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Polyphaga
Taxonomy
Polyphaga was named by Emery (1886). It is extant.
It was assigned to Coleoptera by Crowson (1981), Carpenter (1992), Lawrence and Newton (1995), Bouchard et al. (2011); to Metacoleoptera by Boudinot et al. (2022); and to Eucoleoptera by Cai et al. (2022).
It was assigned to Coleoptera by Crowson (1981), Carpenter (1992), Lawrence and Newton (1995), Bouchard et al. (2011); to Metacoleoptera by Boudinot et al. (2022); and to Eucoleoptera by Cai et al. (2022).
Subtaxa
Agyrtes primoticus, Angarolarva, Bostrichiformia, Bostrychiformia, Clambiformia, Cryptophagites, Cucujiformia, Derodontiformia, Elateriformia, Elmadulescens, Escheria crassipunctata, Escheria dimidiata, Escheria punctulata, Eucinetiformia, Jingxidiscus (syn. Discus), Jurodidae (syn. Sikhotealiniidae), Larvula, Lasiosynidae, Limnius perrini, Malachius vertummni, Melanocantharis, Mesosperchus notatus, Mormolucoides, Nosodendriformia, Plastelater, Polysitum, Potamophilites, Ranis, Rhinorhipiformia, Rhysopsalis, Scirtiformia, Sphaerocantharis (syn. Theornithion, Mikrocarpides), Staphyliniformia (syn. Scarabaeiformia), Syntomopterus, Tomocoleites, Troglops punctatissima
Synonyms
|
Synonymy list
| Year | Name and author |
|---|---|
| 1847 | Malachius vertummni Heer p. 150 figs. Pl 5, figs 1,2 |
| 1861 | Derodontiformia Le Conte |
| 1886 | Polyphaga Emery |
| 1891 | Escheria punctulata Förster p. 361 figs. Pl 11, figs 8a,b |
| 1891 | Escheria dimidiata Förster p. 363 figs. Pl 11, figs 10a,b |
| 1891 | Escheria crassipunctata Förster p. 364 figs. Pl 11, fig 11 |
| 1900 | Agyrtes primoticus Scudder p. 45 figs. Pl 5, fig 6 |
| 1935 | Troglops punctatissima Théobald p. 75 figs. Pl 5, fig 18 |
| 1937 | Hydrobius crassipunctatus Theobald p. 169 figs. Pl XII fig 28 |
| 1937 | Limnius perrini Theobald p. 284 figs. Pl XX fig 1 |
| 1977 | Mesosperchus notatus Ponomarenko p. 110 figs. Pl 9, fig 5; text-fig 59 |
| 1981 | Polyphaga Crowson p. 694 |
| 1992 | Polyphaga Carpenter |
| 1995 | Polyphaga Lawrence and Newton, Jr. |
| 2002 | Tomocoleites Hong p. 147 |
| 2002 | Tomocoleites longigastrulatus Hong p. 147 figs. 2.6.126-135 |
| 2011 | Derodontiformia Bouchard et al. |
| 2011 | Polyphaga Bouchard et al. |
| 2020 | Agyrtes primoticus Strelnikova et al. p. 627 |
| 2022 | Polyphaga Boudinot et al. |
| 2022 | Polyphaga Cai et al. |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Subor. Polyphaga Emery 1886
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G. †Angarolarva Ponomarenko 1985
+
†Angarolarva aquatica Ponomarenko 1985
†Angarolarva incerta Ponomarenko 1990
Infraor. Bostrichiformia Forbes 1926
+
Superfm. Bostrichoidea Latreille 1802
+
Fm. Bostrichidae Latreille 1802 [false powderpost beetle]
+
Subfm. †Alitrepaninae Peng et al. 2022
+
Subfm. Bostrichinae Latreille 1802
+
Tr. Apatini Billberg 1820
Tr. Bostrichini Latreille 1802
G. †Burmostrichus Háva and Zahradník 2025
G. †Discoclavata Poinar 2013
Tr. Xyloperthini Lesne 1921
Subfm. Dinoderinae Thomson 1863
+
G. Dinoderus Stephens 1830
G. †Elongatus Wang et al. 2025
G. Stephanopachys Waterhouse 1888
Invalid names: Dysidinae Lesne 1921 [empty], Euderiinae Lesne 1934 [empty], Lyctidae [replaced], Psoinae Blanchard 1851 [empty]
Fm. Dermestidae Latreille 1804 [skin beetle]
+
Subfm. Dermestinae Latreille 1804
+
Tr. Dermestini Latreille 1804
Tr. †Paradermestini Deng et al. 2017
Invalid names: Marioutini Jacobson 1913 [empty], Thorictini Agassiz 1846 [empty]
Subfm. Orphilinae Le Conte 1861
+
G. †Nothattagenus Li and Cai 2022
Tr. Orphilini Le Conte 1860
Tr. Ranolini Háva 2014
Subfm. Trinodinae Casey 1900
+
Tr. †Cretonodini Kirejtshuk and Azar 2009
Tr. Trinodini Casey 1900
Invalid names: Thylodriini Semenov-Tian-Shanskij 1913 [empty]
G. †Tryoniopsis Dunstan 1923
+
†Tryoniopsis granulata Dunstan 1923
†Tryoniopsis punctata Dunstan 1923
Fm. Ptinidae Latreille 1802 [spider beetle]
+
Subfm. Anobiinae Fleming 1821
+
G. Actenobius Fall 1905
Tr. Anobiini Fleming 1821
G. †Gastrallanobium Wickham 1914
Tr. Gastrallini White 1982
Tr. Hadrobregmini White 1982
G. Nicobium LeConte 1861
Tr. Stegobiini White 1982
Invalid names: Anobium jacquelinae Hawkeswood et al. 2009 [nomen dubium], Nicobiini White 1982 [invalid subgroup]
Subfm. Dorcatominae Thomson 1859
+
G. Caenocara Thomson 1859
Tr. Dorcatomini Thomson 1859
G. Stagetus Wollaston 1861
G. †Venablesia Britton 1960
Invalid names: Petaliini White 1982 [invalid subgroup]
Subfm. Ptininae Latreille 1802
+
G. †Electrognostus Philips and Mynhardt 2011
G. †Okamninus Mynhardt and Philips 2013
Tr. Ptinini Latreille 1802
G. †Xyletinites von Heyden and von Heyden 1866
+
†Xyletinites tumbicola von Heyden and von Heyden 1866
Invalid names: Alvarenganiellinae Viana and Martinez 1971 [empty], Anobiidae Fleming 1821 [synonym], Ptilininae Shuckard 1840 [empty]
Invalid names: Endecatomidae LeConte 1861 [empty]
Unr. Clambiformia Cai and Tihelka 2022
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Superfm. Clamboidea Fischer von Waldheim 1821
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Fm. Clambidae Fischer von Waldheim 1821 [minute beetle]
+
Subfm. Clambinae Fischer von Waldheim 1821
+
G. Clambus Fischer von Waldheim 1821
G. †Eoclambus Kirejtshuk and Azar 2008
G. Sphaerothorax Endrödy-Younga 1959
Invalid names: Calyptomerinae Crowson 1955 [empty], Ptismidae Kirejtshuk and Azar 2016 [synonym]
Fm. Derodontidae LeConte 1861 [tooth-necked fungus beetle]
+
Invalid names: Derodontinae LeConte 1861 [empty]
Fm. Eucinetidae Lacordaire 1857 [plate-thigh beetle]
+
G. Eucinetus Germar 1818
+
†Eucinetus debilispinus Li and Cai 2024
†Eucinetus nikolaevae Kirejtshuk and Ponomarenko 2019
†Eucinetus panghongae Li and Cai 2024
†Eucinetus parvus Du et al. 2020
†Eucinetus zhenhuai Li and Cai 2024
G. †Huaxiacinectus Hong 1995
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†Huaxiacinectus xiangfanggouensis Hong 1995
†Huaxiacinectus xinyaoensis Hong 1995
Fm. †Mesocinetidae Kirejtshuk and Ponomarenko 2010
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G. †Manoelodes Kirejtshuk and Ponomarenko 2010
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†Manoelodes gratiosus Kirejtshuk and Ponomarenko 2010
G. †Manopsis Kirejtshuk and Ponomarenko 2010
+
†Manopsis concavicollis Kirejtshuk and Ponomarenko 2010
G. †Mesocinetus Ponomarenko 1986
+
†Mesocinetus aequalis Kirejtshuk and Ponomarenko 2010
†Mesocinetus angustitibialis Kirejtshuk and Ponomarenko 2010
†Mesocinetus elumbis Kirejtshuk and Ponomarenko 2010
†Mesocinetus mongolicus Ponomarenko 1986
†Mesocinetus ovatus Ponomarenko 1990
†Mesocinetus subjectus Kirejtshuk and Ponomarenko 2010
G. †Parashartegus Kirejtshuk and Ponomarenko 2010
+
†Parashartegus distinctus Kirejtshuk and Ponomarenko 2010
G. †Shartegus Kirejtshuk and Ponomarenko 2010
+
†Shartegus transversus Kirejtshuk and Ponomarenko 2010
Invalid names: Derodontoidea LeConte 1861 [synonym], Eucinetoidea Lacordaire 1857 [synonym]
G. †Cryptophagites Ponomarenko 1990
+
†Cryptophagites capitatus Ponomarenko 1990
†Cryptophagites elongatus Ponomarenko 1990
Infraor. Cucujiformia Lameere 1938
+
Superfm. Cleroidea Latreille 1802
+
Fm. Biphyllidae Leconte 1861 [false skin beetle]
+
G. †Paleobiphyllus Makarov and Perkovsky 2019
+
†Paleobiphyllus ponomarenkoi Makarov and Perkovsky 2019
Fm. Byturidae Gistel 1848
+
Invalid names: Byturinae Gistel 1848 [empty], Platydascillinae Pic 1914 [empty]
Fm. Cleridae Latreille 1802 [checkered beetle]
+
Subfm. Clerinae Latreille 1802
+
G. †Arawakis Opitz 2007
Tr. Clerini Latreille 1802
Tr. Hydnocerini Spinola 1844
G. †Mitrandiria Kolibáč 1997
Unr. Opilonitae Gistel 1848
G. †Zahradnikius Winkler 1992
Invalid names: Clerus piacentinii Piton 1940 [nomen dubium], Clerus succini Giebel 1862 [nomen dubium], Hydnocerinae Spinola 1844 [synonym], Phyllobaeninae Lacordaire 1857 [synonym]
Subfm. Korynetinae Laporte 1836
+
G. †Aberrokorynetes Winkler 1990
G. Epiphloeus Spinola 1841
G. Necrobia Olivier 1795
G. Prosymnus Laporte 1836
G. †Visokorynetes Winkler 1990
Invalid names: Enopliinae Gistel 1856 [synonym], Epiphloeinae Kuwert 1893 [synonym], Tarsosteninae Jacquelin du Val 1860 [synonym]
Subfm. Tillinae Fischer von Waldheim 1813
+
G. †Aiytillus Kolibáč and Perkovsky 2024
G. †Bilbotillus Kolibáč 1997
G. Cymatodera Gray 1832
G. †Smudlotillus Kolibáč 1997
G. Strotocera Schenkling 1903
G. Tillus Olivier 1790
Invalid names: Clerus adonis Heer 1847 [nomen dubium], Tanocleria Hong 2002 [nomen nudum], Trichodes stebingeri Cockerell 1936 [nomen dubium]
Fm. Lophocateridae Crowson 1964
+
G. Antillipeltis Lawrence et al. 2014
+
†Antillipeltis alleni Lawrence et al. 2014
†Antillipeltis iviei Lawrence et al. 2014
G. †Burmacateres Kolibáč and Peris 2020
+
†Burmacateres longicoxa Kolibáč and Peris 2020
+
Invalid names: Mesolophocateres pengweii Yu et al. 2020 [synonym]
Invalid names: Mesolophocateres Yu et al. 2020 [synonym]
G. †Cervicatinius Tan and Ren 2007
+
†Cervicatinius complanus Tan and Ren 2007
Invalid names: Sinopeltis Yu et al. 2012 [synonym]
G. Grynocharis Thomson 1859
G. Lophocateres Olliff 1883
Invalid names: Ancyronini Reitter 1876 [invalid subgroup], Lophocaterini Crowson 1964 [invalid subgroup]
Fm. Melyridae Leach 1815 [soft-winged flower beetle]
+
Subfm. Dasytinae Laporte 1840
+
Tr. Danaceini Thomson 1859
Tr. Dasytini Laporte 1840
Tr. Eparchiini Bernhauer 1934
G. †Eudasytites Wickham 1912
Tr. Listrini Majer 1990
G. †Protodasytes Tihelka et al. 2021
Subfm. Malachiinae Fleming 1821
+
Tr. Colotini Abeille de Perrin 1890
Tr. Malachiini Fleming 1821
Tr. †Palpattalini Tshernyshev 2020
Tr. Troglopini Mulsant and Rey 1867
Invalid names: Attalini Abeille de Perrin 1891 [empty]
Fm. Peltidae Kirby 1837 [shield beetle]
+
G. Peltis Müller 1764
+
†Peltis antehercynica Kolibáč et al. 2016
†Peltis constulata von Heyden 1862
†Peltis costulata von Heyden 1862
†Peltis laminata Wickham 1910
†Peltis minuscula Piton 1935
†Peltis tricostata Heer 1847
Invalid names: Peltinae Kirby 1837 [invalid subgroup], Peltini Latreille 1806 [invalid subgroup], Phenolia incapax Scudder 1876 [nomen dubium]
