PBDB Taxon

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Euungulata

Mammalia

Taxonomy

Euungulata was named by Waddell et al. (2001). It is extant.

It was assigned to Fereuungulata by Waddell et al. (2001); and to Scrotifera by Asher and Helgen (2010).

Sister taxa

Altungulata, Artiodactylamorpha, Cetungulata, Haloboidea, Panperissodactyla, Tipotheria

Synonyms

Eparctocyona was named by McKenna (1975) [Mirorder]. It is extant.

It was synonymized subjectively with Euungulata by Asher and Helgen (2010).

It was assigned to Ungulata by McKenna (1975) and McKenna and Honey (2008).
Ungulata was named by Linnaeus (1766). It is extant.

It was reranked as the subclass Ungulata by Gray (1821); it was reranked as the order Ungulata by Gray (1825), Flower and Lydekker (1891), Parker and Haswell (1897), Gadow (1898), Hay (1902) and Lambe (1908); it was reranked as the order Ungulati by Jaekel (1911); it was reranked as the superorder Ungulata by Hay (1930), Thorpe (1937) and Prothero and Schoch (1989); it was considered an invalid subgroup of Eutheria by Madsen et al. (2001); it was synonymized subjectively with Euungulata by Asher and Helgen (2010).

It was assigned to Quadripedes by Gray (1821); to Educabilia by Bonaparte (1850); to Heterodonta by Ameghino (1889); to Monodelphia by Cope (1889); to Eutheria by Flower and Lydekker (1891); to Ditremata by Ameghino (1894); to Eutheria by Parker and Haswell (1897); to Monodelphia by Hay (1902); to Eutheria by Flower (1883), Gadow (1898) and Gregory (1910); to Holotheria by Jaekel (1911); to Monodelphia by Hay (1930) and Thorpe (1937); to Tokotheria by McKenna (1975); to Eutheria by Prothero and Schoch (1989); to Ungulatomorpha by Archibald (1998); to Mammalia by Gray (1825), Andrews (1906), Lambe (1908) and McKenna and Lofgren (2003); and to Epitheria by Shoshani and Tassy (2005).

Synonymy list

Year	Name and author
1766	Ungulata Linnaeus
1821	Ungulata Gray p. 305
1825	Ungulata Gray p. 342
1850	Ungulata Bonaparte p. 1
1883	Ungulata Flower p. 184
1889	Ungulata Ameghino p. 354
1889	Ungulata Cope p. 875
1891	Ungulata Flower and Lydekker p. 89
1894	Ungulata Ameghino p. 11
1897	Ungulata Parker and Haswell p. 452
1898	Ungulata Gadow p. 45
1902	Ungulata Hay p. 601
1906	Ungulata Andrews p. 1
1908	Ungulata Lambe
1910	Ungulata Gregory p. 466
1911	Ungulati Jaekel p. 237
1930	Ungulata Hay p. 600
1937	Ungulata Thorpe p. 31
1975	Eparctocyona McKenna p. 41
1975	Ungulata McKenna p. 42
1989	Ungulata Prothero and Schoch p. 530
1998	Ungulata Archibald
2001	Euungulata Waddell et al. p. 148 figs. Figure 3
2003	Ungulata McKenna and Lofgren p. 635
2005	Ungulata Shoshani and Tassy p. 9
2008	Eparctocyona McKenna and Honey p. 32
2010	Euungulata Asher and Helgen p. 3 figs. Figure 1

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Deuterostomia
phylum	Chordata	Haeckel 1874
subphylum	Vertebrata
superclass	Gnathostomata
—	Osteichthyes	()
subclass	Sarcopterygii	()
subclass	Dipnotetrapodomorpha	(Nelson 2006)
subclass	Tetrapodomorpha	()
—	Tetrapoda
—	Reptiliomorpha
—	Anthracosauria
subclass	Amphibiosauria	Kuhn 1967
—	Cotylosauria	()
—	Amniota

Rank	Name	Author
subclass	Synapsida
—	Therapsida	()
infraorder	Cynodontia	()
—	Mammaliamorpha
—	Mammaliaformes
class	Mammalia
—	Cladotheria
—	Zatheria
subclass	Tribosphenida	()
subclass	Theria
—	Eutheria	()
—	Placentalia
—	Boreoeutheria
—	Laurasiatheria
—	Scrotifera
—	Euungulata
—	Euungulata

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Unr. Euungulata Waddell et al. 2001

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Unr. Artiodactylamorpha Spaulding et al. 2009

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Unr. Artiodactyla Owen 1848 [even-toed ungulate]

