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Culicomorpha

Insecta - Diptera

Synonymy list
YearName and author
1994Culicomorpha Evenhuis
2007Culicomorpha Blagoderov and Grimaldi p. 11
2007Culicomorpha Yeates et al.
2011Culicomorpha Wiegmann et al.

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—EcdysozoaAguinaldo et al. 1997
—Panarthropoda
phylumArthropodaLatreille 1829
—Deuteropoda
—Mandibulata
—PancrustaceaZrzavý and Štys 1997
—AltocrustaceaRegier et al. 2010
superclassAllotriocarida
—Athalassocarida
—LabiocaridaSchwentner et al. 2017
superclassHexapodaLatreille 1825
RankNameAuthor
—Cercofilata
classInsecta
—Dicondylia
—Paranotalia
subclassPterygota()
—NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
—Holometabola
—Aparaglossata
—Panorpida
superorderAntliophora
orderDiptera
infraorderCulicomorpha
infraorderCulicomorpha

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Infraor. Culicomorpha
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Superfm. Chironomoidea Macquart 1838
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G. †Anisinodus Lukashevich et al. 2010
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†Anisinodus crinitus Lukashevich et al. 2010
Fm. Ceratopogonidae Newman 1834 [biting midge]
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G. †Adelohelea Borkent 1995
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†Adelohelea burmitica Szadziewski and Poinar 2005
†Adelohelea glabra Borkent 1995
†Adelohelea magyarica Borkent 1997
G. †Alautunmyia Borkent 1996
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†Alautunmyia elongata Borkent 1996
G. †Archiculicoides Szadziewski 1996
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†Archiculicoides acraorum Borkent 2000
†Archiculicoides schleei Szadziewski 1996
†Archiculicoides unus Borkent 2000
Subfm. †Atriculicoidinae Szadziewski 1996
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G. †Atriculicoides Remm 1976
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†Atriculicoides cenomanensis Szadziewski and Schlüter 1992
†Atriculicoides ciliatus Borkent 2012
†Atriculicoides dasyheleis Szadziewski 1996
†Atriculicoides globosus Boesel 1937
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Invalid names: Lasiohelea cretea Boesel 1937 [synonym]
†Atriculicoides hispanicus Szadziewski and Arillo 2016
†Atriculicoides incompletus Szadziewski and Schlüter 1992
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Invalid names: Atriculicoides squamiciliatus Remm 1976 [synonym]
†Atriculicoides sanjusti Szadziewski and Arillo 2016
†Atriculicoides sibiricus Szadziewski 1996
†Atriculicoides swinhoei Cockerell 1919
†Atriculicoides szadziewskii Pérez-de la Fuente et al. 2011
†Atriculicoides taimyricus Szadziewski 1996
G. †Burmahelea Szadziewski and Sontag 2019
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†Burmahelea neli Szadziewski and Sontag 2019
Subfm. Ceratopogoninae Newman 1834
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Tr. Ceratopogonini Newman 1834
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G. Alluaudomyia Kieffer 1913
G. Atyphohelea Borkent 1998
G. Austrohelea Wirth and Grogan 1988
G. Baeodasymyia Clastrier and Raccurt 1979
G. Baeohelea Wirth and Blanton 1970
G. †Brachycretacea Szadziewski 1996
G. Brachypogon Kieffer 1899
G. Camptopterohelea Wirth and Hubert 1960
G. Ceratoculicoides Wirth and Ratanaworabhan 1971
G. †Ceratopalpomyia Szadziewski 1988
G. Ceratopogon Meigen 1803
G. †Eohelea Petrunkevitch 1957
G. Fanthamia de Meillon 1939
G. †Fossihelea Szadziewski 1988
G. †Gedanohelea Szadziewski 1988
G. †Mantohelea Szadziewski 1988
G. Meunierohelea Szadziewski 1988
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Invalid names: Chimaerohelea Debenham 1988 [synonym]
G. †Monogedania Szadziewski et al. 2022
G. Monohelea Kieffer 1917
G. Nannohelea Grogan and Wirth 1980
G. †Nelohelea Szadziewski and Sontag 2019
G. †Palaeobrachypogon Borkent 1995
G. †Peronehelea Borkent 1995
G. Schizohelea Kieffer 1917
G. Serromyia Meigen 1818
G. Stilobezzia Kieffer 1911