Fm. Prionoceridae Lacordaire 1857
+
Fm. Rhadalidae LeConte 1861
+
G. Xamerpus Fairmaire 1886
Invalid names: Gietellinae Constantin and Menier 1987 [empty]
Fm. Thanerocleridae Chapin 1924
+
Subfm. Zenodosinae Kolibáč 1992
+
G. †Archaeozenodosus Yu and Kolibáč 2017
G. †Cretozenodosus Cai and Huang 2018
G. †Mesozenodosus Tihelka et al. 2020
G. †Thanerosus Peris and Kolibáč 2022
Invalid names: Thaneroclerinae Chapin 1924 [empty]
Fm. Thymalidae Leveille 1888
+
Invalid names: Decamerinae Crowson 1964 [empty], Thymalinae Leveille 1888 [empty]
Fm. Trogossitidae Latreille 1802 [bark gnawing beetle]
+
Subfm. Lophocaterinae Crowson 1964
+
G. †Cretamerus Peris et al. 2014
G. †Yixianteres Yu et al. 2015
G. †Neolatitergum Zhang 2021
+
†Neolatitergum glabrum Yu et al. 2014
Invalid names: Latitergum Yu et al. 2014 [replaced]
Subfm. Trogossitinae Latreille 1802
+
G. Gymnocheilis Dejean 1835
Tr. †Lithostomatini Kolibáč and Huang 2008
G. Seidlitzella Jakobson 1915
G. Temnoscheila Westwood 1830
G. Tenebroides Piller and Mitterpacher 1783
+
Invalid names: Trogossita Olivier 1790 [objective synonym]
Invalid names: Trogossitini Latreille 1802 [invalid subgroup]
Invalid names: Calitinae Winkler 1924 [empty], Ergoliinae Lacordaire 1854 [empty], Larinotinae Slipinski 1992 [empty], Ostomatidae Reitter 1882 [synonym], Temnochilidae [replaced]
Invalid names: Chaetosomatidae Crowson 1952 [empty], Phloiophilidae Kiesenwetter 1863 [empty], Phycosecidae Crowson 1952 [empty], Protopeltidae Crowson 1966 [empty], Rentoniidae Crowson 1966 [empty]
Superfm. Coccinelloidea Latreille 1807
+
Fm. Anamorphidae Strohecker 1953
+
G. †Kleinzaches Alekseev and Tomaszewska 2018
+
†Kleinzaches germanicianus Alekseev and Tomaszewska 2018
G. †Palaecoryphus Alekseev and Tomaszewska 2018
+
†Palaecoryphus viktori Alekseev and Tomaszewska 2018
G. †Palaeosymbius Arriaga‐Varela et al. 2024
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†Palaeosymbius groehni Arriaga‐Varela et al. 2024
†Palaeosymbius mesozoicus Arriaga‐Varela et al. 2024
Fm. Bothrideridae Erichson 1845
+
Fm. Coccinellidae Latreille 1807 [ladybird beetle]
+
Subfm. Coccinellinae Latreille 1807
+
Tr. Chilocorini Mulsant 1846
Tr. Coccidulini Mulsant 1846
Tr. Epilachnini Mulsant 1846
Tr. Scymnini Mulsant 1846
Tr. Sticholotidini Weise 1901
Invalid names: Coccidulinae Mulsant 1846 [synonym], Scymninae Mulsant 1846 [synonym]
Invalid names: Chilocorinae Mulsant 1846 [empty]
Fm. Corylophidae Le Conte 1852 [minute hooded beetle]
+
Subfm. Corylophinae LeConte 1852
+
Tr. Corylophini Le Conte 1852
Tr. Orthoperini Jacquelin du Val 1857
Tr. Parmulini Poey 1854
Tr. †Xenostanini Li et al. 2022
Invalid names: Peltinodinae Paulian 1950 [empty], Sericoderinae Matthews 1888 [empty]
Fm. Discolomatidae Horn 1878
+
Invalid names: Cephalophaninae John 1954 [empty], Discolomatinae Horn 1878 [empty], Notiophyginae Jacobson 1915 [empty], Pondonatinae John 1954 [empty]
Fm. Endomychidae Leach 1815 [handsome fungus beetle]
+
Subfm. Leiestinae Thomson 1863
+
G. †Burmalestes Tomaszewska 2018
G. †Cretolestes Tomaszewska et al. 2018
G. †Glesirhanis Shockley and Alekseev 2014
G. Leiestes Chevrolat 1836
G. †Palaeoestes Kirejtshuk and Nel 2009
G. †Phymaphoroides Motschulsky 1856
G. Trycherus Gerstaecker 1857
Subfm. Lycoperdininae Redtenbacher 1844
+
G. Eumorphus Weber 1801
G. Gerstaeckerus Tomaszweska 2005
+
Invalid names: Engonius Gestaecker 1857 [replaced], Miocarabus Hong 1984 [synonym]
Subfm. Merophysiinae Seidlitz 1872
+
G. †Cretaparamecus Tomaszewska et al. 2018
Invalid names: Holoparamecinae Seidlitz 1888 [synonym]
Invalid names: Epipocinae Gorham 1873 [empty]
Fm. Latridiidae Erichson 1842 [minute brown scavenger beetle]
+
Subfm. Corticariinae Curtis 1829
+
G. Corticaria Marsham 1802
G. †Corticariites Kirejtshuk 2019
G. Corticarina Reitter 1881
G. Melanophthalma Motschulsky 1866
G. †Succinimontia Zherikhin 1977
Subfm. Latridiinae Erichson 1842
+
G. †Archelatrius Kirejtshuk and Azar 2009
G. Cartodere Thomson 1859
G. Dienerella Reitter 1911
G. †Dieneremia Reike et al. 2013
G. Enicmus Thomson 1859
G. Latridius Herbst 1793
G. Revelieria Perris 1869
G. Stephostethus LeConte 1878
Fm. Teredidae Seidlitz 1888
+
Invalid names: Anommatinae Ganglbauer 1899 [empty], Xylariophilinae Pal and Lawrence 1986 [empty]
Fm. †Tetrameropsidae Kirejtshuk and Azar 2008
+
Invalid names: Alexiidae Imhoff 1856 [empty], Eupsilobiidae Casey 1895 [empty], Euxestidae Grouvelle 1908 [empty], Murmidiidae Jacquelin du Val 1858 [empty], Mycetaeidae Jacquelin du Val 1857 [empty]
Superfm. Cucujoidea Latreille 1802
+
Fm. Cryptophagidae Kirby 1837 [silken fungus beetle]
+
Subfm. Atomariinae LeConte 1861
+
Tr. Atomariini Le Conte 1861
Invalid names: Hypocoprini Reitter 1879 [empty]
Subfm. Cryptophaginae Kirby 1837
+
Invalid names: Lebanophytidae Kirejtshuk 2013 [synonym]
Fm. Cucujidae Latreille 1802
+
Fm. Cyclaxyridae Gimmel et al. 2009
+
G. Cyclaxyra Broun 1893
G. †Neolitochropus Lyubarsky and Perkovsky 2016
+
†Neolitochropus bedovoyi Lyubarsky and Perkovsky 2011
+
Invalid names: Ephistemus crassipes Lyubarsky and Perkovsky 2019 [synonym], Neolitochropus hoffeinsorum Lyubarsky and Perkovsky 2016 [synonym], Olibrolitus katyae Lyubarsky and Perkovsky 2020 [synonym]
†Neolitochropus michalskii Háva 2020
Invalid names: Olibrolitus Lyubarsky and Perkovsky 2020 [synonym]
Fm. Hobartiidae Sen Gupta and Crowson 1966
+
Invalid names: Hobartiinae Sen Gupta and Crowson 1969 [empty]
Fm. Laemophloeidae Ganglbauer 1899
+
G. Laemophloeus Dejean 1835
G. †Lithocoryne Scudder 1900
+
†Lithocoryne arcuata Wickham 1913
†Lithocoryne coloradensis Wickham 1914
†Lithocoryne gravis Scudder 1900
Invalid names: Laemophloeinae [invalid subgroup], Propalticidae Crowson 1952 [synonym]
Subfm. †Meligethiellinae Kirejtshuk and Ponomarenko 1990
+
G. †Meligethiella Medvedev 1969
+
†Meligethiella glabra Kirejtshuk and Ponomarenko 1990
†Meligethiella kovalevi Kirejtshuk and Ponomarenko 1990
†Meligethiella soroniiformis Medvedev 1969
G. †Ostomalynus Kirejtshuk and Ponomarenko 1990
+
†Ostomalynus ovalis Kirejtshuk and Ponomarenko 1990
Fm. Passandridae Blanchard 1845 [parasitic flat bark beetle]
+
Fm. Phalacridae Leach 1815
+
Subfm. Phalacrinae Leach 1815
+
G. Olibrus Erichson 1845
G. Phalacrus Paykull 1800
G. Stilbus Seidlitz 1872
Invalid names: Phaenocephalinae Matthews 1899 [empty]
Fm. Phloeostichidae Reitter 1911
+
G. †Pleuroceratos Poinar and Kirejtshuk 2008
+
†Pleuroceratos burmiticus Poinar and Kirejtshuk 2008
†Pleuroceratos jiewenae Tihelka et al. 2020
†Pleuroceratos tertius Háva 2023
Invalid names: Hymaeinae Sen Gupta and Crowson 1866 [empty], Phloeostichinae Reitter 1911 [empty]
Fm. Silvanidae Kirby 1837
+
Subfm. Silvaninae Kirby 1837
+
G. Airaphilus Redtenbacher 1858
G. Austronausibius Halstead 1980
G. Cathartosilvanus Grouvelle 1912
G. †Mistran Alekseev and Bukejs 2016
Invalid names: Hyliotinae [empty]
Invalid names: Agapythidae Sen Gupta and Crowson 1969 [empty], Brachypteridae Erichson 1845 [empty], Cavognathidae Sen Gupta and Crowson 1966 [empty], Clavicornia [invalid subgroup], Heteromera [invalid subgroup], Lamingtoniidae Sen Gupta and Crowson 1969 [empty], Myraboliidae Lawrence and Britton 1991 [empty], Priasilphidae Crowson 1973 [empty], Sinisilvanidae Hong 2002 [invalid subgroup], Tasmosalpingidae Lawrence and Britton 1991 [empty]
Superfm. Erotyloidea Latreille 1802
+
Fm. Boganiidae Sen Gupta and Crowson 1966
+
Invalid names: Paracucujinae Endrody-Younga and Crowson 1986 [empty]
Fm. Erotylidae Latreille 1802 [pleasing fungus beetle]
+
Subfm. Languriinae Crotch 1873
+
Tr. Thallisellini Sen Gupta 1968
Invalid names: Setariolinae Crowson 1952 [empty], Toraminae Sen Gupta 1967 [empty]
Subfm. Xenoscelinae Ganglbauer 1899
+
G. †Microzavaljus Lyubarsky and Perkovsky 2018
G. †Warnis Lyubarsky et al. 2016
G. †Xenohimatium Lyubarsky and Perkovsky 2012
G. †Xenophagus Lyubarsky and Perkovsky 2017
G. Zavaljus Reitter 1880
Invalid names: Dacninae Gistel 1856 [empty], Encaustinae Crotch 1876 [empty], Loberinae [empty], Megalodacninae Sen Gupta 1969 [empty]
Superfm. Lymexyloidea Fleming 1821
+
Fm. Lymexylidae Fleming 1821 [ship-timber beetle]
+
G. †Anancites Perkovsky 2022
+
†Anancites hukawngensis Chen and Zhang 2020
Invalid names: Adamas Chen and Zhang 2020 [replaced]
Subfm. Atractocerinae Laporte 1840
+
G. Atractocerus Palisot de Beauvois 1802
G. †Cratoatractocerus Wolf-Schwenninger 2011
G. †Cretoquadratus Chen 2019
G. †Paratractocerus Nazarenko and Perkovsky 2020
G. Raractocetus Kurosawa 1985
G. †Vetatractocerus Yamamoto 2019
Superfm. Nitiduloidea Latreille 1802
+
Fm. Helotidae Reitter 1876
+
G. Helota Macleay 1825
+
†Helota palmus Zhang 1989
†Helota senilis Zhang et al. 1994
Invalid names: Tarrodacne Zhang 1989 [synonym]
G. †Lobatihelota Li et al. 2023
+
†Lobatihelota iridescens Li et al. 2023
†Lobatihelota lescheni Li et al. 2023
Fm. Kateretidae Kirby 1837
+
G. Cercus Latreille 1796
G. Heterhelus Jacquelin du Val 1858
+
Subg. Heterhelus (Heterhelus) Jacquelin du Val 1858
†Heterhelus expressus Kirejtshuk and Nel 2008
G. †Pelretes Tihelka et al. 2021
+
†Pelretes bicolor Zhao et al. 2022
†Pelretes vivificus Tihelka et al. 2021
G. †Protokateretes Zhao et al. 2023
+
†Protokateretes antiquus Peris and Jelínek 2019
†Protokateretes magnascapulae Zhao et al. 2024
†Protokateretes megacephalus Zhao et al. 2023
†Protokateretes rectangulum Zhao et al. 2024
Fm. Monotomidae Laporte 1840 [root eating beetle]
+
Subfm. Monotominae Laporte 1840
+
Tr. †Cretakarenniini Peris and Delclòs 2015
Tr. Europini Sen Gupta 1988
Tr. Lenacini Crowson 1952
Tr. †Rhizophtomini Kirejtshuk and Azar 2009
Invalid names: Rhizophagidae Redtenbacher 1845 [synonym]
Fm. Nitidulidae Latreille 1802 [sap beetle]
+
Subfm. †Apophisandrinae Molino-Olmedo 2017
+
G. †Apophisandra Molino-Olmedo 2017
G. †Baltoraea Kurochkin and Kirejtshuk 2010
G. †Cretabaltoraea Peris et al. 2024
Subfm. Cillaeinae Kirejtshuk and Audisio 1986
+
Tr. Cillaeini Kirejtshuk and Audisio 1986
Tr. Conotelini Kirejtshuk and Kovalev 2022