Subfm. †Antiacodontinae

G. †Sarcolemur Cope 1875

Superfm. †Cainotherioidea Camp and VanDerHoof 1940

Fm. †Cainotheriidae Cope 1881

Subfm. †Cainotheriinae Camp and VanderHoof 1940

Subfm. †Oxacroninae Hurzeler 1936

G. †Palembertina Weppe et al. 2020

Fm. †Robiacinidae Weppe et al. 2019

G. †Robiacina Sudre 1969

Superfm. †Choeropotamoidea

Fm. †Choeropotamidae Owen 1845

G. †Amphirhagatherium Deperet 1908

G. †Anthracobunodon Heller 1934

G. †Choeropotamus Desmarest 1822

G. †Cuisitherium Sudre et al. 1983

G. †Diplopus Kowalewsky 1873

G. †Hallebune Erfurt and Sudre 1995

G. †Haplobunodon Deperet 1908

G. †Lophiobunodon Deperet 1908

G. †Masilabune Tobien 1980

G. †Rhagatherium Pictet 1857

G. †Thaumastognathus Trouessart 1898

Fm. †Climacoceratidae Hamilton 1978

G. †Climacoceras MacInnes 1936

†Climacoceras africanus MacInnes 1936

†Climacoceras gentryi Hamilton 1978

G. †Prolibytherium Arambourg 1961

†Prolibytherium fusus Danowitz et al. 2016

†Prolibytherium magnieri Arambourg 1961

Invalid names: Climacoceratinae Hamilton 1978 [empty], Sperrgebietomerycinae Morales et al. 1999 [empty]

Fm. †Diacodexeidae Krishtalka and Stucky 1985

G. †Bunophorus Sinclair 1914

†Bunophorus etsagicus Cope 1882

†Bunophorus grangeri Sinclair 1914

Invalid names: Wasatchia dorseyana Sinclair 1914 [synonym], Wasatchia lysitensis Sinclair 1914 [synonym]

†Bunophorus macropternus Cope 1882

†Bunophorus pattersoni Krishtalka and Stucky 1986

†Bunophorus robustus Sinclair 1914

†Bunophorus sinclairi Guthrie 1966

Invalid names: Bunophorus gazini Guthrie 1971 [synonym]

G. †Diacodexis Cope 1882

†Diacodexis antunesi Estravís and Russell 1989

†Diacodexis corsaensis Checa 2004

†Diacodexis gazini Godinot 1978

†Diacodexis gigasei Smith 1996

†Diacodexis gracilis Krishtalka and Stucky 1985

†Diacodexis ilicis Gingerich 1989

†Diacodexis indicus Bajpai et al. 2005

†Diacodexis kelleyi Krishtalka and Stucky 1985

†Diacodexis metsiacus Cope 1882

†Diacodexis minutus Krishtalka and Stucky 1985

†Diacodexis morrisi Hooker 2010

†Diacodexis parvus Kumar et al. 2010

†Diacodexis primus Krishtalka and Stucky 1985

†Diacodexis secans Cope 1881

Invalid names: Diacodexis laticuneus Cope 1882 [synonym], Diacodexis olseni Sinclair 1914 [synonym], Mioclaenus brachystomus Cope 1882 [synonym]

†Diacodexis varleti Sudre 1983

Invalid names: Trigonolestes Cope 1894 [synonym], Wasatchia Sinclair 1914 [synonym]

G. †Gujaratia Bajpai et al. 2005

†Gujaratia pakistanensis Thewissen et al. 1983

G. †Jiangsudon Metais et al. 2008

†Jiangsudon shanghuangensis Metais et al. 2008

G. †Magwetherium Ducrocq et al. 2016

†Magwetherium burmense Ducrocq et al. 2016

G. †Neodiacodexis West and Atkins 1970

†Neodiacodexis emryi West and Atkins 1970

G. †Simpsonodus Krishtalka and Stucky 1986

†Simpsonodus chacensis Cope 1875

G. †Tapochoerus McKenna 1959

†Tapochoerus egressus Stock 1934

†Tapochoerus mcmillini Walsh 2000

Invalid names: Bunophorinae Stucky and Krishtalka 1990 [empty]

Superfm. †Dichobunoidea Gill 1872

G. †Acoessus Cope 1881

Fm. †Cebochoeridae Lydekker

G. †Acotherulum Gervais 1850

G. †Cebochoerus Gervais 1852

G. †Choeromorus Gervais 1848

G. †Gervachoerus Sudre 1978

G. †Moiachoerus Golpe-Posse 1972

Fm. †Dichobunidae Gill 1872

G. †Aksyiria Gabunia 1973

G. †Cadutherium Métais et al. 2007

G. †Chorlakkia Gingerich et al. 1979

Subfm. †Dichobuninae Turner 1849

G. †Dulcidon Van Valen 1965

Subfm. †Eurodexeinae Erfurt and Sudre 1996

G. †Haqueina Dehm and Oettingen-Spielberg 1958

Subfm. †Helohyinae Marsh 1877

Subfm. †Hyperdichobuninae Gentry and Hooker 1988

Subfm. †Lantianiinae Metais et al. 2004

G. †Mioachoerus

G. †Neochorlakkia Ducrocq et al. 2022

G. †Pakibune Thewissen et al. 1987

G. †Paraphenacodus Gabunia 1971

G. †Wutuhyus Tong and Wang 2006

Invalid names: Diacodexinae Gazin 1955 [empty]

