. †Kolpochoerus van Hoepen and van Hoepen 1932
+
†Kolpochoerus afarensis Cooke 1978
†Kolpochoerus cookei Brunet and White 2001
†Kolpochoerus denheinzelini Brunet and White 2001
†Kolpochoerus evronensis Haas 1970
†Kolpochoerus limnetes Hopwood 1926
†Kolpochoerus majus Hopwood 1934
†Kolpochoerus maroccanus Ennouchi 1953
†Kolpochoerus olduvaiensis Leakey 1942
†Kolpochoerus phillipi Souron et al. 2015
G. †Metridiochoerus Hopwood 1926
+
†Metridiochoerus andrewsi Hopwood 1926
†Metridiochoerus compactus Van Hoepen and Van Hoepen 1932
†Metridiochoerus hopwoodi Leakey 1958
†Metridiochoerus modestus van Hoepen and van Hoepen 1932
†Metridiochoerus shawi Dale 1948
Invalid names: Tapinochoerus van Hoepen and van Hoepen 1932 [synonym]
Tr. †Phacochoerini Gray 1868
+
G. †Potamochoeroides Dale 1948
G. †Stylochoerus van Hoepen and van Hoepen 1932
G. Phacochoerus Cuvier 1826 [warthog]
+
Phacochoerus aethiopicus Pallas 1766 [desert warthog]
Phacochoerus africanus Gmelin 1788 [common warthog]
G. Potamochoerus Gray 1854
+
Potamochoerus larvatus Cuvier 1822
Potamochoerus porcus Linnaeus 1758 [red river hog]
Invalid names: Choiropotamus Gray 1852 [replaced]
G. Sus Linnaeus 1758 [pig]
+
†Sus antediluvianus Kaup 1833
†Sus arvernensis Deperet 1885
Sus barbatus Müller 1838 [bearded pig]
†Sus brachygnathus Dubois 1908
Sus bucculentus Heude 1892 [Vietnam warty pig]
Sus cebifrons Heude 1888 [Visayan warty pig]
Sus celebensis Müller and Schlegel 1843 [Sulawesi pig]
Sus cristatus Wagner 1839
†Sus falconeri Lydekker 1884
†Sus giganteus Falconer and Cautley 1847
†Sus hyotherioides Schlosser 1903
†Sus hysudricus Falconer and Cautley 1847
†Sus lydekkeri Zdansky 1928
†Sus microdon Schlosser 1903
†Sus palaeochoerus Kaup 1833
Sus philippensis Nehring 1886 [Philippine warty pig]
Sus scrofa Linnaeus 1758 [wild boar]
†Sus sivalensis Falconer and Cautley 1847
†Sus stehlini Schlosser 1903
†Sus strozzii Forsyth Major 1881
Sus verrucosus Müller 1840 [Javan warty pig]
Sus vittatus Müller and Schlegel 1842 [banded pig]
†Sus waylandi Cooke and Coryndon 1970
Invalid names: Babyrousini Gray 1868 [empty]
Subfm. †Tetraconodontinae Lydekker 1886
+
G. †Conohyus Pilgrim 1926
+
†Conohyus grivensis Gaillard 1899
†Conohyus huenermanni Heissig 1989
†Conohyus olujici Bernor and Radovcic 2004
†Conohyus simorrensis Lartet 1851
†Conohyus sindiensis Lydekker 1884
†Conohyus steinheimensis Fraas 1870
Invalid names: Adaetontherium Lewis 1934 [synonym]
G. †Lophocoerus Pilgrim 1926
G. †Notochoerus Broom 1925
+
†Notochoerus capensis Broom 1925
†Notochoerus clarkei Suwa and White 2004
G. †Nyanzachoerus Leakey 1958
+
†Nyanzachoerus jaegeri Coppens 1971
†Nyanzachoerus kanamensis Leakey 1958
†Nyanzachoerus khinzir Boisserie et al. 2014
†Nyanzachoerus nakaliensis Tsubamoto et al. 2020
G. †Parachleuastochoerus Golpe-Posse 1972
+
†Parachleuastochoerus crusafonti Golpe-Posse 1972
†Parachleuastochoerus huenermanni Heissig 1989
†Parachleuastochoerus kretzoii Fortelius et al. 2005
†Parachleuastochoerus valentini Filhol 1882
G. †Tetraconodon Falconer 1868
+
†Tetraconodon magnum Falconer 1868
†Tetraconodon malensis Htike et al. 2005
Invalid names: Babyrousinae Thenius 1970 [empty], Chaeropotamina Bonaparte 1850 [empty], Phacochoeridae Gray 1868 [synonym]
G. †Taucanamo Simpson 1945
+