Subfm. Cybocephalinae Jacquelin du Val 1858
+
G. Cybocephalus Erichson 1844
G. †Pastillocenicus Kirejtshuk and Nel 2008
Invalid names: Cybocephalini Jacquelin du Val 1858 [invalid subgroup]
Subfm. Nitidulinae Latreille 1802
+
Fm. Cychramini Gistel 1848
Tr. Cyllodini Everts 1898
Tr. Nitidulini Latreille 1802
Invalid names: Carpophilini Erichson 1842 [empty]
Subfm. Prometopiinae Böving and Craighead 1931
+
G. †Palaeometopia Kirejtshuk 2007
G. Prometopia Erichson 1843
G. †Sinonitidulina Hong 1983
+
†Sinonitidulina liugouensis Hong 1983
†Sinonitidulina luanpingensis Hong 1983
†Sinonitidulina punctata Hong 1983
Invalid names: Calonecrinae Kirejtshuk 1982 [empty], Cateretinae Kirby 1837 [empty]
Fm. Smicripidae Horn 1879 [palmetto beetle]
+
G. †Mesosmicrips Kirejtshuk 2017
+
†Mesosmicrips cretacea Cai and Huang 2016
†Mesosmicrips sunae Lyubarsky and Perkovsky 2023
G. Smicrips LeConte 1878
+
†Smicrips europeus Kirejtshuk and Nel 2008
†Smicrips fudalai Kupryjanowicz et al. 2019
†Smicrips gorskii Bukejs and Kirejtshuk 2015
Fm. Sphindidae Jacquelin du Val 1860 [cryptic slime mold beetle]
+
G. †Trematosphindus Li and Cai 2021
+
†Trematosphindus kirejtshuki Legalov et al. 2024
†Trematosphindus newtoni Li and Cai 2021
Invalid names: Aspidiphoridae Kiesenwetter 1877 [synonym], Odontosphindinae Sen Gupta and Crowson 1979 [empty], Sphindiphorinae Sen Gupta and Crowson 1979 [empty]
Fm. †Parandrexidae Kirejtshuk 1994
+
Subfm. †Cretoparacucujinae Kirejtshuk 2023
+
Subfm. †Parandrexinae Kirejtshuk 1994
+
G. †Parandrexis Martynov 1926
+
†Parandrexis beipiaoensis Hong 1983
†Parandrexis parvula Martynov 1926
†Parandrexis rotundicollis Kirejtshuk 1994
†Parandrexis subtilis Kirejtshuk 1994
Unr. Phytophaga
+
Superfm. Chrysomeloidea Latreille 1802
+
Fm. Cerambycidae Latreille 1802 [longhorned beetle]
+
G. †Alifasciola Zhang 1989
Subfm. Cerambycinae Latreille 1802
G. †Cerambycinus Germar 1839
G. †Cerambycites Deichmüller 1886
G. †Delostromatium Zhang 1989
Subfm. Dorcasominae Lacordaire 1868
G. †Ferrucornus Hart et al. 2024
G. †Glischra Zhang et al. 1994
G. †Hornia Zhang et al. 1994
G. Hylotrupes Audinet-Serville 1834
Subfm. Lamiinae Latreille 1825
Subfm. Lepturinae Latreille 1802
G. †Mesosites Deichmüller 1881
G. †Miopyrestes Zhang et al. 1994
Subfm. Necydalinae Latreille 1825
Subfm. Parandrinae Blanchard 1845
G. †Paraphilus Zhang et al. 1994
G. †Perimesosa Zhang et al. 1994
G. †Podozoodes Zhang et al. 1994
Subfm. Prioninae Latreille 1802
G. †Saxosa Zhang 1989
G. †Scaptolenopsis Wickham 1914
G. †Sinopraecipuus Yu et al. 2015
Subfm. Spondylidinae Audinet-Serville 1832
G. †Trosites Zhang 1989
G. †Ventiala Hart et al. 2024
Invalid names: Pseudonepidae Jordan 1967 [synonym], Spondylididae Audinet-Serville 1832 [synonym]
Fm. Chrysomelidae Latreille 1802 [leaf beetle]
+
Subfm. Alticinae Newman 1834 [flea beetle]
Subfm. Bruchinae Latreille 1802
Subfm. Cassidinae Gyllenhal 1813 [tortoise beetle]
+
Invalid names: Hispinae Gyllenhal 1813 [synonym]
Subfm. Chrysomelinae Latreille 1802
G. †Chrysomelites Heer 1864
G. †Chrysomelopsis Handlirsch 1906
G. †Chrysothoracus Zhang 1989
G. †Coelocassida Birket-Smith 1977
G. Colaphellus Weise 1916
Subfm. Criocerinae Latreille 1804
Subfm. Cryptocephalinae Gyllenhal 1813
Subfm. Donaciinae Kirby 1837
Subfm. Eumolpinae Hope 1840
G. †Eumolpites Heer 1865
Subfm. Galerucinae Latreille 1802
G. Gastrolina Baly 1859
G. †Hadroscelus Quedenfeldt 1885
G. Hydrothassa Thomson 1859
Subfm. Lamprosomatinae Lacordaire 1848
G. Longitarsus Berthold 1827
Subfm. Sagrinae Leach 1815 [frog-legged beetle]
Invalid names: Clythrinae Kirby 1837 [empty], Halticinae Newman 1834 [empty]
Fm. Megalopodidae Latreille 1802
+
Subfm. Palophaginae Kuschel and May 1990
Invalid names: Megalopodinae Latreille 1802 [empty], Zeugophorinae Böving and Craighead 1931 [empty]
Fm. Orsodacnidae Thomson 1859
+
Subfm. Aulacoscelidinae Chapuis 1874
Invalid names: Orsodacninae Thomson 1859 [empty]
Fm. Vesperidae Mulsant 1839
+
Subfm. Philinae Thomson 1861
Invalid names: Anoplodermatinae Guerin-Meneville 1840 [empty], Vesperinae Mulsant 1839 [empty]
Invalid names: Cerambycoidea Latreille 1802 [synonym], Oxypeltidae Lacordaire 1868 [empty]
Superfm. Curculionoidea Latreille 1802 [weevil]
+
Fm. Anthribidae Billberg 1820 [fungus weevil]
+
Subfm. Anthribinae Billberg 1820
Subfm. Choraginae Kirby 1819
G. Hormiscus Waterhouse 1845
Subfm. †Juranthribinae Legalov 2011
Subfm. †Protoscelinae Medvedev 1968
Subfm. Urodontinae Thomson 1859
Fm. Attelabidae Billberg 1820 [leaf rolling weevil]
+
Subfm. Apoderinae Jekel 1860
Subfm. Attelabinae Billberg 1820
Subfm. Rhynchitinae Gistel 1848 [tooth-nosed snout weevil]
+
Invalid names: Isotheinae Scudder 1893 [synonym]
Invalid names: Rhynchitidae Gistel 1848 [synonym]
Fm. Belidae Schoenherr 1826
+
Subfm. †Montsecbelinae Legalov 2015
Subfm. Oxycoryninae Schönherr 1840
Invalid names: Aglycyderinae Wollaston 1864 [empty]
Fm. Brentidae Billberg 1820
+
Subfm. Apioninae Schoenherr 1823
Subfm. Brentinae Billberg 1820
Subfm. Cyladinae Schoenherr 1823
Subfm. Eurhynchinae Lacordaire 1863
Subfm. Ithycerinae Schoenherr 1823
Subfm. Nanophyinae Gistel 1848
Invalid names: Ithyceridae Schoenherr 1823 [synonym]
Fm. Caridae Thompson 1992
+
Subfm. †Baissorhynchinae Zherikhin 1993
Subfm. Carinae Thompson 1992
Subfm. Chilecarinae Legalov 2009
Fm. Cimberididae de Gozis 1882
+
Tr. Cimberidini de Gozis 1882
Tr. Doydirhynchini Pierce 1916
Tr. †Kuschelomacerini Riedel 2010
Tr. †Rhamphophorini Poinar and Brown 2021
Fm. Curculionidae Latreille 1802 [snout beetle]
+
Subfm. Bagoinae Thomson 1859
Subfm. Brachycerinae Billberg 1820
G. †Calandrites Scudder 1893
Subfm. Ceutorhynchinae Gistel 1848
Subfm. Conoderinae Schoenherr 1833
Subfm. Cossoninae Schoenherr 1825
Subfm. Cryptorhynchinae Schoenherr 1825
Subfm. Curculioninae Latreille 1802
G. †Curculionites Heer 1847
Subfm. Cyclominae Schoenherr 1826
Subfm. Dryophthorinae Schoenherr 1825
+
Invalid names: Orthognathinae Lacordaire 1866 [synonym], Rhynchophorinae Schonherr 1833 [synonym]
Subfm. Ectemnorrhininae
Subfm. Erirhininae Schoenherr 1825
Subfm. Hylobiinae Kirby 1837
Subfm. Hyperinae Marseul 1863
Subfm. Lixinae Schonherr 1823
G. †Meristos Murray 1860
Subfm. Mesoptiliinae Lacordaire 1863
Subfm. Molytinae Schonherr 1823
G. †Mononychites Haupt 1956
G. †Ophryastites Scudder 1893
Subfm. Orobitidinae Thomson 1859
Tr. †Palaeocryptorhynchini Poinar and Legalov 2014
G. †Palaeotanymecides Mani 1947
Subfm. Platypodinae Shuckard 1839 [ambrosia beetle]
+
Invalid names: Tesserocerinae Strohmeyer 1914 [synonym]
Subfm. Scolytinae Latreille 1804 [bark beetle]
G. Sitones Germar 1817
G. †Syntomostylus Scudder 1893
Invalid names: Bagous bicolor Förster 1891 [nomen dubium], Balaninus barthelemyi Hope 1847 [nomen dubium], Baridinae Schoenherr 1836 [empty], Brachyceridae Billberg 1820 [synonym], Brachycerus germanus Heer 1847 [nomen dubium], Brachycerus lecoqui Oustalet 1870 [nomen dubium], Brachycerus nanus Heer 1864 [nomen dubium], Cleonus foersteri Theobald 1937 [nomen dubium], Cleonus inflexus Oustalet 1874 [nomen dubium], Cleonus poli Hope 1847 [nomen dubium], Coniatus minusculus Oustalet 1874 [nomen dubium], Coniatus scudderi Théobald 1935 [nomen dubium], Cossonus marioni Oustalet 1874 [nomen dubium], Cryptorhynchus gypsi Oustalet 1874 [nomen dubium], Cryptorhynchus incertus Förster 1891 [nomen dubium], Cryptorrhynchinae Schoenherr 1825 [empty], Curculionites exiguus Oustalet 1874 [nomen dubium], Curculionites lividus Heer 1856 [nomen dubium], Curculionites parvulus Heer 1856 [nomen dubium], Dryophthoridae Schoenherr 1825 [synonym], Erirhinus chantrei Oustalet 1874 [nomen dubium], Erirrhininae Schoenherr 1825 [empty], Gymnetron rotundicolle Förster 1891 [nomen dubium], Hydronomus nasutus Oustalet 1874 [nomen dubium], Lachnopus dilatatus Theobald 1937 [nomen dubium], Lachnopus humatus Scudder 1893 [nomen dubium], Larinus pici Piton 1935 [nomen dubium], Lixus oeningensis Heer 1864 [nomen dubium], Otiorhynchus flaccus Scudder 1893 [nomen dubium], Otiorhynchus tumbae Scudder 1890 [nomen dubium], Pachylobius compressus Scudder 1893 [nomen dubium], Pachylobius depraedatus Scudder 1893 [nomen dubium], Pachylobius martyi Piton and Théobald 1936 [nomen dubium], Pachylobius yungi Piton 1936 [nomen dubium], Phloeotribus zimmermanni Wickham 1916 [nomen dubium], Phyllobius antecessor Scudder 1893 [nomen dubium], Phyllobius avus Scudder 1893 [nomen dubium], Phyllobius carcerarius Scudder 1893 [nomen dubium], Phyllobius incertus Théobald 1935 [nomen dubium], Phytonomus incertus Théobald 1935 [nomen dubium], Rhinocyllus marmoreus Théobald 1935 [nomen dubium], Rhynchaenus solieri Hope 1847 [nomen dubium], Rhyssomatus tabescens Scudder 1893 [nomen dubium], Rhytirhinini [empty], Scythropus somniculosus Scudder 1893 [nomen dubium], Sphenophorus elegans Théobald 1935 [nomen dubium], Tanymecus gautieri Piton 1936 [nomen dubium], Tanysphyrus deletus Oustalet 1874 [nomen dubium]
Fm. †Mesophyletidae Poinar 2008
+
Subfm. †Aepyceratinae Poinar et al. 2017
Subfm. †Mesophyletinae Poinar 2008
Fm. Nemonychidae Bedel 1882 [pine flower snout beetle]
+
Subfm. †Brenthorrhininae Arnoldi 1977
Subfm. †Cretonemonychinae Gratshev and Legalov 2009
Subfm. †Eobelinae Arnoldi 1977
Subfm. Rhinorhynchinae Voss 1922
Subfm. †Slonikinae Zherikhin 1977
Invalid names: Nemonychinae Bedel 1882 [empty]
Fm. †Obrieniidae Zherikhin and Gratshev 1993
+
Subfm. †Kararhynchinae Zherikhin and Gratshev 1993
Subfm. †Obrieniinae Zherikhin and Gratshev 1993
Subfm. †Pseudobrieniinae Legalov 2012
Invalid names: Akulosamphus Aymard 1856 [nomen nudum], Brenthidae Billberg 1820 [empty], Erirhinidae Schoenherr 1825 [empty], Loricera glacialis Scudder 1877 [nomen dubium]
Superfm. Tenebrionoidea Latreille 1802
+
Fm. Aderidae Csiki 1909
+
G. †Palaeocnopus Alekseev and Grzymala 2015
+
†Palaeocnopus densipunctatus Alekseev and Grzymala 2015
†Palaeocnopus glabricornis Alekseev and Grzymala 2015
†Palaeocnopus mara Alekseev and Grzymala 2015
†Palaeocnopus saeticornis Alekseev and Grzymala 2015
G. Vanonus Casey 1895
+
†Vanonus aestiorum Alekseev and Grzymala 2015