Fm. †Helohyidae Marsh 1877

G. †Achaenodon Cope 1873

Invalid names: Protelotherium Osborn 1895 [synonym]

G. †Archaenodon Cope 1873

G. †Discritochoerus Gazin 1956

G. †Dyscritochoerus Gazin 1956

G. †Gobiohyus Matthew and Granger 1925

G. †Heliosus Burger and Jolley 2020

G. †Helohyus Marsh 1872

Invalid names: Lophiohyus Sinclair 1914 [synonym], Thinotherium Cope 1870 [synonym], Thinotherium Marsh 1872 [synonym]

G. †Ithygrammodon Osborn et al. 1878

G. †Lophiohys

G. †Pakkokuhyus Holroyd and Ciochon 1995

G. †Parahyus Marsh 1876

G. †Progenitohyus Ducrocq et al. 1997

Fm. †Homacodontidae Marsh 1849

G. †Antiacodon Marsh 1872

Invalid names: Nanomeryx Marsh 1894 [synonym]

G. †Asiohomacodon Tsubamoto et al. 2003

G. †Auxontodon Gazin 1958

G. †Bunomeryx Wortman 1898

G. †Gagadon Stucky and Covert 2014

G. †Hexacodus Gazin 1952

G. †Homacodon Marsh 1872

G. †Hylomeryx Peterson 1919

Invalid names: Sphenomeryx Peterson 1919 [synonym]

G. †Limeryx Metais et al. 2005

G. †Mesomeryx Peterson 1919

G. †Microsus Leidy 1870

G. †Mytonomeryx Gazin 1955

G. †Pentacemylus Peterson 1932

G. †Texodon West 1982

G. †Tsaganohyus Kondrashov et al. 2004

Invalid names: Bunomerycini Gentry and Hooker 1988 [empty], Homacodontini Marsh 1894 [empty]

Fm. †Leptochoeridae Marsh 1894

G. †Ibarus Storer 1984

G. †Laredochoerus Westgate 1994

G. †Leptochoerus Leidy 1856

Invalid names: Laopithecus Marsh 1875 [synonym]

G. †Stibarus Cope 1873

Invalid names: Menotherium Cope 1874 [synonym], Nanochoerus Macdonald 1955 [synonym]

Invalid names: Diacodexis woltonensis Krishtalka and Stucky 1985 [invalid subgroup]

Fm. †Mixtotheriidae Pearson 1927

G. †Mixtotherium Filhol 1880

Fm. †Raoellidae Sahni et al. 1981

G. †Indohyus Ranga Rao 1971

Invalid names: Kunmunella Sahni and Khare 1971 [synonym], Metkatius Kumar and Sahni 1985 [synonym], Raoella Sahni and Khare 1971 [synonym]

G. †Khirtharia Pilgrim 1940

Invalid names: Bunodentus Ranga Rao 1972 [synonym]

G. †Rajouria Rana et al. 2021

G. †Eolantianus Averianov 1996

†Eolantianus russelli Averianov 1996

G. †Erlianhyus Li and Li 2022

†Erlianhyus primitivus Li and Li 2022

G. †Hsanotherium Ducrocq et al. 2000

†Hsanotherium parvum Ducrocq et al. 2000

G. †Myanmarius Tsubamoto et al. 2013

†Myanmarius chitseini Tsubamoto et al. 2013

Subor. Omnivora

G. †Hadrohyus Leidy 1872

†Hadrohyus supremus Leidy 1872

G. †Raphenacodus Gabunia 1971

Unr. Ruminantiamorpha Spaulding et al. 2009

Fm. †Protoceratidae Marsh 1891 [protoceratid]

G. †Calops Marsh 1894

G. †Heteromeryx Matthew 1905

G. †Leptoreodon Wortman 1898

Invalid names: Camelomeryx Scott 1898 [synonym], Merycodesmus Scott 1898 [synonym]

G. †Leptotragulus Scott and Osborn 1887

Invalid names: Parameryx Marsh 1894 [synonym]

G. †Poabromylus Peterson 1932

Subfm. †Protoceratinae

G. †Pseudoprotoceras Cook 1934

Subfm. †Synthetoceratinae Frick 1937

G. †Toromeryx Wilson 1974

G. †Trigenicus Douglass 1903

Invalid names: Leptotragulinae [invalid subgroup]