G. †Wirthohelea Szadziewski 1988
Invalid names: Stilobezziini Wirth 1952 [synonym]
Tr. Culicoidini Kieffer 1911
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G. Culicoides Latreille 1809
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Invalid names: Devalquia Choufani and Nel 2013 [synonym]
G. Washingtonhelea Wirth and Grogan 1988
Tr. Heteromyiini Wirth 1962
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G. Heteromyia Say 1825
G. Metahelea Edwards 1929
G. Neurohelea Kieffer 1925
G. Physohelea Grogan and Wirth 1979
Tr. Johannsenomyiini Crampton 1925
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G. Johannsenomyia Malloch 1915
G. Macropeza Meigen 1818
G. Mallochohelea Wirth 1962
Tr. Palpomyiini Enderlein 1936
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G. Bezzia Kieffer 1899
G. Palpomyia Meigen 1818
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Invalid names: Eopalpomyitis Hong et al. 2000 [synonym], Miopalpomyia Pierce 1966 [synonym], Neopalpomyia Pierce 1966 [synonym], Parapalpomyia Pierce 1966 [synonym]
G. Phaenobezzia Haeselbarth 1965
Invalid names: Ceratopogon escheri Giebel 1856 [nomen dubium], Culicoides carri Pierce 1966 [nomen dubium], Culicoides fossilis Pierce 1966 [nomen dubium], Culicoides laurae Pierce 1966 [nomen dubium], Culicoides mioceneus Pierce 1966 [nomen dubium], Johannsenomyia hotchkissae Pierce 1966 [nomen dubium], Neoculicoides Pierce 1966 [nomen dubium], Neoculicoides jeanneae Pierce 1966 [nomen dubium], Sphaeromiini Newman 1834 [empty]
Subfm. Forcipomyiinae Lenz 1934
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G. Dasyhelea Kieffer 1913
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Invalid names: Sebessia Remm 1979 [synonym]
Tr. Forcipomyiini Lenz 1934
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G. Atrichopogon Kieffer 1906
G. Forcipomyia Meigen 1818
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Invalid names: Amberaspinus Evenhuis 1994 [synonym]
G. †Gerontodacus Borkent 2019
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†Gerontodacus krzeminskii Choufani et al. 2015
†Gerontodacus punctus Borkent 2000
†Gerontodacus skalskii Szadziewski and Arillo 1998
†Gerontodacus succineus Szadziewski 1996
G. †Heleageron Borkent 1995
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†Heleageron arenatus Borkent 1995
†Heleageron grimaldii Borkent 1996
Subfm. †Lebanoculicoidinae Borkent 2000
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G. †Lebanoculicoides Szadziewski 1996
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†Lebanoculicoides bloudani Choufani et al. 2015
†Lebanoculicoides daheri Choufani et al. 2015
†Lebanoculicoides excantabris Pérez-de la Fuente et al. 2011
†Lebanoculicoides mesozoicus Szadziewski 1996
Subfm. Leptoconopinae Noe 1907
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G. †Archiaustroconops Szadziewski 1996
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†Archiaustroconops alavensis Szadziewski and Arillo 1998
†Archiaustroconops andersoni Szadziewski et al. 2015
†Archiaustroconops annae Choufani et al. 2015
†Archiaustroconops besti Borkent et al. 2013
†Archiaustroconops borkenti Pérez-de la Fuente et al. 2011
†Archiaustroconops ceratoformis Szadziewski 1996
†Archiaustroconops cretaceous Szadziewski 1996
†Archiaustroconops dominiakae Choufani et al. 2015
†Archiaustroconops gracilis Szadziewski and Poinar 2005
†Archiaustroconops hammanensis Choufani et al. 2015
†Archiaustroconops hamus Borkent 2000
†Archiaustroconops kotejai Szadziewski and Poinar 2005
G. Austroconops Wirth and Lee 1958
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†Austroconops asiaticus Szadziewski 2004
†Austroconops borkenti Szadziewski and Schlüter 1992
†Austroconops fossilis Szadziewski 1996
†Austroconops gladius Borkent 2000
†Austroconops gondwanicus Szadziewski 1996
†Austroconops megaspinus Borkent 2000
†Austroconops perrichoti Dominiak et al. 2018
†Austroconops sibiricus Szadziewski 1996
G. †Baskintoconops Pielowska-Ceranowska 2023