†Vanonus ulmerigicus Alekseev and Grzymala 2015
Invalid names: Circaeidae Iablokoff-Khnzorian 1961 [synonym], Xylophilidae [replaced]
Fm. Anthicidae Latreille 1819 [ant like flower beetle]
+
Subfm. Macratriinae LeConte 1862
+
Tr. †Camelomorphini Kirejtshuk et al. 2008
Tr. Macratriini Le Conte 1862
Invalid names: Copobaeninae Abdullah 1969 [empty]
Fm. Boridae Thomson 1859
+
Invalid names: Borinae Thomson 1859 [empty], Synercticinae Lawrence and Pollock 1994 [empty]
Fm. Ciidae Leach 1819 [minute tree-fungus beetle]
+
Invalid names: Sphindociinae Lawrence 1974 [empty]
Fm. †Elodophthalmidae Kirejtshuk and Azar 2008
+
G. †Elodophthalmus Kirejtshuk and Azar 2008
+
†Elodophthalmus gracilis Kirejtshuk and Azar 2008
†Elodophthalmus harmonicus Kirejtshuk and Azar 2008
†Elodophthalmus maksoudae Cai et al. 2024
Subfm. Ischaliidae Blair 1920 [false fire-colored beetle]
+
G. Ischalia Pascoe 1860
+
Subg. Ischalia (Eupleurida) LeConte 1862
Subg. Ischalia (Ischalia) Pascoe 1860
Subg. †Ischalia (Telnovia) Alekseev and Bukejs 2017
G. †Prototrichalus Molina-Olmedo et al. 2020
+
†Prototrichalus jingpo Telnov and Kundrata 2023
†Prototrichalus meiyingae Molina-Olmedo et al. 2020
†Prototrichalus milleri Molina-Olmedo et al. 2020
†Prototrichalus sepronai Molina-Olmedo et al. 2020
G. †Kulindrobor Tihelka et al. 2021
+
†Kulindrobor enigmaticus Tihelka et al. 2021
†Kulindrobor magnus Tihelka et al. 2021
Fm. Melandryidae Leach 1815 [false darkling beetle]
+
Subfm. Melandryinae Leach 1815
+
Tr. Dircaeini Mulsant 1856
G. †Electroabdera Alekseev 2014
Tr. Hypulini Gistel 1848
Tr. Orchesiini Mulsant 1856
G. †Quasianisoxya Alekseev 2015
Tr. Serropalpini Latreille 1829
G. Symphora LeConte 1866
Tr. Xylitini Thomson 1864
Invalid names: Osphyinae Mulsant 1856 [empty]
Fm. Meloidae Gyllenhal 1810 [blister beetle]
+
Subfm. Meloinae Gyllenhal 1810
+
Tr. Epicautini Parker and Boving 1924
Tr. Lyttini Solier 1851
Tr. Meloini Gyllenhal 1810
Tr. Mylabrini Rafinesque 1815
Tr. Tetraonycini Böving and Craighead 1931
Invalid names: Eleticinae Wellman 1910 [empty]
Fm. Mordellidae Latreille 1802 [tumbling flower beetle]
+
Subfm. †Apotomourinae Bao et al. 2018
+
G. †Apotomoura Bao et al. 2018
G. †Multispinus Bao et al. 2018
G. Falsomordellistena Ermisch 1941
G. Glipostena Ermisch 1941
G. Mordellaria Ermisch 1950
Subfm. Mordellinae Latreille 1802
+
G. †Angimordella Bao et al. 2019
Tr. Conaliini Ermisch 1956
Tr. †Mediumiugini Peris and Ruzzier 2013
Tr. Mordellini Latreille 1802
Tr. Mordellistenini Ermisch 1941
Tr. Raynoldsiellini Franciscolo 1957
Tr. Reynoldsiellini Franciscolo 1957
Tr. Stenaliini Franciscolo 1955
G. †Petrimordella Batelka et al. 2023
+
†Petrimordella indata Statz 1952
†Petrimordella inundata Wickham 1914
†Petrimordella lapidicola Wickham 1909
†Petrimordella nearctica Wickham 1914
†Petrimordella nigrapilosa Statz 1952
†Petrimordella oligocenica Nel 1985
†Petrimordella priscula Cockerell 1925
†Petrimordella protogaea Wickham 1914
†Petrimordella rasnitsyni Odnosum and Perkovsky 2016
†Petrimordella scudderiana Wickham 1914
†Petrimordella smithiana Wickham 1913
†Petrimordella stygia Wickham 1914
Subfm. †Praemordellinae Shchegoleva-Barovskaya 1929
+
G. †Bellimordella Liu et al. 2008
G. †Cretanaspis Huang and Yang 1999
G. †Mirimordella Liu et al. 2007
G. †Praemordella Shchegoleva-Barovskaya 1929
G. †Wuhua Wang and Zhang 2011
Invalid names: Ctenidiinae Franciscolo 1951 [empty], Praemordellidae Shchegoleva-Barovskaya 1929 [synonym]
Fm. Mycetophagidae Leach 1815 [hairy fungus beetle]
+
Invalid names: Bergininae Leng 1920 [empty], Esarcinae Reitter 1882 [empty]
Fm. Mycteridae Oken 1843 [palm and flower beetle]
+
Subfm. Eurypinae Thomson 1860
+
G. †Bertinotus Kirejtshuk and Nel 2009
G. †Europoeurypus Alekseev et al. 2020
G. †Glesoconomorphus Alekseev et al. 2019
G. Omineus Lewis 1895
Invalid names: Lacconotinae Le Conte 1862 [synonym]
Invalid names: Hemipeplinae Lacordaire 1854 [empty]
Fm. Oedemeridae Latreille 1810 [false blister beetle]
+
Subfm. Oedemerinae Latreille 1810
+
Tr. Asclerini Gistel 1848
G. †Darwinylus Peris 2017
Tr. Ditylini Mulsant 1858
Tr. Oedemerini
Invalid names: Saperda absyrti Heer 1847 [nomen dubium]
Fm. Prostomidae Thomson 1859 [jugular-horned beetle]
+
G. Prostomis Latreille 1825
G. †Vetuprostomis Engel and Grimaldi 2008
+
†Vetuprostomis angularis Li and Cai 2022
†Vetuprostomis consimilis Engel and Grimaldi 2008
†Vetuprostomis gaoi Li and Cai 2022
Fm. Pyrochroidae Latreille 1807
+
Invalid names: Cretaceimelittomoides Vulcano and Pereira 1987 [nomen nudum], Cretaceimelittomoides cearensis Vulcano and Pereira 1987 [nomen nudum], Pilipalpinae Abdullah 1964 [empty], Tydessinae Nikitsky 1986 [empty]
Fm. Pythidae Solier 1834
+
Fm. Rhipiphoridae Gemminger 1870 [wedge shaped beetle]
+
G. Rhipiphorus Bosc 1792
Invalid names: Rhipidiinae Gerstaecker 1855 [empty], Rhipiphorinae Gemminger 1870 [empty]
Fm. Ripiphoridae Gemminger 1870 [wedge shaped beetle]
+
Subfm. Pelecotominae Seidlitz 1875
+
G. †Burmitoma Batelka et al. 2018
G. Clinops Gerstaecker 1855
G. †Flabellotoma Batelka et al. 2016
G. †Plesiotoma Batelka et al. 2018
G. †Spinotoma Hsiao and Huang 2017
Invalid names: Micholaeminae Viana 1971 [synonym]
Subfm. Ripidiinae Gerstaecker 1855
+
Tr. Eorhipidiini Iablokoff-Khnzorian 1986
G. †Pauroripidius Kaupp and Nagel 2001
Tr. Ripidiini Gerstaecker 1855
G. †Ripilarva Batelka and Beutel 2024
Invalid names: Ripidius megalophus Dalman 1826 [nomen dubium]
Subfm. Ripiphorinae Gemminger 1870
+
Tr. Macrosiagonini von Heyden 1908
Tr. Ripiphorini Gemminger 1870
Invalid names: Hemirhipidiinae Heller 1921 [empty], Ptilophorinae Gerstaecker 1855 [empty]
Fm. Salpingidae Leach 1815 [narrow-waisted bark beetle]
+
Subfm. Inopeplinae Grouvelle 1908
+
G. Diagrypnodes Waterhouse 1876
G. Eopeplus Kirejtshuk and Nel 2009
Subfm. Salpinginae Leach 1815
+
Tr. Istrisiini Nikitsky 1992
G. †Protolissodema Alekseev 2013
G. Salpingus Illiger 1801
G. Sphaeriestes Stephens 1829
Invalid names: Aegialitinae LeConte 1862 [empty], Dacoderinae LeConte 1862 [empty], Othniinae LeConte 1861 [empty]
Fm. Scraptiidae Gistel 1848 [false flower beetle]
+
Subfm. Scraptiinae Gistel 1848
+
G. †Archescraptia Abdullah 1964
G. †Palaeoscraptia Abdullah 1964
Tr. Scraptiini Gistel 1848
G. †Scraptiomima Medvedev 1969
+
†Scraptiomima brachycornis Medvedev 1969
†Scraptiomima minima Ponomarenko 1990
Fm. Stenotrachelidae Thomson 1859
+
Invalid names: Cephaloinae LeConte 1862 [empty], Nematoplinae LeConte 1862 [empty], Stenotrachelinae Thomson 1859 [empty], Stoliinae Nikitsky 1985 [empty]
Fm. Tenebrionidae Latreille 1802 [darkling beetle]
+
Subfm. Alleculinae Laporte 1840 [comb-clawed beetle]
+
Tr. Alleculini Laporte 1840
Tr. Cteniopodini Solier 1835
+
Invalid names: Cisteleniae Latreille 1802 [replaced], Omophlini Mulsant 1856 [synonym]
Invalid names: Dascillus exiguus Zhang 1989 [nomen dubium]
Subfm. Blaptinae Leach 1815
+
Tr. Amphidorini LeConte 1862
Tr. Blaptini Leach 1815
Tr. Opatrini Brullé 1832
Tr. Pedinini Eschscholtz 1829
Subfm. Diaperinae Latreille 1802
+
Tr. Diaperini Latreille 1802
Tr. Gnathidiini Gieben 1921
Tr. Hypophlaeini Billberg 1820
Tr. Scaphidemini Reitter 1922
Subfm. Lagriinae Latreille 1825
+
Tr. Belopini Reitter 1917
Tr. †Gonialaenini Nabozhenko et al. 2019
Tr. Laenini Seidlitz 1895
Tr. Lagriini Latreille 1825
Tr. Lupropini Ardoin 1958
Tr. Prateini Aalbu et al. 2023
Subfm. Nilioninae Oken 1843 [false ladybird beetle]
+
G. Nilio Latreille 1829
Invalid names: Nilionini Thomson 1860 [invalid subgroup]
Subfm. Pimeliinae Latreille 1802
+
Tr. Asidini Fleming 1821
Tr. Coniontini Waterhouse 1858
Tr. Nycteliini Solier 1834
Tr. Nyctoporini Lacordaire 1859
Tr. Pimeliini Latreille 1802
Tr. Sepidiini Eschscholtz 1829
Tr. Stenosini Schaum 1859
Tr. Tentyriini Eschscholtz 1831
Tr. Zophosini Solier 1834
Invalid names: Akidini Billberg 1820 [empty]
G. †Rhinohelaeites Haupt 1950
+
†Rhinohelaeites longipes Haupt 1950
†Rhinohelaeites punctatulus Haupt 1950
Subfm. Stenochiinae Kirby 1837
+
G. †Anthracohelops Haupt 1950
G. †Caryosoma Haupt 1950
G. †Eodromus Pongrácz 1935
G. †Mimohelops Haupt 1950
G. †Parakeleusticus Haupt 1950
G. †Pseudohelops Haupt 1950
G. †Pyrochalcaspis Haupt 1950
Subfm. Tenebrioninae Latreille 1802
+
Tr. Alphitobiini Reitter 1917
Tr. Amarygmini Gistel 1848
Tr. Bolitophagini Kirby 1837
Tr. Eulabini Horn 1870
Tr. Heleini Fleming 1821
Tr. Helopini Latreille 1802
Tr. Melanimonini Seidlitz 1894
Tr. Metaclisini Steiner 2016
Tr. Palorini Matthews 2003
Tr. Scotobiini Solier 1838
Tr. Tenebrionini Latreille 1802
Tr. Toxicini Oken 1843
Tr. Triboliini Gistel 1848
Tr. Ulomini Blanchard 1845
Invalid names: Titaenini Fauvel 1905 [empty], Ulus minutus Wickham 1914 [nomen dubium]
G. †Tenebrionites Cockerell 1920
+
†Tenebrionites alatus Cockerell 1927
†Tenebrionites anglicus Cockerell 1920
†Tenebrionites inclinans Cockerell 1925
Invalid names: Ancylochira oligostriata Piton and Théobald 1937 [nomen dubium], Tenebrio calculensis Scudder 1895 [nomen dubium]
Fm. Tetratomidae Billberg 1820 [polypore fungus beetle]
+
Subfm. Eustrophinae Gistel 1848
+
G. †Cretosynstrophus Cai et al. 2016
Tr. Eustrophini Gistel 1848
Tr. Holostrophini Nikitsky 1998
Subfm. Hallomeninae Gistel 1848
+
G. †Elasontagius Alekseev and Bukejs 2020
G. †Pseudohallomenus Nikitsky 1977
Subfm. Tetratominae Billberg 1820
+
Tr. †Archaeolupropini Nabozhenko et al. 2023
G. Tetratoma Fabricius 1790
Invalid names: Penthinae Lacordaire 1859 [empty], Piseninae Miyatake 1960 [empty]
Fm. Zopheridae Solier 1834 [ironclad beetle]
+
Subfm. Colydiinae Billberg 1820
+
G. †Bifurderum Cheng et al. 2020
Tr. Colydiini Billberg 1820
G. †Coslonatus Li and Cai 2021
G. †Cryphalites Cockerell 1917
Tr. Gempylodini Sharp 1893
G. †Stegastochlidus Poinar and Vega 2020
Tr. Synchitini Erichson 1845
Subfm. Zopherinae Solier 1834
+
Tr. Monommatini Blanchard 1845
Tr. Pycnomerini Erichson 1845
Tr. Usechini Horn 1867
Tr. †Zopheromimini Alekseev and Nabozhenko 2023
Invalid names: Colydiidae Billberg 1820 [replaced]