Unr. Ruminantia Scopoli 1777 [ruminant]

Fm. †Agriochoeridae Leidy 1869 [agriochoerid]

Fm. †Andegamerycidae Ginsburg and Morales 1989

G. †Delahomeryx Stevens et al. 1969

G. †Krabitherium Métais et al. 2007

G. †Merycotherium Bojanus 1834

G. †Mosaicomeryx Mennecart and Métais 2015

G. †Paalitherium Métais et al. 2017

G. †Palaeohypsodontus Trofimov 1958

Unr. Pecora Linnaeus 1758

G. †Prodremotherium Filhol 1877

G. †Sanitherium von Meyer 1866

Invalid names: Diamantohyus Stromer 1926 [synonym], Xenochoerus Zdarsky 1909 [synonym]

G. †Thandaungia Métais 2006

Infraor. Tragulina Flower 1883

Invalid names: Anoplotheriadae Gray 1821 [empty], Giraffoidea Gray 1821 [empty], Hispanomerycidae Moya 1986 [empty]

Subor. Suiformes Jaeckel 1911

Invalid names: Anthracotherioidea Gill 1872 [empty], Palaeodonta Matthew [synonym]

Superfm. Suoidea Gray 1821

G. †Doliochoerus Filhol 1882

†Doliochoerus quercyi Filhol 1882

G. †Egatochoerus Ducrocq 1994

†Egatochoerus jaegeri Ducrocq 1994

G. †Odoichoerus Tong and Zhao 1986

†Odoichoerus uniconus Tong and Zhao 1986

G. †Palaeochoerus Pomel 1847

†Palaeochoerus gergovianus De Blainville 1846

†Palaeochoerus minus Gope Posse 1972

†Palaeochoerus platyops Cope 1881

†Palaeochoerus pusilus Ginsburg 1974

†Palaeochoerus typus Pomel 1847

G. †Schizochoerus Crusafont-Pairo and Lavocat 1954

†Schizochoerus anatoliensis van der Made 1997

†Schizochoerus arambourgi Ozansoy 1965

†Schizochoerus sinapensis van der Made 1997

†Schizochoerus vallesiensis Crusafont and Lavocat 1954

Fm. Suidae Gray 1821 [pig]

Subfm. †Cainochoerinae Pickford 1995

G. †Dinochoerus Peterson 1905

G. †Eocenchoerus Liu 2001

G. †Eumaiochoerus Hurzeler 1982

G. †Gerontochoerus Leakey 1943

G. †Hemichoerus Filhol 1882

G. †Hippopotamodon Lydekker 1877

Invalid names: Dicoryphochoerus Pilgrim 1926 [synonym], Limnostonyx Ginsburg 1988 [synonym]

G. †Hyosus Pilgrim 1926

Subfm. †Hyotheriinae Cope 1888

Subfm. †Kubanochoerinae Gabunia 1958

G. †Kubanochoerus Gabunia 1955

Subfm. †Listriodontinae Simpson 1945

G. †Lophochoerus Pilgrim 1926

G. †Mabokopithecus von Koenigswald 1969

G. †Mesochoerus Shaw and Cooke 1941

G. †Microstonyx Pilgrim 1926

Subfm. †Namachoerinae Pickford 1995

Subfm. †Palaeochoerinae Matthew 1924

G. †Paradoxodonides Strand 1943

G. †Promesochoerus Leakey 1965

G. †Pronotochoerus Leakey 1943

Subfm. †Schizochoerinae Golpe Posse 1972

Subfm. Suinae Gray 1821

Subfm. †Tetraconodontinae Lydekker 1886

Invalid names: Babyrousinae Thenius 1970 [empty], Chaeropotamina Bonaparte 1850 [empty], Phacochoeridae Gray 1868 [synonym]

G. †Taucanamo Simpson 1945

†Taucanamo grandaevum Fraas 1870

†Taucanamo inonuensis Pickford and Ertürk 1979

†Taucanamo primum van der Made 1997

†Taucanamo sansaniense Lartet 1851

Invalid names: Palaeochoeridae Matthew 1924 [invalid subgroup], Sanitheriidae Simpson 1945 [empty], Suina Gray 1868 [invalid subgroup], Suinamorpha Spaulding et al. 2009 [synonym]