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†Baskintoconops maaloufi Pielowska-Ceranowska 2023
G. †Fossileptoconops Szadziewski 1996
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†Fossileptoconops lebanicus Szadziewski 1996
G. †Jordanoconops Szadziewski 2000
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†Jordanoconops weitschati Szadziewski 2000
G. Leptoconops Skuse 1889
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Subg. Leptoconops (Leptoconops) Skuse 1889
Subg. †Leptoconops (Palaeoconops) Borkent 2001
G. †Minyohelea Borkent 1995
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†Minyohelea bacula Borkent 2000
†Minyohelea casca Borkent 1997
†Minyohelea falcata Borkent 2000
†Minyohelea lebanica Szadziewski 1996
†Minyohelea minuta Szadziewski 1996
†Minyohelea nexuosa Pielowska-Ceranowska 2021
†Minyohelea pumilis Borkent 1995
†Minyohelea schleei Szadziewski 1996
†Minyohelea wirthi Szadziewski 1996
Invalid names: Lebanoconops Szadziewski 1996 [synonym]
Invalid names: Austroconopinae Borkent et al. 1987 [synonym]
G. †Protoculicoides Boesel 1937
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†Protoculicoides revelatus Borkent 2019
Invalid names: Ceratopogon crassipalpis Meunier 1912 [nomen dubium], Ceratopogon madagascariensis Meunier 1912 [nomen dubium], Ceratopogon tenuipes Meunier 1912 [nomen dubium], Ceratopogon viscatus Meunier 1912 [nomen dubium], Heleidae [synonym], Simulia pasithea von Heyden 1870 [nomen dubium], Simulium terribilis Förster 1891 [nomen dubium], Sinopogonitinae Hong 2002 [invalid subgroup]
Fm. Chironomidae Macquart 1838 [non biting midge]
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Subfm. †Aenneinae Ansorge 1999
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G. †Aenne Ansorge 1999
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†Aenne liasina Ansorge 1999
†Aenne triassica Krzemiński and Krzemińska 1999
G. †Cretaenne Azar et al. 2008
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†Cretaenne inexpectata Azar et al. 2008
†Cretaenne kobeyssii Azar et al. 2008
†Cretaenne rasnicyni Lukashevich and Przhiboro 2011
Subfm. Aphroteniinae Brundin 1966
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Tr. †Electroteniini Kalugina 1980
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G. †Electrotenia Kalugina 1980
Subfm. Buchonomyiinae Brundin and Sæther 1978
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G. Buchonomyia Fittkau 1955
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†Buchonomyia succinea Seredszus and Wichard 2003
G. †Dungeyella Jarzembowski et al. 2008
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†Dungeyella gavini Jarzembowski et al. 2008
G. †Furcobuchonomyia Baranov et al. 2017
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†Furcobuchonomyia pankowskii Giłka and Zakrzewska 2017
†Furcobuchonomyia saetheri Baranov et al. 2017
G. Calopsectra Kieffer 1909
Subfm. Chironominae Macquart 1838
+
Tr. †Asiatendipini Hong 2002
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G. †Asiatendipes Hong 2002
G. †Bythomyia Zhang 1989
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†Bythomyia oryctes Zhang 1989
Tr. Chironomini Macquart 1838
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G. Axarus Roback 1980
G. Chironomus Meigen 1803
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Invalid names: Palaeolycus Etheridge and Olliff 1890 [synonym], Tendipes Meigen 1908 [synonym]
G. Cryptochironomus Kieffer 1918
G. Endochironomus Kieffer 1918
G. Glyptotendipes Kieffer 1913
G. †Hintelmanniella Baranov and Haug 2021
G. Paratendipes Kieffer 1911
G. †Prolipiniella Doitteau and Nel 2007
Invalid names: Tendipedini [synonym]
G. Cladopelma Kieffer 1921
Tr. †Fushunitendipini Hong 2002
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G. †Beifangitendipes Hong 2002
G. †Clavicornius Hong 2002
G. †Furvitendipes Hong 2002
G. †Fushunitendipes Hong 1981
G. †Latitendipes Hong 2002
G. †Spinitendipes Hong 2002
G. †Tenuigastrulus Hong 2002
Invalid names: Longigastrula Hong 2002 [nomen nudum]
G. Phaenopsectra Kieffer 1921