Invalid names: Afreminae Levey 1985 [empty], Archeocrypticidae Kaszab 1964 [empty], Asiamordella Hong 2002 [nomen nudum], Chalcodryidae Watt 1974 [empty], Cossyphodinae Wasmann 1899 [empty], Eurygenius wickhami Cockerell 1917 [nomen dubium], Fushunicucujus Hong 2002 [nomen nudum], Lagrioididae Abdullah and Abdullah 1968 [empty], Perimylopidae St. George 1939 [empty], Pterogeniidae Crowson 1953 [empty], Trachelostenidae Lacordaire 1859 [empty], Trictenotomidae Blanchard 1845 [empty]
Infraor. Elateriformia Crowson 1960
+
Superfm. Buprestoidea Leach 1815
+
Fm. Buprestidae Leach 1815 [jewel beetle]
+
G. †Agrilium Westwood 1854
+
†Agrilium cyllabacus Westwood 1854
†Agrilium cyllarus Westwood 1854
†Agrilium stomphax Westwood 1854
†Agrilium strombus Westwood 1854
†Agrillium angustum Ponomarenko 1985
†Agrillium antiquum Ponomarenko 1985
†Agrillium minor Ponomarenko 1985
G. †Archelater Carpenter 1992
+
†Archelater aeneoviridis Pongrácz 1935
†Archelater major Pongrácz 1935
†Archelater megaloprepus Pongrácz 1935
†Archelater rutilogrammus Pongrácz 1935
Subfm. Buprestinae Leach 1815
+
Tr. Anthaxiini Gory and Laporte 1839
G. †Brachyspathus Wickham 1917
G. †Brevista Alexeev 1995
Tr. Buprestini Leach 1815
G. †Buprestites Heer 1847
Tr. Chrysobothrini Gory and Laporte 1837
G. †Cretothyrea Alexeev 1996
G. †Fuesslinia Heer 1847
G. †Illolampra Zhang et al. 1994
G. †Jurabuprestis Alexeev 2000
G. †Kzylordynia Alexeev 1995
G. †Lomatus Murray 1860
Tr. Melanophilini Bedel 1921
G. †Metabuprestium Alexeev 1995
G. †Micranthaxia Heer 1852
G. †Protogenia Heer 1847
G. †Pseudothyrea Handlirsch 1906
Tr. Stigmoderini Lacordaire 1857
Tr. Thomassetiini Bellamy 1987
G. †Trapezitergum Yu et al. 2013
Invalid names: Glaphyropteridae Pongrácz 1935 [invalid subgroup]
Subfm. Chrysochroinae Laporte 1835
+
Tr. Chrysochroini Laporte 1835
Tr. Dicercini Gistel 1848
Tr. Paraleptodemini Cobos 1975
Tr. Poecilonotini Jakobson 1913
Tr. Sphenopterini Lacordaire 1857
G. Coroebus Gory and Laporte 1839
G. †Ctenicerium Westwood 1854
+
†Ctenicerium blissus Westwood 1854
+
Invalid names: Buprestium dardanus Westwood 1854 [synonym], Elater morrisi Giebel 1856 [objective synonym]
†Ctenicerium gigas Handlirsch 1906
†Ctenicerium stygnus Westwood 1854
†Ctenicerium valgus Westwood 1854
Subfm. †Parathyreinae Alekseev 1994
+
G. †Acmaeoderimorpha Alekseev 1994
G. †Ancestrimorpha Alekseev 1994
G. †Cretalbiana Alexeev 2013
G. †Cretofrontolina Alexeev 2009
G. †Karatausia Alekseev 1994
G. †Mongoligena Alekseev 1994
G. †Mongolobuprestis Alekseev 1994
G. †Paleas Alekseev 1994
G. †Paramongoligena Alekseev 1994
G. †Parathyrea Alekseev 1994
G. †Pseudochrysobothris Alekseev 1994
G. †Pseudomongoligena Alexeev 2000
G. †Sinoparathyrea Pan et al. 2011
G. †Stigmoderimorpha Alekseev 1994
G. †Umerata Alekseev 1994
G. †Planocoleus Hong 1982
+
†Planocoleus ensatus Hong 1982
†Planocoleus glabratus Hong 1982
†Planocoleus mjangadiensis Ponomarenko 1986
Subfm. Polycestinae Lacordaire 1857
+
Tr. Acmaeoderini Kerremans 1893
Tr. Haplostethini Le Conte 1861
G. †Mongoligenula Yu et al. 2015
Tr. Polyctesini Cobos 1955
Tr. Ptosimini Kerremans 1902
Invalid names: Ancylochira pinguis Piton 1940 [nomen dubium], Ancylochira striatopunctata Piton 1940 [nomen dubium], Ancylochira violaceocyanescens Piton 1940 [nomen dubium], Chrysobothrites Heer 1864 [nomen nudum], Chrysochroa punctata Piton 1940 [nomen dubium], Iridotaenia hovassei Piton 1940 [nomen dubium], Stigmodera unicolor Piton 1940 [nomen dubium]
Fm. Schizopodidae LeConte 1859
+
Subfm. Schizopodinae LeConte 1861
+
Tr. †Cretadystaxiini Legalov 2025
Tr. †Electrapatini Yablokov-Khnzorian 1962
Invalid names: Mesohelophorus mongolicus Ponomarenko 1986 [nomen dubium]
Superfm. Byrrhoidea Latreille 1804
+
Fm. Byrrhidae Latreille 1804 [pill beetle]
+
G. †Amphicyrtella Ponomarenko 1990
+
†Amphicyrtella clavator Ponomarenko 1990
†Amphicyrtella granulum Ponomarenko 1990
Subfm. Byrrhinae Latreille 1804
+
Tr. Byrrhini Latreille 1804
Tr. Morychini El Moursy 1961
Tr. Pedilophorini Casey 1912
Tr. Simplocariini Mulsant and Rey 1869
G. †Mesobyrrhus Huang and Zhang 1997
+
†Mesobyrrhus parvus Huang and Zhang 1997
†Mesobyrrhus tanae Huang and Zhang 1997
G. †Mesosimplocaria Ponomarenko 1990
+
†Mesosimplocaria ovalis Ponomarenko 1990
†Mesosimplocaria parmula Ponomarenko 1990
G. †Nosotetocus Scudder 1892
+
†Nosotetocus debilis Scudder 1900
†Nosotetocus marcovi Scudder 1892
†Nosotetocus vespertinus Scudder 1900
Subfm. Syncalyptinae Mulsant and Rey 1869
+
G. Curimopsis Ganglbauer 1902
Tr. Syncalyptini Mulsant and Rey 1869
Invalid names: Nosodendron tritavum Scudder 1890 [nomen dubium]
Invalid names: Chelonarium montanum Wickham 1914 [nomen dubium]
Superfm. Dascilloidea Gue'rin-Me'neville 1843
+
Fm. Dascillidae Guerin-Meneville 1843 [soft bodied plant beetle]
+
Subfm. Dascillinae Guerin-Meneville 1843
+
G. †Cretodascillus Jin et al. 2013
G. Dascillus Latreille 1796
G. †Parelateriformius Yan and Wang 2010
G. †Lyprodascillus Zhang 1989
+
†Lyprodascillus peritelus Zhang et al. 1994
†Lyprodascillus sypharus Zhang 1989
Invalid names: Karumiidae Escalera 1913 [empty]
Superfm. Dryopoidea
+
Fm. Chelonariidae Blanchard 1845
+
G. Chelonarium Fabricius 1801
+
†Chelonarium andabata Alekseev and Bukejs 2021
†Chelonarium dingansich Alekseev and Bukejs 2021
†Chelonarium dominicanum Poinar and Háva 2021
Fm. Dryopidae Billberg 1820 [long toed water beetle]
+
G. Dryops Olivier 1791
+
Dryops ernesti des Gozis 1886
Invalid names: Parnus Fabricius 1792 [synonym]
G. Helmis Latreille 1802
Invalid names: Parnidae Leach 1817 [replaced]
Fm. Elmidae Curtis 1830 [riffle beetle]
+
G. Cleptelmis Sanderson 1953
+
Invalid names: Larainae LeConte 1861 [empty]
Fm. Eulichadidae Crowson 1973 [forest stream beetle]
+
Subfm. Eulichadinae Crowson 1973
+
G. †Cretasyne Yan et al. 2013
G. †Mesaplus Hong 1983
G. †Mesodascilla Martynov 1926
Invalid names: Lasiosyne punctata Yan 2012 [nomen dubium], Lasiosyne shartegiensis Yan 2012 [nomen dubium]
Fm. Heteroceridae MacLeay 1825 [Variegated mud-loving beetle]
+
G. †Excavotarsus Li et al. 2020
+
†Excavotarsus lini Li et al. 2020
†Excavotarsus minor Li et al. 2020
G. †Heterocerites Ponomarenko 1986
+
†Heterocerites kobdoensis Ponomarenko 1986
†Heterocerites magnus Prokin and Ren 2011
Invalid names: Elythomerinae Pacheco 1964 [empty]
Fm. Limnichidae Erichson 1846 [minute marsh loving beetle]
+
G. Platypelochares Champion 1923
Invalid names: Hyphalinae Britton 1971 [empty], Thaumastodinae Champion 1924 [empty]
Fm. Lutrochidae Kasap and Crowson 1975 [robust marsh loving beetle]
+
Fm. †Mastigocoleidae Tihelka et al. 2022
+
G. †Mastigocoleus Tihelka and Cai 2022
+
†Mastigocoleus resinicola Tihelka and Cai 2022
†Mastigocoleus rhinoceros Tihelka and Cai 2022
Fm. Psephenidae Lacordaire 1854 [water penny beetle]
+
Subfm. Eubrianacinae Jacobson 1913
+
G. Eubrianax Kiesenwetter 1874
G. †Succinumanax Bao et al. 2018
G. †Vetujinbrianax Cai and Huang 2018
Subfm. Eubriinae Lacordaire 1857
+
Subfm. Psephenoidinae Hinton 1939
+
G. Malacopsephenoides Jeng and Sato 2006
G. Psephenoides Gahan 1914
Fm. Ptilodactylidae Laporte 1836
+
Invalid names: Aploglossinae Champion 1897 [empty], Araeopidiinae Lawrence 1991 [empty]
Invalid names: Actinocerites Hong 2002 [nomen nudum], Brachypodites Hong 2002 [nomen nudum], Callirhipidae Emden 1924 [empty], Cneoglossidae Champion 1897 [empty], Expansifemoria Hong 2002 [nomen nudum]
Superfm. Elateroidea Leach 1815
+
Fm. Artematopodidae Lacordaire 1857 [soft bodied plant beetle]
+
Invalid names: Artematopodinae Lacordaire 1857 [empty], Eurypogonidae [replaced], Notocupes undatabdominus Lin 1980 [nomen dubium]
Fm. †Berendtimiridae Winkler 1987
+
Subfm. †Berendtimirinae Winkler 1987
+
G. †Berendtimirus Winkler 1987
G. †Jantarokrama Kirejtshuk and Kovalev 2015
Subfm. †Retromalisinae Fanti et al. 2023
+
G. †Maigus Fanti et al. 2023
G. †Retromalisus Kazantsev 2020
Fm. Brachypsectridae LeConte and Horn 1883 [Texas beetle]
+
G. †Vetubrachypsectra Qu and Cai 2019
+
†Vetubrachypsectra burmitica Qu and Cai 2019
†Vetubrachypsectra huchengi Li et al. 2022
Fm. Cantharidae Imhoff 1856 [soldier beetle]
+
Subfm. Cantharinae Imhoff 1856
+
G. †Brevipterus Yang et al. 2022
G. †Cnathrion Kazantsev and Perkovsky 2019
Tr. Podabrini Gistel 1856
Subfm. Chauliognathinae LeConte 1861
+
Tr. Chauliognathini Leconte 1861
Tr. Ichthyurini Champion 1915
G. †Katyacantharis Kazantsev and Perkovsky 2019
+
†Katyacantharis zherikhini Kazantsev and Perkovsky 2019
Subfm. Malthininae Kiesenwetter 1852
+
G. †Archaeomalthodes Hsiao et al. 2016
G. †Kuskaella Fanti and Kupryjanowicz 2017
Tr. Malchinini Brancucci 1980
Tr. Malthinini Kiesenwetter 1852
Tr. Malthodini Böving and Craighead 1931
Tr. †Mimoplatycini Kazantsev 2013
Tr. †Nothotytthonychini Fanti 2022
Invalid names: Cantharidiae Latreille 1802 [replaced], Dysmorphocerinae Brancucci 1980 [empty], Telephoridae [replaced]
Fm. Cerophytidae Latreille 1834
+
G. †Amberophytum Yu et al. 2019
+
†Amberophytum birmanicum Yu et al. 2019
†Amberophytum maculatum Poinar et al. 2021
G. †Aphytocerus Zherikhin 1977
+
†Aphytocerus communis Zherikhin 1977
†Aphytocerus dolganicus Zherikhin 1977
G. †Necromera Martynov 1926
+
†Necromera admiranda Chang and Kirejtshuk 2011
†Necromera baeckmanni Martynov 1926
†Necromera brevicornis Dolin 1980
†Necromera inflata Dolin 1980
†Necromera intermedia Dolin 1980
†Necromera latissima Dolin 1980
†Necromera longa Hong and Wang 1990
†Necromera longicornis Dolin 1980
†Necromera muscula Dolin 1980
Invalid names: Idiomerus Dolin 1980 [synonym], Leptocnemus Hong and Wang 1990 [synonym]
Invalid names: Aphytocerinae Kirejtshuk 2013 [invalid subgroup], Praelateriidae Dolin 1973 [synonym]
Fm. †Cretophengodidae Li et al. 2021
+
Fm. Elateridae Leach 1815 [click beetle]
+
Subfm. Agrypninae Candèze 1857
+
G. †Cretopachyderes Kundrata et al. 2021
Tr. †Cryptocardiini Dolin 1980
Tr. Hemirhipini Candèze 1857
Tr. Oophorini Gistel 1848
Tr. Pseudomelanactini Arnett 1967
Tr. Pyrophorini Candèze 1863
Invalid names: Drilini Blanchard 1845 [empty]
G. †Cryptocoelus Dolin and Nel 2002
+
†Cryptocoelus baissensis Alexeev 2011
†Cryptocoelus buffoni Dolin and Nel 2002