Fm. †Tapirulidae Cope 1879

G. †Obotherium Bai et al. 2023

†Obotherium parvum Bai et al. 2023

†Obotherium tongi Bai et al. 2023

G. †Tapiruloides Bai et al. 2023

†Tapiruloides usuensis Bai et al. 2023

G. †Tapirulus Gervais 1850

†Tapirulus depereti Stehlin 1910

†Tapirulus hyracinus Gervais 1850

†Tapirulus majori Stehlin 1910

†Tapirulus perrierensis Sudre 1978

†Tapirulus schlosseri Stehlin 1910

G. †Tragulohyus Gervais 1874

Subor. Tylopoda Illiger 1811

Fm. †Amphimerycidae Stehlin 1910

G. †Amphimeryx Pictet 1853

G. †Pseudamphimeryx Stehlin 1919

Superfm. †Anoplotherioidea Bonaparte 1850

Fm. †Anoplotheriidae Gray 1821

Fm. †Xiphodontidae Flower 1883

Fm. †Oromerycidae Gazin 1955

G. †Camelodon Granger 1910

G. †Eotylopus Matthew 1910

G. †Malaquiferus Gazin 1955

G. †Merycobunodon Golz 1976

G. †Montanatylopus Prothero 1986

G. †Oromeryx Marsh 1894

G. †Protylopus Wortman 1898

Invalid names: Camelidamorpha Spaulding et al. 2009 [synonym], Cameloidea [empty], Oreodontoidea Gill 1872 [empty], Xiphodontoidea Flower 1883 [empty]

Invalid names: Achaenodontidae [empty], Ancodonta Matthew [empty], Antiacodontidae Krishtalka and Stucky 1985 [empty], Booidea [empty], Bunomerycidae Gentry and Hooker 1988 [empty], Cetartiodactyla Montgelard et al. 1997 [synonym], Eurytheriidae Cope 1889 [empty], Haplobunodontidae Sudre 1978 [empty], Hyodonta Matthew 1929 [empty], Leptochoerinae [empty], Merycoidodontoidea [empty], Neoselenodontia Webb and Taylor 1980 [empty], Nonruminantia Gregory 1910 [empty], Trigonolestidae [empty]

Unr. †Panperissodactyla Welker et al. 2015

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Or. †Panameriungulata de Muizon and Cifelli 2000

Fm. †Didolodontidae Simpson 1934

G. †Archaeohyracotherium Ameghino 1906

†Archaeohyracotherium mediale Ameghino 1902

G. †Argyrolambda Ameghino 1904

G. †Depaulacoutoia Cifelli and Ortiz-Jaureguizar 2014

Invalid names: Paulacoutoia Cifelli 1983 [replaced]

G. †Didolodus Ameghino 1897

†Didolodus magnus Gelfo 2010

†Didolodus minor Simpson 1948

†Didolodus multicuspis Ameghino 1897

Invalid names: Argyrolambda conidens Ameghino 1904 [synonym], Argyrolambda conulifera Ameghino 1904 [synonym], Didolodus colligatus Ameghino 1902 [synonym], Didolodus crassicuspis Ameghino 1901 [synonym], Didolodus dispar Ameghino 1904 [synonym], Lonchoconus lanceolatus Ameghino 1901 [synonym], Nephacodus latigonus Ameghino 1902 [synonym]

Invalid names: Cephanodus Ameghino 1902 [synonym], Lonchoconus Ameghino 1901 [synonym], Nephacodus Ameghino 1902 [synonym]

G. †Enneoconus Ameghino 1901

G. †Ernestokokenia Ameghino 1901

†Ernestokokenia chaishoer Simpson 1935

†Ernestokokenia nitida Ameghino 1901

Invalid names: Enneoconus parvidens Ameghino 1901 [synonym], Ernestokokenia marginata Ameghino 1901 [synonym]

†Ernestokokenia patagonica Ameghino 1901

Invalid names: Euprotogonia trigonalis Ameghino 1901 [synonym]

†Ernestokokenia yirunhor Simpson 1935

Invalid names: Notoprotogonia Ameghino 1904 [synonym]

G. †Escribania Bonaparte et al. 1993

†Escribania chubutensis Bonaparte et al. 1993

†Escribania talonicuspis Gelfo et al. 2007

G. †Lamegoia Paula Couto 1952

†Lamegoia conodonta Paula Couto 1952

G. †Paulogervaisia Ameghino 1901

†Paulogervaisia inusta Ameghino 1901

†Paulogervaisia mamma Ameghino 1901

†Paulogervaisia porca Ameghino 1901

G. †Proectocion Ameghino 1904

†Proectocion argentinus Ameghino 1904

†Proectocion precisus Ameghino 1904

Invalid names: Oxybunotherium Pascual 1965 [synonym]

G. †Raulvaccia Bonaparte et al. 1993

†Raulvaccia peligrensis Bonaparte et al. 1993

G. †Ricardocifellia Mones 2015

†Ricardocifellia protocenica Paula Couto 1952

G. †Salladolodus Soria and Hoffstetter 1983

†Salladolodus deuterotherioides Soria and Hoffstetter 1983

G. †Saltaodus Gelfo et al. 2020

†Saltaodus sirolli Gelfo et al. 2020

G. †Umayodus Gelfo and Sigé 2011

†Umayodus raimondi Gelfo and Sigé 2011

Invalid names: Bunolitopternidae Schlosser 1823 [synonym], Didolodidae Scott 1913 [replaced]