†Taucanamo grandaevum Fraas 1870
†Taucanamo inonuensis Pickford and Ertürk 1979
†Taucanamo primum van der Made 1997
†Taucanamo sansaniense Lartet 1851
Invalid names: Palaeochoeridae Matthew 1924 [invalid subgroup], Sanitheriidae Simpson 1945 [empty], Suina Gray 1868 [invalid subgroup], Suinamorpha Spaulding et al. 2009 [synonym]
Fm. †Tapirulidae Cope 1879
+
G. †Obotherium Bai et al. 2023
+
†Obotherium parvum Bai et al. 2023
†Obotherium tongi Bai et al. 2023
G. †Tapirulus Gervais 1850
+
†Tapirulus depereti Stehlin 1910
†Tapirulus hyracinus Gervais 1850
†Tapirulus majori Stehlin 1910
†Tapirulus perrierensis Sudre 1978
†Tapirulus schlosseri Stehlin 1910
G. †Tragulohyus Gervais 1874
Subor. Tylopoda Illiger 1811
+
Superfm. †Anoplotherioidea Bonaparte 1850
+
Fm. †Anoplotheriidae Gray 1821
+
Subfm. †Anoplotheriinae Gray 1821
+
G. †Anoplotherium Cuvier 1804
+
Invalid names: Eurytherium Gervais 1852 [synonym], Hoplotherium Meyer 1841 [synonym], Oplotherium Laizer and Parieu 1838 [synonym], Pleregnathus Laizer and Parieu 1838 [synonym]
G. †Diplobune Rutimeyer 1862
+
Invalid names: Hyracodon Filhol 1876 [synonym], Hyracodontherium Filhol 1877 [synonym], Hyrocodon Filhol 1873 [synonym]
G. †Duerotherium Cuesta and Badiola 2009
G. †Ephelcomenus Hurzeler 1938
G. †Robiatherium Sudre 1988
Invalid names: Anoplotheriina Bonaparte 1850 [empty], Robiaciinae Sudre 1977 [empty]
Fm. †Xiphodontidae Flower 1883
+
G. †Dichodon Owen
+
†Dichodon cartieri Rutimeyer 1891
†Dichodon cervinus Owen 1848
†Dichodon cuspidatus Owen 1847
†Dichodon frohnstettense von Meyer 1852
†Dichodon frohnstettensis Meyer 1852
†Dichodon simplex Kowalevsky 1874
G. †Haplomeryx Schlosser 1886
+
†Haplomeryx egerkingensis Stehlin 1910
†Haplomeryx euzetensis Depéret 1917
†Haplomeryx obliquus Cuvier 1822
†Haplomeryx picteti Stehlin 1910
†Haplomeryx zitteli Schlosser 1886
G. †Leptotheridium Stehlin 1910
+
†Leptotheridium lugeoni Stehlin 1910
†Leptotheridium traguloides Stehlin 1910
G. †Paraxiphodon Sudre 1978
+
†Paraxiphodon cournovensis Sudre 1978
†Paraxiphodon teulonensis Sudre 1978
G. †Xiphodon Cuvier 1822
+
†Xiphodon castrensis Kowalevsky 1873
†Xiphodon gracile Cuvier 1822
†Xiphodon intermedium Stehlin 1910
Fm. †Oromerycidae Gazin 1955
+
G. †Merycobunodon Golz 1976
+
†Merycobunodon littoralis Golz 1976
†Merycobunodon walshi Murphey and Kelly 2017
G. †Protylopus Wortman 1898
+
†Protylopus annectens Peterson 1919
†Protylopus pearsonensis Golz 1976
†Protylopus petersoni Wortman 1898
+
Invalid names: Protylopus minor Prothero 1998 [synonym], Protylopus parvus Prothero 1998 [synonym]
†Protylopus robustus Golz 1976
†Protylopus stocki Golz 1976
Invalid names: Camelidamorpha Spaulding et al. 2009 [synonym], Cameloidea [empty], Hydrophorae Gray 1821 [synonym], Oreodontoidea Gill 1872 [empty], Xiphodontoidea Flower 1883 [empty]
Invalid names: Antiacodontidae Krishtalka and Stucky 1985 [empty], Bunomerycidae Gentry and Hooker 1988 [empty], Cetartiodactyla Montgelard et al. 1997 [synonym], Eurytheriidae Cope 1889 [empty], Haplobunodontidae Sudre 1978 [empty], Leptochoerinae [empty], Merycoidodontoidea [empty], Neoselenodontia Webb and Taylor 1980 [empty], Nonruminantia Gregory 1910 [empty], Selenodontia Scopoli 1777 [empty], Trigonolestidae [empty]
Diagnosis
No diagnoses are available