†Cryptocoelus dolini Alexeev 2011
†Cryptocoelus gianteus Chang et al. 2007
†Cryptocoelus lukashevichae Alexeev 2011
†Cryptocoelus major Dolin and Nel 2002
†Cryptocoelus shcherbakovi Alexeev 2011
†Cryptocoelus sinitshenkovae Alexeev 2011
Subfm. Dendrometrinae Gistel 1848
+
Tr. Dendrometrini Gistel 1848
Tr. Dimini Candèze 1863
Tr. Oxynopterini Candèze 1857
Tr. Prosternini Gistel 1856
Tr. Semiotini Jacobson 1913
Invalid names: Plastocerini Crownson 1972 [empty]
G. †Elateridium Tillyard 1918
+
†Elateridium angustius Tillyard 1918
†Elateridium subulatum Dunstan 1923
†Elateridium transversum Dunstan 1923
†Elateridium wianamattense Tillyard 1916
Invalid names: Elaterites Tillyard 1916 [replaced]
Subfm. Elaterinae Leach 1815
+
Tr. Agriotini Laporte 1840
G. †Ajkaelater Szabó et al. 2022
Tr. Ampedini Gistel 1848
G. †Crioraphes Yablokov-Khnzorian 1961
Tr. Elaterini Leach 1815
Tr. Megapenthini Gurjeva 1973
Tr. Melanotini Candèze 1859
Tr. Physorhinini Candèze 1859
Tr. Pomachiliini Candèze 1859
G. †Elaterites Heer 1847
+
†Elaterites amissus Heer 1847
†Elaterites bruchi Cockerell 1926
†Elaterites dicrepidioides Deichmüller 1881
†Elaterites laconoides Cockerell 1920
†Elaterites lavateri Heer 1847
†Elaterites longus Haupt 1956
†Elaterites microstictus Cockerell 1926
†Elaterites murchisoni Giebel 1856
†Elaterites obsoletus Heer 1847
†Elaterites palaeophilus Cockerell 1920
†Elaterites perditulus Cockerell 1920
†Elaterites sculptilis Cockerell 1920
Subfm. Hypnoidinae Schwarz 1906
+
G. Cryptohypnus Eschscholtz 1830
G. Hypnoidus Dillwyn 1829
G. Ligmargus Stibick 1976
Subfm. Negastriinae Nakane and Kishii 1956
+
G. †Ganestrius Dolin 1976
Tr. Negastriini Nakane and Kishii 1956
G. Paradonus Stibick 1971
G. †Protoquasimus Dolin 1976
G. †Paralithomerus Chang et al. 2008
+
†Paralithomerus exquisitus Chang et al. 2008
†Paralithomerus parallelus Chang et al. 2008
Subfm. Pityobiinae Hyslop 1917
+
G. †Cretopityobius Otto 2019
Invalid names: Tibionemini Motyka et al. 2023 [empty]
Subfm. †Protagrypninae Dolin 1973
+
Tr. †Desmatini Dolin 1975
Tr. †Hypnomorphini Dolin 1975
Tr. †Pollostelaterini Alexeev 2011
Tr. †Protagrypnini Dolin 1973
G. †Protocardiophorus Dolin 1976
+
†Protocardiophorus ancestralis Dolin 1976
†Protocardiophorus jurassicus Dolin 1980
G. †Pseudocardiophorites Dolin 1976
+
†Pseudocardiophorites angustatus Dolin 1980
†Pseudocardiophorites fragilis Dolin 1976
†Pseudocardiophorites hayeki Dolin 1976
†Pseudocardiophorites infractus Dolin 1976
†Pseudocardiophorites quadricollis Dolin 1976
Invalid names: Acmaeodera burmitina Cockerell 1917 [nomen dubium], Cebrionidae Latreille 1802 [replaced], Cebrioninae Latreille 1802 [empty], Elater maculatus Gistl 1831 [nomen dubium], Hemiopinae Fleutiaux 1941 [empty], Omalisinae Lacordaire 1857 [empty], Physodactylinae Lacordaire 1857 [empty], Subprotelaterinae Fleutiaux 1920 [empty], Tetralobinae Laporte 1840 [empty]
Fm. Eucnemidae Eschscholtz 1829 [false click beetle]
+
Subfm. Eucneminae Eschscholtz 1829
+
Tr. Dendrocharini Muona 1993
Tr. Dyscharachthini Muona 1993
Tr. Eucnemini Eschscholtz 1829
G. †Falsothambus Muona 2020
Tr. Mesogenini Muona 1993
Tr. Proutianini Muona 1993
G. †Yallmoleia Li et al. 2025
Subfm. Macraulacinae Fleutiaux 1923
+
Tr. Echthrogasterini Cobos 1965
Tr. Euryptychini Mamaev 1976
Tr. Jenibuntorini Muona 1993
Tr. Macraulacini Fleutiaux 1923
Tr. Nematodini Leiler 1976
Tr. Orodotini Muona 1993
Tr. †Throscogeniini Yablokov-Khnzorian 1962
G. †Xanrofeurya Li et al. 2025
Subfm. Melasinae Fleming 1821
+
Tr. Calyptocerini Muona 1993
Tr. Dirhagini Reitter 1911
Tr. Epiphanini Muona 1993
Tr. Melasini Fleming 1821
Tr. Neocharini Muona 1993
Tr. Xylobiini Reitter 1911
Subfm. Palaeoxeninae Muona 1993
+
G. †Anoixis Chang et al. 2010
G. †Cupressicharis Muona 2020
G. Palaeoxenus Horn 1891
Subfm. Schizophilinae Muona 1993
+
G. †Apoclion Chang et al. 2010
G. †Paradesmatus Chang et al. 2009
Invalid names: Perothopinae Lacordaire 1857 [empty], Phlegoninae Muona 1993 [empty], Phyllocerinae Reitter 1905 [empty], Pseudomeninae Muona 1993 [empty]
Fm. Lampyridae Rafinesque 1815
+
Subfm. Lamprohizinae Kazantsev 2010
+
G. †Eolamprohiza Kazantsev 2024
G. Lamprohiza Motschulsky 1853
G. Phausis Le Conte 1851
Subfm. Lampyrinae Rafinesque 1815
+
Tr. Lampyrini Rafinesque 1815
Tr. Lucidotini Lacordaire 1857
Tr. Photinini LeConte 1881
Subfm. Luciolinae Lacordaire 1857
+
G. †Eoluciola Kazantsev 2012
G. †Flammarionella Cai et al. 2024
G. Luciola Laporte 1833
G. †Protoluciola Kazantsev 2015
Invalid names: Luciolini Lacordaire 1857 [invalid subgroup]
Subfm. Ototretinae McDermott 1964
+
G. †Electrotreta Kazantsev 2012
Invalid names: Ototretadrilinae Crowson 1972 [synonym]
Invalid names: Amydetinae Olivier 1907 [empty], Cyphonocerinae Crowson 1972 [empty], Photurinae Lacordaire 1857 [empty], Pterotinae LeConte 1861 [empty]
Fm. Lycidae Laporte 1836 [net-winged beetle]
+
Subfm. Dexorinae Bocak and Bocakova 1989
+
Tr. †Burmolycini Bocak et al. 2019
Tr. †Cretolycini Tihelka et al. 2019
Subfm. Erotinae LeConte 1881
+
Tr. Dictyopterini Houlbert 1922
Tr. Erotini LeConte 1881
Tr. Lopherotini Kazantsev 2012
Invalid names: Dictyopterinae Houlbert 1922 [synonym]
Subfm. Lycinae Laporte 1836
+
Tr. Calopterini Green 1949
Tr. Lycini Laporte 1836
Tr. Platerotini Kleine 1929
Invalid names: Metriorrhynchini Kleine 1926 [empty]
Invalid names: Ateliinae Kleine 1928 [empty], Calochrominae Lacordaire 1857 [empty], Leptolycinae Leng and Mutchler 1922 [empty], Metriorrhynchinae Kleine 1926 [empty]
Fm. †Mysteriomorphidae Alekseev and Ellenberger 2019
+
G. †Mysteriomorphus Alekseev and Ellenberger 2019
+
†Mysteriomorphus longelytrus Qu et al. 2023
†Mysteriomorphus pelevini Alekseev and Ellenberger 2019
Invalid names: Trapezioceps Qu et al. 2023 [synonym]
Fm. Omethidae Leconte 1861
+
Invalid names: Driloniinae Crowson 1972 [empty], Matheteinae LeConte 1881 [empty], Omethinae LeConte 1861 [empty], Telegeusinae Leng 1920 [empty]
Fm. Throscidae Laporte 1840
+
G. Pactopus Leconte 1868
+
†Pactopus americanus Wickham 1914
†Pactopus avitus Britton 1960
†Pactopus burmensis Muona 2019
†Pactopus fafneri Muona 1993
†Pactopus fasolti Muona 1993
G. Potergus de Bonvouloir 1871
+
†Potergus frohi Muona 1993
†Potergus logei Muona 1993
†Potergus superbus Muona 2019
G. Trixagus Kugelann 1794
+
†Trixagus barclayi Kirejtshuk 2019
†Trixagus majusculus Kovalev et al. 2012
†Trixagus parvulus Muona 2019
Trixagus sericeus Le Conte 1868
†Trixagus sosnovskyi Yablokov-Khnzorian 1962
Invalid names: Palaeothroscus Yablokov-Khnzorian 1962 [synonym], Throscus Latreille 1796 [synonym]
Invalid names: Megacephalites Hong 2002 [nomen nudum], Palaeothroscini Yablokov-Khnzorian 1962 [invalid subgroup], Throscinae Laporte 1840 [empty], Trixagidae Gistel 1848 [synonym]
Invalid names: Artematopoidea Lacordaire 1857 [synonym], Eudicronychinae Girard 1971 [empty], Rhagophthalmidae Olivier 1907 [empty]
Invalid names: Mesosperchus angulatus Ponomarenko 1985 [nomen dubium], Podabrocephalidae Pic 1930 [empty]
G. †Jingxidiscus Hong 1990
+
†Jingxidiscus lushangfenensis Hong 1981
Invalid names: Discus Hong 1981 [replaced]
Fm. Jurodidae Ponomarenko 1985
+
G. †Jurodes Ponomarenko 1985
+
†Jurodes daohugouensis Yan et al. 2014
†Jurodes ignoramus Ponomarenko 1985
†Jurodes minor Ponomarenko 1990
†Jurodes pygmaeus Yan et al. 2014
†Jurodes shartegiensis Yan 2014
†Jurodes shef Yan et al. 2021
Invalid names: Sikhotealiniidae Lafer 1996 [synonym]
G. †Larvula Ponomarenko 1985
+
†Larvula cassa Ponomarenko 1985
†Larvula triassica Prokin and Ponomarenko 2013
Fm. †Lasiosynidae Kirejtshuk et al. 2010
+
G. †Anacapitis Yan 2009
+
†Anacapitis incertus Yan 2009
†Anacapitis karataviensis Yan 2009
†Anacapitis oblongus Yan 2009
†Anacapitis plata Tan and Ren 2009
Invalid names: Brachysyne Tan and Ren 2009 [synonym]
G. †Glaphyropterodes Handlirsch 1906
+
†Glaphyropterodes gehreti Heer 1852
†Glaphyropterodes sibiricus Ponomarenko 1985
G. †Lasiosyne Tan et al. 2007
+
†Lasiosyne cataphracta Yan 2012
†Lasiosyne daohugouensis Kirejtshuk et al. 2010
†Lasiosyne decora Yan 2012
†Lasiosyne euglyphea Tan et al. 2007
+
Invalid names: Pappisyne eucallus Tan and Ren 2009 [synonym], Pappisyne lasiospatha Tan and Ren 2009 [synonym]
†Lasiosyne fedorenkoi Kirejtshuk et al. 2010
†Lasiosyne gratiosa Kirejtshuk et al. 2010
†Lasiosyne insculpta Yan 2012
†Lasiosyne integera Guo et al. 2022
†Lasiosyne longitarsus Yan 2012
†Lasiosyne multituberata Lin 1980
†Lasiosyne parva Guo et al. 2022
†Lasiosyne quadricollis Kirejtshuk et al. 2010
Invalid names: Pappisyne Tan and Ren 2009 [synonym]
Invalid names: Lasiosyne laxa Yan et al. 2014 [nomen dubium]
Unr. Nosodendriformia Cai and Tihelka 2022
+
Superfm. Nosodendroidea Erichson 1846
+
Fm. Nosodendridae Erichson 1846 [wounded tree beetle]
+
G. †Archaenosodendron Li and Cai 2021
+
†Archaenosodendron angulare Li and Cai 2021
†Archaenosodendron cretaceum Deng et al. 2019
†Archaenosodendron explanatum Li and Cai 2021
†Archaenosodendron remotidens Li and Cai 2021
G. Nosodendron Latreille 1804
G. †Plastelater Handlirsch 1906
+
†Plastelater glabratus Ponomarenko 1985
†Plastelater neptuni Giebel 1856
G. †Polysitum Dunstan 1923
+
†Polysitum elongatum Ponomarenko 1985
†Polysitum hosbayari Ponomarenko 1986
†Polysitum kuznetskiense Rohdendorf 1961
†Polysitum latum Ponomarenko 1985
†Polysitum majus Ponomarenko 1990
†Polysitum minutus Dunstan 1923
†Polysitum ovale Ponomarenko 1990
†Polysitum punctatus Dunstan 1923
†Polysitum rugosum Ponomarenko 1986
†Polysitum sharategense Ponomarenko 2014
†Polysitum wudenghaoensis Hong 1976
Unr. Rhinorhipiformia Cai et al. 2022
+
Unr. Scirtiformia Lawrence et al. 2011
+
Superfm. Scirtoidea Fleming 1821
+
Fm. †Boleopsidae Kirejtshuk and Nel 2013
+
Fm. Scirtidae Fleming 1821 [marsh beetle]
+
Subfm. Scirtinae Fleming 1821
+
G. Elodes Latreille 1796
G. Microcara Thomson 1859
G. Scirtes Illiger 1807
G. †Varcalium Li et al. 2022
Invalid names: Cyphoninae Stephens 1829 [synonym], Cyphonini Stephens 1829 [invalid subgroup], Helodinae Shuckard 1839 [synonym], Helodopsini Yablokov-Khnzorian 1961 [invalid subgroup], Microcarini Yablokov-Khnzorian 1961 [invalid subgroup]