Subor. †Litopterna Ameghino 1889

Fm. †Adianthidae Ameghino 1891

Subfm. †Adianthinae

Subfm. †Indaleciinae Bond and Vucetich 1983

G. †Proheptoconus Bordas 1936

G. †Eolicaphrium Ameghino 1902

†Eolicaphrium primarium Ameghino 1902

Subor. †Lopholipterna Cifelli 1983

Invalid names: Macrauchenioidea Cifelli 1983 [empty], Proterotherioidea Ameghino 1887 [empty]

Fm. †Macraucheniidae Ameghino 1889

G. †Amilnedwardsia Ameghino 1901

Subfm. †Cramaucheniinae Ameghino 1902

G. †Ernestohaeckelia Ameghino 1901

G. †Llullataruca McGrath et al. 2018

G. †Macrauchenidia Cabrera 1939

Subfm. †Macraucheniinae Gervais 1855

G. †Macrauchenopsis Paula Couto 1945

G. †Notodiaphorus Loomis 1914

G. †Pseudomacrauchenia Kraglievich 1930

G. †Rutimeyeria Ameghino 1901

G. †Victorlemoineia Ameghino 1901

Invalid names: Polymorphinae Simpson 1945 [empty]

Fm. †Mesorhinidae Ameghino 1891

G. †Coelosoma Ameghino 1891

G. †Mesorhinus Ameghino 1885

G. †Pseudocoelosoma Ameghino 1891

Fm. †Notonychopidae Soria 1989

G. †Notonychops Soria 1989

G. †Requisia Bonaparte and Morales 1997

G. †Wainka Simpson 1935

Fm. †Proterotheriidae Ameghino 1887

G. †Anisolambda Ameghino 1901

Invalid names: Josepholeidya Ameghino 1901 [synonym]

Subfm. †Anisolambdinae Cifelli 1983

G. †Anisolophus Burmeister 1885

Tr. †Brachytherini Ameghino 1891

G. †Caliphrium

G. †Deuterotherium Ameghino 1895

G. †Diplasiotherium Rovereto 1914

G. †Eoauchenia Ameghino 1887

G. †Eoproterotherium Ameghino 1904

G. †Eoprotherotherium Ameghino 1904

G. †Epecuenia Cabrera 1939

G. †Guilielmofloweria Ameghino 1901

G. †Heteroglyphis Roth 1899

G. †Lambdaconus Ameghino 1897

G. †Licaphrops Ameghino 1904

G. †Lophogododon Ameghino 1904

Subfm. †Megadolodinae Cifelli and Villarroel 1997

G. †Mesolicaphrium McGrath et al. 2020

G. †Neodolodus Hoffstetter and Soria 1986

G. †Neolicaphrium Frenguelli 1921

G. †Olisanophus McGrath et al. 2020

G. †Paramacrauchenia Bordas 1939

G. †Phoradiadius Simpson et al. 1962

G. †Picturotherium Kramarz and Bond 2005

G. †Polyacrodon Roth 1899

Invalid names: Decaconus Ameghino 1901 [synonym], Oroacrodon Ameghino 1904 [objective synonym], Periacrodon Ameghino 1904 [synonym]

G. †Prolicaphrium Ameghino 1902

G. †Promylophis Shockey et al. 2022

Subfm. †Proterotheriinae Ameghino 1885

Tr. †Proterotherini Ameghino 1891

G. †Prothoatherium Ameghino 1902

G. †Ricardolydekkeria Ameghino 1901

Invalid names: Heterolambda Ameghino 1904 [synonym], Lopholambda Ameghino 1904 [synonym]

G. †Tetramerorhinus Ameghino 1894

G. †Thoatherium Ameghino 1887

Invalid names: Merycodon Mercerat 1891 [synonym], Rhagodon Mercerat 1891 [synonym]

G. †Villarroelia Cifelli and Guerrero Diaz 1997

G. †Xesmodon Berg 1899

Invalid names: Glyphodon Roth 1899 [replaced]

Invalid names: Anisolambda nodulosa Roth 1904 [nomen vanum], Bunodontheridae Mercerat 1891 [synonym]

Fm. †Protolipternidae Cifelli 1983

G. †Asmithwoodwardia Ameghino 1901

G. †Miguelsoria Cifelli 1983

G. †Protolipterna Cifelli 1983

Fm. †Sparnotheriodontidae Soria 1980

G. †Notiolofos Bond et al. 2009

G. †Notolophus Bond et al. 2006

G. †Sparnotheriodon Soria 1980

G. †Thoatheriopsis Soria 2001

†Thoatheriopsis mendocensis Soria 2001

Invalid names: Eolitopterna Soria 2001 [empty], Lambdaconus elegans Roth 1904 [nomen dubium], Paulogervaisia celata Ameghino 1901 [nomen dubium], Proacrodon Roth 1899 [nomen vanum], Proacrodon transformatus Roth 1899 [nomen vanum], Proadiantus gibbus Ameghino 1901 [nomen vanum]