Invalid names: Elodiidae Shuckard 1839 [synonym], Sinodryopitidae Hong 2002 [invalid subgroup]
Invalid names: Decliniidae Nikitsky et al. 1994 [empty]
G. †Sphaerocantharis Bode 1953
+
†Sphaerocantharis defossa Bode 1953
†Sphaerocantharis sibirica Ponomarenko 1990
Invalid names: Mikrocarpides Bode 1953 [synonym], Theornithion Bode 1953 [synonym]
Infraor. Staphyliniformia Lameere 1900
+
Superfm. Histeroidea Gyllenhal 1808
+
Fm. †Cretohisteridae Zhou et al. 2018
+
Fm. Histeridae Gyllenhal 1808 [clown beetle]
+
Subfm. Abraeinae MacLeay 1819
+
Tr. Abraeini MacLeay 1819
Tr. Acritini Wenzel 1944
Tr. †Pantostictini Zhou et al. 2020
G. †Pseudacritus Simon-Pražák et al. 2024
G. †Amplectister Caterino and Maddison 2018
+
†Amplectister tenax Caterino and Maddison 2018
†Amplectister terapoides Yamamoto and Caterino 2022
Subfm. Dendrophilinae Reitter 1909
+
Tr. Anapleini Olexa 1982
Tr. Bacaniini Kryzhanovskij 1976
G. †Druantia Caterino 2021
G. †Olexum Simon Pražák and Lackner 2023
Tr. Paromalini Reitter 1909
G. †Yethiha Caterino 2021
Subfm. Histerinae Gyllenhal 1808
+
Tr. Histerini Gyllenhal 1808
G. †Platycretus Simon Pražák and Lackner 2023
Subfm. Onthophilinae MacLeay 1819
+
G. †Carinumerus Caterino 2021
G. †Cretonthophilus Caterino et al. 2015
G. Onthophilus Leach 1817
G. †Phasmister Caterino 2021
Invalid names: Chlamydopsinae Bickhardt 1914 [empty], Hetaeriinae Marseul 1857 [empty], Niponiinae Fowler 1912 [empty], Tribalinae Bickhardt 1914 [empty], Trypeticinae Bickhardt 1914 [empty]
Invalid names: Sphaeritidae Shuckard 1839 [empty]
Superfm. Hydrophiloidea Latreille 1802 [water scavenger beetle]
+
Fm. Helophoridae Leach 1815
+
G. Helophorus Fabricius 1775
+
Subg. Helophorus (Atracthelophorus) Kuwert 1890
Subg. Helophorus (Gephelophorus) Sharp 1915
Subg. Helophorus (Meghelophorus) Kuwert 1886
+
Invalid names: Helophorus (Praehelephorus) Lomnicki 1894 [synonym]
Subg. †Helophorus (Mesohelophorus) Ponomarenko 1977
Subg. †Helophorus (Mesosperchus) Ponomarenko 1977
Subg. Helophorus (Orphelophorus) d'Orchymont 1927
Subg. Helophorus (Rhopalhelophorus) Kuwert 1886
Subg. Helophorus (Trichelophorus) Kuwert 1886
Helophorus auricollis Eschscholtz 1822
Helophorus brevipalpis Bedel 1881
Helophorus browni McCorkle 1973
Helophorus discrepans Rey 1885
Helophorus eclectus d'Orchymont 1945
Helophorus lacustris Le Conte 1850
Helophorus nubilus Fabricius 1777
Helophorus oblongus Le Conte 1850
+
Invalid names: Helophorus (Rhopalhelophorus) kuwerti Lomnicki 1894 [synonym]
Helophorus orientalis Motschulsky 1860
Helophorus sempervarians Angus 1970
Helophorus splendidus Sahlberg 1880
Helophorus tuberculatus Gyllenhal 1808
Fm. Hydrochidae Thomson 1859
+
G. †Ponohydrochus Prokin and Strelnikova 2021
+
†Ponohydrochus buryaticus Prokin and Strelnikova 2021
Fm. Hydrophilidae Latreille 1802 [water scavenger beetle]
+
Subfm. Acidocerinae Zaitzev 1908
+
G. Helochares Mulsant 1846
G. †Hydroyixia Fikáček et al. 2014
Invalid names: Horelophopsinae Hansen 1997 [synonym]
G. †Aposphinctus Bode 1953
+
†Aposphinctus sibiricus Ponomarenko 1990
Invalid names: Hexameristus Bode 1953 [synonym], Spalacoides Bode 1953 [synonym], Streblocardioides Bode 1953 [synonym]
G. Hydrocharis Berthold 1827
Subfm. Hydrophilinae Latreille 1802
+
Tr. Berosini Mulsant 1844
Tr. Hydrophilini Latreille 1802
Tr. Laccobiini Houlbert 1922
Invalid names: Amphiopini Kuwert 1890 [empty], Cymbiodyta austerus Statz 1939 [nomen dubium]
G. †Paraspercheus Ponomarenko 1977
+
†Paraspercheus asiaticus Ponomarenko 1977
†Paraspercheus vitimensis Ponomarenko 1977
Subfm. Rygmodinae Orchymont 1916
+
Invalid names: Andotypini Hansen 1991 [synonym], Borborophorini Hansen 1991 [synonym], Tormissini Hansen 1991 [synonym]
Subfm. Sphaeridiinae Latreille 1802
+
Tr. Coelostomatini Heyden 1891
Tr. Megasternini Mulsant 1844
Tr. Sphaeridiini Latreille 1802
Invalid names: Omicrini Smetana 1975 [empty], Protosternini Hansen 1991 [empty]
Invalid names: Escheria Heer 1847 [nomen dubium], Escheria bella Heer 1862 [nomen dubium], Escheria ovalis Heer 1847 [nomen dubium], Escheriidae Kolbe 1880 [synonym], Mesosperchus schultzi Ponomarenko 1985 [nomen dubium], Spercheinae Erichson 1837 [empty], Sphaeridium melanarium Gistl 1831 [nomen dubium]
G. †Laetopsia Fikáček et al. 2012
+
†Laetopsia baissensis Ponomarenko 1987
†Laetopsia bontsaganica Prokin 2009
†Laetopsia hydraenoides Prokin et al. 2010
†Laetopsia mongolica Ponomarenko 1987
†Laetopsia shatrovskiyi Prokin et al. 2010
Fm. Spercheidae Erichson 1837
+
G. Spercheus Kugelann 1798
+
†Spercheus punctatus Ponomarenko 2019
†Spercheus wightensis Ponomarenko 2019
Invalid names: Epimetopidae Zaitzev 1908 [empty], Georissidae Laporte 1819 [empty]
Superfm. Scarabaeoidea Latreille 1802 [scarab beetle]
+
Fm. †Alloioscarabaeidae Bai et al. 2012
+
Fm. Bolboceratidae Mulsant 1842 [dor beetle]
+
Fm. Diphyllostomatidae Holloway 1972
+
Fm. Geotrupidae Latreille 1802 [earth-boring dung beetle]
+
Subfm. Geotrupinae Latreille 1802
+
G. Ceratophyus Fischer von Waldheim 1823
Tr. †Cretogeotrupini Nikolajev 1996
Tr. Geotrupini Latreille 1802
G. †Lithogeotrupes Nikolajev 2008
G. †Parageotrupes Nikolajev and Ren 2010
G. Typhaeus Leach 1815
Invalid names: Lethrinae Mulsant and Rey 1871 [empty], Taurocerastinae [empty]
G. †Geotrupoides Handlirsch 1906
+
†Geotrupoides jiaoheense Hong 1992
†Geotrupoides kezuoensis Hong 1992
†Geotrupoides lithographicus Deichmüller 1886
†Geotrupoides saxosus Zhang 1997
†Geotrupoides songyingziensis Hong 1984
Fm. Glaphyridae MacLeay 1819 [bumble bee scarab beetle]
+
G. †Cretohypna Yan et al. 2012
+
†Cretohypna cristata Yan et al. 2012
†Cretohypna puncta Zhao et al. 2016
†Cretohypna robusta Zhao et al. 2016
†Cretohypna yixianensis Nikolajev and Ren 2015
G. †Lithohypna Nikolajev et al. 2011
+
†Lithohypna chifengensis Nikolajev et al. 2011
†Lithohypna laoningensis Nikolajev 2014
†Lithohypna lepticephala Nikolajev and Ren 2012
†Lithohypna longula Nikolajev and Ren 2012
†Lithohypna tuberculata Nikolajev and Ren 2012
†Lithohypna yuxiana Nikolajev and Ren 2012
G. †Mesohypna Nikolajev and Ren 2013
+
†Mesohypna lopatini Nikolajev and Ren 2013
†Mesohypna probata Nikolajev 2014
G. †Paraglaphyrus Nikolajev 2015
+
†Paraglaphyrus ovalis Nikolajev 2015
†Paraglaphyrus robustus Nikolajev 2015
†Paraglaphyrus subtilis Nikolajev 2015
†Paraglaphyrus yixianensis Nikolajev 2015
Fm. Glaresidae Prudhomme de Borre 1886 [enigmatic scarab beetle]
+
G. †Cretoglaresis Nikolajev 2007
+
†Cretoglaresis nana Nikolajev 2007
†Cretoglaresis ovalis Nikolajev 2009
G. Glaresis Erichson 1848
+
†Glaresis burmitica Cai and Huang 2018
†Glaresis cretacea Nikolajev 2007
†Glaresis orthochilus Bai et al. 2010
†Glaresis tridentata Bai et al. 2014
Invalid names: Aphodiites Heer 1864 [nomen dubium]
G. †Holcorobeus Nikritin 1977
+
†Holcorobeus atrosulcatus Nikolajev 1992
†Holcorobeus evittatus Zhang 1992
†Holcorobeus nigrovittatus Nikolajev 1992
†Holcorobeus vittatus Nikritin 1977
Invalid names: Holcorobeus (Holcorobeus) Nikritin 1977 [invalid subgroup]
G. †Hongscarabaeus Krell 2000
+
†Hongscarabaeus brunneus Hong 1982
Invalid names: Proscarabaeus Hong 1982 [replaced]
Fm. Hybosoridae Erichson 1847 [scavenger scarab beetle]
+
Subfm. Anaidinae Nikolajev 1996
+
G. Anaides Westwood 1842
G. †Crassisorus Nikolajev et al. 2012
G. †Cretanaides Nikolajev 1996
G. †Protanaides Nikolajev 2010
Subfm. Ceratocanthinae Martinez 1968
+
G. †Palaeopycnus Lu et al. 2022
Invalid names: Ivieolini Howden and Gill 2000 [empty]
Subfm. Hybosorinae Erichson 1847
+
G. †Cretohybosorus Nikolajev 1999
G. †Fortishybosorus Yan et al. 2013
G. Hybosorus MacLeay 1819
G. †Jurahybosorus Nikolajev 2005
G. †Leptosorus Nikolajev 2006
G. Phaeochrous Laporte 1840
G. †Protohybosorus Nikolajev 2010
Invalid names: Anaidini Nikolajev 1996 [empty], Hybosorini Erichson 1847 [empty]
Fm. Lucanidae Latreille 1804 [stag beetle]
+
Subfm. Aesalinae MacLeay 1819
+
Tr. Aesalini MacLeay 1819
Tr. Ceratognathini Sharp 1899
Tr. Nicagini LeConte 1861
G. †Cretolucanus Nikolajev 2007
+
†Cretolucanus longus Nikolajev 2007
†Cretolucanus ordinarius Nikolajev 2007
†Cretolucanus sibericus Nikolajev 2007
Subfm. Lampriminae MacLeay 1819
+
G. †Electraesalopsis Bai et al. 2017
G. †Prostreptocerus Yu and Cai 2024
G. †Protognathinus Chalumeau and Brochier 2001
Subfm. Lucaninae Latreille 1804
+
G. †Anisoodontus Wu et al. 2022
G. †Miocenidorcus Riou 1999
Tr. Platycerini Mulsant 1842
G. †Prolucanus Qi et al. 2022
Invalid names: Chiasognathini Burmeister 1847 [empty], Lucanus fossilis Wickham 1913 [nomen dubium], Penichrolucaninae Arrow 1950 [synonym], Platycerus sepultus Germar 1837 [nomen dubium]
G. †Mesoscarabaeus Hong 1982
+
†Mesoscarabaeus corneus Hong 1982
†Mesoscarabaeus morulosus Hong 1982
Fm. Ochodaeidae Streubel 1846 [sand-loving scarab beetle]
+
Subfm. †Cretochodaeinae Nikolajev 1995
+
G. †Cretochodaeus Nikolajev 1995
G. †Mioochodaeus Nikolajev 1995
Fm. Passalidae Leach 1815 [bess beetle]
+
Subfm. Aulacocyclinae Kaup 1868
+
Subfm. Passalinae Leach 1815
+
Tr. Macrolinini Kaup 1871
Tr. Passalini Leach 1815
Tr. †Protopassalini Santos et al. 2020
Fm. †Passalopalpidae Boucher and Bai 2016
+
G. †Mesoceratocanthus Nikolajev et al. 2010
+
†Mesoceratocanthus tuberculifrons Nikolajev et al. 2010
Fm. Pleocomidae Leconte 1861 [rain beetle]
+
Fm. Scarabaeidae Latreille 1802 [scarab beetle]
+
Subfm. Aegialiinae Laporte 1840
+
G. Aegialia Latreille 1807
G. †Archeopsammoporus Minkina 2020
G. †Cretaegialia Nikolayev 1993
Subfm. Aphodiinae Leach 1815
+
Tr. Aphodiini Leach 1815
Tr. Eupariini Schmidt 1910
Tr. †Psammaegialiini Nikolajev et al. 2014
Tr. Psammodiini Mulsant 1842
Tr. Rhyparini Schmidt 1910
Subfm. Cetoniinae Leach 1815 [flower chafer]
+
Tr. Cetoniini Leach 1815
Tr. Schizorhinini Burmeister 1842
Tr. Trichiini Fleming 1821
Tr. Valgini Mulsant 1842
Subfm. †Cretoscarabaeinae Nikolajev 1995
+
G. †Cretorabaeus Nikolajev 1995
G. †Cretoscarabaeus Nikolajev 1995
Subfm. Dynastinae MacLeay 1819 [rhinoceras beetle]
+
G. †Calcitoryctes Krell 2011
G. Cyclocephala Dejean 1821
G. Ligyrus Burmeister 1847
Tr. Oryctini Mulsant 1842
G. †Oryctoantiquus Ratcliffe and Smith 2005
Tr. Pentodontini Mulsant 1842
Invalid names: Pentodon proserpinae Heer 1862 [nomen dubium]
Subfm. †Lithoscarabaeinae Nikolajev 1992
+
G. †Baisarabaeus Nikolajev 2005
G. †Lithoscarabaeus Nikolajev 1992
Subfm. Melolonthinae Leach 1819
+
Tr. †Cretomelolonthini Nikolajev 1998
Tr. Diplotaxini Kirby 1837
G. †Eophyllocerus Haupt 1950
Tr. Euchirini Hope 1840
Tr. Hopliini Latreille 1829
Tr. Macrodactylini Kirby 1837
Tr. Melolonthini Leach 1819
Tr. Rhizotrogini Burmeister 1855
Subfm. Rutelinae MacLeay 1819
+
Tr. Adoretini Burmeister 1844
Tr. Anomalini Streubel 1839
G. †Anomalites Frič 1885
Tr. Anoplognathini MacLeay 1819
G. †Pelidnotites Cockerell 1920
G. †Petraeianus Zhang 1989
G. †Petulantis Zhang et al. 1994
Subfm. Scarabaeinae Latreille 1802
+
G. Anachalcos Hope 1837
Tr. Ateuchini Perty 1830
G. Canthochilum Chapin 1934
Tr. Coprini Leach 1815
Tr. Deltochilini Lacordaire 1856
Tr. Gymnopleurini Lacordaire 1856
G. Heliocopris Hope 1837
G. Metacatharsius Paulian 1939
Tr. Oniticellini Kolbe 1905
Tr. Onitini Laporte 1840
Tr. Onthophagini Burmeister 1846
Tr. Phanaeini Hope 1838
Tr. Scarabaeini Latreille 1802
Invalid names: Onthophagus spitsbergeniensis Krell 2010 [nomen dubium]
Invalid names: Allidiostomatinae Arrow 1940 [empty], Coprophaginae [empty], Dynamopodinae Arrow 1911 [empty], Oniticellus amplicollis Heer 1862 [nomen dubium], Orphninae Erichson 1847 [empty], Pachypodinae Erichson 1840 [empty], Phaenomeridinae Erichson 1847 [empty]
Fm. †Septiventeridae Bai et al. 2013
+
Invalid names: Belohinidae Paulian 1959 [empty], Copris subterraneus Heer 1862 [nomen dubium], Gymnopleurus deperditus Heer 1862 [nomen dubium], Holcorobeini Nikolajev 1992 [invalid subgroup], Onthophagus urusheeri Krell 2000 [nomen dubium]
Superfm. Staphylinoidea Latreille 1802
+
Fm. Agyrtidae Thomson 1859 [primitive carrion beetle]
+
G. †Mesecanus Newton 1982
+
†Mesecanus communis Ponomarenko 1977
†Mesecanus lintouensis Lin 1985
Invalid names: Mesagyrtes Ponomarenko 1977 [replaced]
Fm. Colonidae Horn 1880
+
Fm. Hydraenidae Mulsant 1844 [minute moss beetle]
+
Subfm. Hydraeninae Mulsant 1844
+
G. †Archaeodraena Jäch and Yamamoto 2017
G. †Davidraenites Kirejtshuk 2019
G. †Eolimnebius Kirejtshuk 2019
Tr. Hydraenini Mulsant 1844
G. †Hydraenites Kirejtshuk 2019
Tr. Limnebiini [minute moss beetle]
G. †Metacoxites Kirejtshuk 2019
G. †Ochtebiites Ponomarenko 1977
+
†Ochtebiites altus Ponomarenko 1977
†Ochtebiites incertus Ponomarenko 1977
†Ochtebiites manlaicus Ponomarenko 1980
†Ochtebiites minor Ponomarenko 1985
Invalid names: Prosthetopinae Perkins 1994 [empty]
Fm. Leiodidae Fleming 1821 [round fungus beetle]
+
Subfm. Cholevinae Kirby 1837
+
Tr. Anemadini Hatch 1927
Tr. Cholevini Kirby 1837
Tr. Oritocatopini Jeannel 1936
Tr. Ptomaphagini Jeannel 1911
Subfm. Leiodinae Fleming 1821
+
Tr. Leiodini Fleming 1821
Tr. Pseudoliodini Portevin 1926
Tr. Scotocryptini
Tr. Sogdini Lopatin 1961
Invalid names: Leptininae LeConte 1866 [empty], Platypsyllinae Ritsema 1869 [empty]
Fm. Ptiliidae Erichson 1845 [feather-winged beetle]
+
Subfm. Ptiliinae Erichson 1845
+
Tr. Acrotrichini Reitter 1909
Tr. Discheramocephalini Grebennikov 2009
Tr. Ptenidiini Flach 1889
Tr. Ptiliini Erichson 1845
Tr. Ptinellini Reitter 1906
Invalid names: Nanosellini Barber 1924 [empty]
Fm. Staphylinidae Latreille 1802 [rove beetle]
+
Subfm. Aleocharinae Fleming 1821
+
Tr. Aleocharini Fleming 1821
Tr. Athetini Casey 1910
Tr. Corotocini Fenyes 1918
Tr. Homalotini Heer 1839
Tr. Hygronomini Thomson 1859
Tr. Hypocyphtini Laporte 1835
Tr. Lomechusini Fleming 1821
Tr. Mesoporini Cameron 1959
Tr. Oxypodini Thomson 1859
Tr. Trichopseniini Le Conte and Horn 1883
G. †Aleocharopsis Wickham 1913
+
†Aleocharopsis caseyi Wickham 1913
†Aleocharopsis secunda Wickham 1913
G. †Apticax Schomann and Solodovnikov 2012
+
†Apticax solidus Schomann and Solodovnikov 2012
†Apticax volans Schomann and Solodovnikov 2012
G. Boletobius Leach 1819
G. †Cardiolita Strand 1933
Subfm. Dasycerinae Reitter 1887
+
G. †Cedasyrus Yin and Cai 2020
G. †Protodasycerus Yamamoto 2016
G. †Vetudasycerus Cai et al. 2017
G. Ellipsotomus Motschulsky 1845
Subfm. Euaesthetinae Thomson 1859
+
Tr. Euaesthetini Thomson 1859
Tr. Nordenskioldiini Bernhauer and Schubert 1911
G. Heterosoma Bernhauer 1903
G. †Laostaphylinus Zhang 1988
+
†Laostaphylinus fuscus Zhang 1988
†Laostaphylinus nigritellus Zhang 1988
Subfm. Micropeplinae Leach 1815
+
G. Arrhenopeplus Koch 1937
G. Kalissus Leconte 1874
G. Micropeplus Latreille 1809
G. †Protopeplus Cai and Huang 2014
Subfm. Mycetoporinae Thomson 1859
+
G. Bolitobius Leach 1819
G. †Cretafrica Mnguni et al. 2024
G. †Cuneocharis Ryvkin 1990
G. †Glabrimycetoporus Yue et al. 2009
G. Ischnosoma Stephens 1829
G. Lordithon Thomson 1859
G. Mycetoporus Mannerheim 1830
G. Parabolitobius Li et al. 2000
G. †Undiatina Ryvkin 1990
Invalid names: Bolitobiini Horn 1877 [synonym]
Subfm. Omaliinae MacLeay 1825
+
G. †Archodromus Tikhomirova 1968
Tr. Coryphiini Jakobson 1908
G. †Daiodromus Ryvkin 1990
G. †Eophyllodrepa Ryvkin 1985
Tr. Eusphalerini Hatch 1957
G. †Globoides Tikhomirova 1968
G. †Mesodeliphrum Ryvkin 1990
G. †Morsum Ryvkin 1985
Tr. Omaliini MacLeay 1825
G. †Porrhodromus Tikhomirova 1968
G. †Prodaia Ryvkin 1990
G. †Protostaphylinus Lin 1976
G. †Pseudolesteua Schaufuss 1890
Subfm. Osoriinae Erichson 1839
+
Tr. Eleusinini Sharp 1887
Tr. Leptochirini Sharp 1887
Tr. Osoriini Erichson 1839
G. †Sinolispinodes Zhang 1989
Subfm. Oxyporinae Fleming 1821
+
G. †Cretoxyporus Cai and Huang 2014
G. Oxyporus Fabricius 1775
G. †Protoxyporus Cai and Huang 2014
Subfm. Oxytelinae Fleming 1821
+
Tr. Blediini Ádám 2001
Tr. Coprophilini Heer 1839
Tr. Deleasterini Reitter 1909
G. †Megalymma Tikhomirova 1980
G. †Mesoxytelus Tikhomirova 1968
G. †Morda Ryvkin 1990
Tr. Oxytelini Fleming 1821
Tr. Syntomiini Böving and Craighead 1931
Tr. Thinobiini Sahlberg 1876
G. †Turgaphloeus Ryvkin 1990
Invalid names: Euphaniini Reitter 1909 [empty], Piestini Erichson 1839 [empty]
Subfm. Paederinae Fleming 1821
+
Tr. Lathrobiini Laporte 1835
G. †Mesostaphylinus Zhang 1988
Tr. Paederini Fleming 1821
Tr. Pinophilini Nordmann 1837
Subfm. Phloeocharinae Erichson 1839
+
G. †Angucharcotes Li et al. 2022
G. Charhyphus Sharp 1887
+
Invalid names: Triga Fauvel 1878 [replaced], Trigites Handlirsch 1907 [synonym]
G. Phloeocharis Mannerheim 1830
G. Vicelva Moore and Legner 1973
Subfm. Piestinae Erichson 1839
+
G. †Eopiestus Cai and Lu 2016
G. †Paleosiagonium Yue et al. 2016
G. †Propiestus Yamamoto et al. 2018
Subfm. Pselaphinae Latreille 1802
+
Unr. Batrisitae Reitter 1882
Unr. Clavigeritae Leach 1815
G. †Duterotyrus Schaufuss 1890
Unr. Euplectitae Streubel 1839
Unr. Faronitae Reitter 1882
Unr. Goniaceritae Reitter 1882
G. †Greys Schaufuss 1890
G. †Penarhytus Peris et al. 2014
G. †Protrichonyx Parker 2016
G. Pselaphaulax Reitter 1909
Unr. Pselaphitae Latreille 1802
Subfm. Pseudopsinae Ganglbauer 1895
+
G. †Cretaceonanobius Liu et al. 2022
G. †Cretopseudopsis Liu et al. 2020
G. Pseudopsis Newman 1834
Subfm. Scaphidiinae Latreille 1807
+
Tr. Scaphidiini Latreille 1806
G. †Scaphidiopsis Handlirsch 1906
Tr. Scaphisomatini Casey 1893
Subfm. Scydmaeninae Leach 1815 [antlike stone beetle]
+
Unr. Cephenniitae Reitter 1882
G. †Cryptodiodon Schaufuss 1890
G. †Electroscydmaenus Schaufuss 1890
Unr. †Hapsomelitae Poinar and Brown 2004
G. †Hetereuthia Schaufuss 1890
G. †Heuretus Schaufuss 1890
Unr. Mastigitae Fleming 1821
G. †Microlynx Kohring and Schlüter 1989
G. †Palaeothia Schaufuss 1890
G. †Pinitoides Motschulsky 1856
Unr. Scydmaenitae Leach 1815
G. †Scydmaenoides Motschulsky 1856
G. †Semnodioceras Schaufuss 1890
Invalid names: Mastiginae Fleming 1821 [empty]
G. †Seniaulus von Heyden and von Heyden 1866
+
†Seniaulus scaphioides von Heyden and von Heyden 1866
Subfm. Silphinae Latreille 1807
+
Tr. Necrodini Portevin 1926
G. Oiceoptoma Leach 1815
G. Phosphuga Leach 1817
G. †Pliosilpha Gersdorf 1969
Tr. Silphini Latreille 1807
Subfm. Staphylininae Latreille 1802
+
Tr. Arrowinini Solodovnikov and Newton 2005
G. Chiquiticus Reyes-Hernández and Solodovnikov 2024
G. †Cretoprosopus Solodovnikov and Yue 2013
Tr. Diochini Casey 1906
G. †Megolisthaerus Solodovnikov and Yue 2010
Tr. Othiini Thomson 1859
G. †Paleothius Solodovnikov and Yue 2013
Tr. †Thayeralinini Solodovnikov and Yue 2013
Tr. Xantholinini Erichson 1839
Invalid names: Platyprosopinae Lynch Arribálzaga 1884 [synonym], Platyprosopini Lynch Arribálzaga 1884 [empty], Xantholininae Erichson 1839 [synonym]
Subfm. Steninae Macleay 1825
+
G. †Afristenus Mnguni et al. 2022
G. Dianous Leach 1819
G. †Eocenostenus Cai et al. 2014
G. †Festenus Żyła et al. 2017
Subfm. Tachyporinae MacLeay 1825
+
G. †Abscondus Tikhomirova 1968
Tr. Tachinusini Fleming 1821
Tr. Tachyporini MacLeay 1825
G. †Tachyporoides Tikhomirova 1968
Tr. Vatesini Seevers 1958
Invalid names: Acylophorus immotus Scudder 1900 [nomen dubium]
Subfm. Trigonurinae Reiche 1821
+
G. †Abolescus Tikhomirova 1968
G. †Kovalevia Ryvkin 1990
G. †Trigunda Ryvkin 1990
Invalid names: Empelinae Newton and Thayer 1992 [empty], Habrocerinae Mulsant and Rey 1877 [empty], Leptotyphlinae Fauvel 1874 [empty], Microsilphinae Crowson 1950 [empty], Neophoninae Fauvel 1905 [empty], Silphidae Latreille 1807 [synonym], Triga coeni Scudder 1900 [nomen dubium]
Invalid names: Dasyceridae Reitter 1887 [empty], Empelidae [empty]
Invalid names: Scarabaeiformia Crowson 1960 [synonym]
Invalid names: Agyrtes primoticus Scudder 1900 [nomen dubium], Derodontiformia Le Conte 1861 [invalid subgroup], Escheria crassipunctata Förster 1891 [nomen dubium], Escheria dimidiata Förster 1891 [nomen dubium], Escheria punctulata Förster 1891 [nomen dubium], Eucinetiformia [empty], Limnius perrini Theobald 1937 [nomen dubium], Malachius vertummni Heer 1847 [nomen dubium], Mesosperchus notatus Ponomarenko 1977 [nomen dubium], Tomocoleites Hong 2002 [nomen nudum], Troglops punctatissima Théobald 1935 [nomen dubium]
Diagnosis
No diagnoses are available