Superor. Perissodactylamorpha Rose et al. 2020

Or. Perissodactyla Owen 1848 [odd-toed ungulate]

Fm. †Anthracobunidae Wells and Gingerich 1983

G. †Anthracobune Pilgrim 1940

G. †Ishatherium Sahni and Kumar 1980

G. †Jozaria Wells and Gingerich 1983

G. †Lammidhania Gingerich 1977

G. †Obergfellia Cooper et al. 2014

G. †Pilgrimella Dehm and Oettingen-Spielberg 1958

Fm. †Brontotheriidae Marsh 1873 [brontothere]

G. †Balochititanops Missiaen et al. 2011

Subfm. †Brontotheriinae Marsh 1873

G. †Danjiangia Wang 1995

Subfm. †Embolotheriinae Osborn 1929

Subfm. †Eotitanopinae Osborn 1913

G. †Lambdotherium Cope 1880

G. †Metatelmatherium Granger and Gregory 1938

G. †Mulkrajanops Kumar and Sahni 1985

G. †Pakotitanops West 1980

Subfm. †Palaeosyopinae Steinmann and Döderlein 1890

G. †Sivatitanops Pilgrim 1925

G. †Sthenodectes Gregory 1912

G. †Xylotitan Mihlbachler and Samuels 2016

Invalid names: Brontopinae Osborn 1914 [invalid subgroup], Epimanteoceratinae Granger and Gregory 1943 [empty], Menodontidae Cope 1881 [invalid subgroup], Titanotheriidae [synonym]

Fm. †Cambaytheriidae Bajpai et al. 2005

G. †Kalitherium Bajpai et al. 2006

G. †Cambaytherium Bajpai et al. 2005

†Cambaytherium gracilis Smith et al. 2016

†Cambaytherium marinus Bajpai et al. 2006

†Cambaytherium thewissi Bajpai et al. 2005

Invalid names: Cambaytherium bidens Bajpai et al. 2005 [synonym], Cambaytherium minor Bajpai et al. 2005 [synonym], Indobune vastanensis Rose et al. 2006 [synonym]

Invalid names: Indobune Rose et al. 2006 [synonym]

Or. †Desmostylia Reinhart 1953 [desmostylian]

G. †Behemotops Domning et al. 1986

Or. †Desmostyloidea Abel 1933

G. †Jamilcotatus Aranda-Manteca and Barnes 1998

G. †Kronokotherium Pronina 1957

G. †Seuku Beatty and Cockburn 2015

Invalid names: Cornwalliidae Shikama 1966 [empty], Desmostyliformes Hay 1923 [synonym]

Superfm. Equoidea Hay 1902

G. †Cymbalophus Hooker 1980

Fm. Equidae Gray 1821 [horse]

Fm. †Pachynolophidae Pavlow 1888

Fm. †Palaeotheriidae Bonaparte 1850

G. †Ghazijhippus Missiaen and Gingerich 2014

†Ghazijhippus talibhasani Missiaen and Gingerich 2014

G. †Hallensia Franzen and Haubold 1986

†Hallensia louisi Hooker 1994

†Hallensia matthesi Franzen and Haubold 1986

Subor. †Lophodontomorpha Hooker and Dashzeveg 2004

Infraor. †Euperissodactyla Hooker and Dashzeveg 2004

G. †Orientolophus Ting 1993

G. †Mesolambdolophus Holbrook and Lapergola 2011

†Mesolambdolophus setoni Holbrook and Lapergola 2011

G. †Nakusia Ginsburg et al. 1999

†Nakusia shahrigensis Ginsburg et al. 1999

G. †Paschatherium Russell 1963

†Paschatherium dolloi Teilhard de Chardin 1927

†Paschatherium levei Hooker and Russell 2012

†Paschatherium marianae Estravís and Russell 1992

†Paschatherium plaziati Marandat 1989

†Paschatherium russelli Godinot 1978

†Paschatherium yvettae Gheerbrant et al. 1997

G. †Perissobune Missiaen and Gingerich 2014

†Perissobune intizarkhani Missiaen and Gingerich 2014

†Perissobune munirulhaqi Missiaen and Gingerich 2014

G. †Skopaiolophus Métais et al. 2006

†Skopaiolophus burmese Métais et al. 2006

Subor. †Solidungula Cuvier

Fm. †Anchitheridae Leidy 1869 [anchitheriine horse]

Unr. Tapiromorpha Haeckel 1873

Unr. †Ancylopoda Cope 1889

G. †Cardiolophus Gingerich 1991

Unr. †Ceratomorpha Wood 1937

G. †Homogalax Hay 1899

Fm. †Isectolophidae Peterson 1919

G. †Isectolophus Scott and Osborn 1887

Invalid names: Parisectolophus Peterson 1919 [synonym]

G. †Kalakotia Ranga Rao 1972

Invalid names: Aulaxolophus Ranga Rao 1972 [synonym]

Fm. †Lophialetidae Matthew and Granger 1925

G. †Systemodon Cope 1881

Invalid names: Breviodontidae [empty], Moropomorpha [synonym], Rhodopagidae [empty]

G. †Teilhardimys Kretzoi and Kretzoi 2000

†Teilhardimys brisswalteri Hooker and Russell 2012

†Teilhardimys musculus Kretzoi and Kretzoi 2000

Invalid names: Microhyus musculus Teilhard de Chardin 1927 [replaced]

†Teilhardimys reisi Antunes et al. 1987

Invalid names: Microhyus Teilhard de Chardin 1927 [replaced]

Subor. †Titanotheriomorpha

Superfm. †Brontotherioidea Marsh 1873

Invalid names: Chelodactyla Scott 1913 [empty], Equina Bonaparte 1850 [empty], Hippopotamus iravaticus Noetling 1897 [nomen dubium], Hyracotheriidae Cope 1881 [empty], Mesaxonia Marsh 1884 [empty], Rhinocerotida Owen 1845 [empty], Selenida McKenna and Bell 1997 [invalid subgroup], Stereopterna Ameghino 1889 [empty], Triplopodidae Cope 1881 [empty]

G. †Radinskya McKenna et al. 1989

†Radinskya yupingae McKenna et al. 1989

Invalid names: Anthracobunia Ginsburg et al. 1999 [empty]

Unr. Phenacodonta McKenna 1975

G. †Olbitherium Tong et al. 2004

†Olbitherium millenariusum Tong et al. 2004

Fm. †Phenacodontidae Cope 1881 [condylarth]

G. †Almogaver Crusafont-Pairo and Villalta 1954

G. †Copecion Gingerich 1989

†Copecion brachypternus Cope 1882

†Copecion davisi Gingerich 1989

G. †Eodesmatodon Zheng and Chi 1978

†Eodesmatodon spanios Zheng and Chi 1978

G. †Lophocion Wang and Tong 1997

†Lophocion asiaticus Wang and Tong 1997

†Lophocion grangeri Bai et al. 2019

Subfm. †Meniscotheriinae Cope 1882

G. †Ectocion Cope 1882

Invalid names: Gidleyina Simpson 1935 [synonym], Prosthecion Patterson and West 1973 [synonym]

G. †Meniscotherium Cope 1874

Invalid names: Hyracops Marsh 1892 [synonym]

G. †Phenacodus Cope 1873

†Phenacodus bisonensis Gazin 1956

†Phenacodus grangeri Simpson 1935

Invalid names: Tetraclaenodon transitus Dorr 1958 [synonym]

†Phenacodus intermedius Granger 1915

†Phenacodus magnus Thewissen 1990

†Phenacodus matthewi Simpson 1935

Invalid names: Phenacodus gidleyi Simpson 1935 [synonym]

†Phenacodus sulcatus Cope 1874

†Phenacodus trilobatus Cope 1881

Invalid names: Phenacodus robustus Simpson 1937 [synonym]

†Phenacodus vortmani Cope 1880

Invalid names: Phenacodus almiensis Gazin 1942 [synonym], Phenacodus apternus Cope 1882 [synonym], Phenacodus copei Granger 1915 [synonym]

†Phenacodus zuniensis Cope 1881

Invalid names: Eohyus Marsh 1894 [synonym], Opisthotomus Cope 1875 [synonym], Phenacodus hemiconus Cope 1882 [nomen dubium], Phenacodus primaevus Cope 1873 [nomen dubium], Trispondylus Cope 1884 [replaced]

G. †Tetraclaenodon Scott 1892

†Tetraclaenodon puercensis Cope 1881

Invalid names: Mioclaenus floverianus Cope 1888 [synonym], Phenacodus calceolatus Cope 1883 [synonym], Protogonia plicifera Cope 1882 [synonym], Protogonia subquadrata Cope 1881 [synonym], Protogonia zuniensis Cope 1888 [synonym], Tetraclaenodon symbolicus Gidley 1935 [synonym]

†Tetraclaenodon septentrionalis Thewissen 1990

Invalid names: Euprotogonia Cope 1893 [synonym], Protogonia Cope 1881 [synonym]

Invalid names: Eparctocyona McKenna 1975 [synonym], Ungulata Linnaeus 1766 [synonym]

Diagnosis

No diagnoses are available