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†Phaenopsectra meunieri Seredszus and Wichard 2007
Tr. Pseudochironomini Saether 1977
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G. †Eomicromimus Giłka et al. 2023
G. †Eoriethia Giłka et al. 2023
G. Megacentron Freeman 1961
G. †Mesoacentron Giłka et al. 2021
G. Microtendipes Kieffer 1915
G. †Palaeocentron Giłka et al. 2021
G. Riethia Kieffer 1917
Tr. Tanytarsini Zavrel 1917
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G. †Archistempellina Giłka and Zakrzewska 2013
G. †Corneliola Giłka and Zakrzewska 2013
G. †Eonandeva Giłka and Zakrzewska 2015
G. Micropsectra Kieffer 1909
G. Nandeva Wiedenbrug et al. 1998
Subtr. Tanytarsina Zavrel 1917
Subtr. Zavreliina Saether 1977
Invalid names: Frutexitendipini Hong 2002 [invalid subgroup], Hamicaudini Hong 2002 [invalid subgroup], Lacusitendipini Hong 2002 [invalid subgroup], Longicopulini Hong 2002 [invalid subgroup]
G. †Chironomopsis Handlirsch 1906
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†Chironomopsis arrogans Giebel 1856
†Chironomopsis extinctus Brodie 1845
G. †Coelochironoma Zhang et al. 1986
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†Coelochironoma xantha Zhang et al. 1986
G. Corynocera Zetterstedt 1837
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Corynocera ambigua Zetterstedt 1837
†Corynocera duffi Deevey 1955
Invalid names: Dryadotanytarsus Andersen 1943 [synonym]
Subfm. †Cretodiamesinae Kalugina 1976
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G. †Cretodiamesa Kalugina 1976
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†Cretodiamesa taimyrica Kalugina 1976
G. †Cricotopiella Meunier 1916
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†Cricotopiella rostrata Meunier 1916
Subfm. Diamesinae Kieffer 1922
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G. Diamesa Walts 1837
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†Diamesa extincta Melander 1949
Tr. Diamesini Kieffer 1922
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G. Lappodiamesa Serra-Tosio 1968
G. Pagastia Oliver 1959
Tr. †Eugenodiamesini Lukashevich and Przhiboro 2015
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G. †Eugenodiamesa Lukashevich and Przhiboro 2015
Invalid names: Heptagyiini Brundin 1966 [empty]
G. Dicrotendipes Kieffer 1913
G. Eurycnemus van der Wulp 1874
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†Eurycnemus appendiculatus Meunier 1904
†Eurycnemus hyalinus Meunier 1904
†Eurycnemus pilosellus Meunier 1904
†Eurycnemus stagnum Meunier 1904
†Eurycnemus tenellus Meunier 1904
†Eurycnemus vulgaris Meunier 1904
G. †Haematotanypus Azar et al. 2008
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†Haematotanypus libanicus Azar et al. 2008
G. †Huabeitendipes Hong 1995
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G. †Jentzschiella Meunier 1899
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†Jentzschiella jentzschi Evenhuis 1994
G. †Jurochlus Kalugina 1985
+
†Jurochlus adustus Lukashevich and Przhiboro 2012
†Jurochlus limbatus Lukashevich and Przhiboro 2012
†Jurochlus lineatus Lukashevich and Przhiboro 2012
†Jurochlus rigor Kalugina 1985
†Jurochlus sibiricus Kalugina 1985
†Jurochlus trivittatus Lukashevich and Przhiboro 2012
G. †Langtonius Lukashevich and Przhiboro 2018
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†Langtonius cynaricaudatus Lukashevich and Przhiboro 2018
G. †Liauningius Hong 1982
+
G. †Libanochlites Brundin 1976
+
†Libanochlites eocenicus Baranov et al. 2019
†Libanochlites neocomicus Brundin 1976
G. †Manlayamyia Kalugina 1980
+
†Manlayamyia litorina Kalugina 1980
G. †Mogsonomus Kalugina 1985
+
†Mogsonomus tophus Kalugina 1985
G. †Nomochirus Gil Collado 1926
+
†Nomochirus sampelayoi Gil Collado 1926
Subfm. Orthocladiinae Lenz 1921
+
G. †Acutiforcipia Ashe and O'Connor 2019
+
†Acutiforcipia crusnotabile Seredszus and Wichard 2009
†Acutiforcipia cuspilonga Seredszus and Wichard 2009
†Acutiforcipia fittkaui Seredszus and Wichard 2009
G. Antillocladius Saether 1981
G. Brillia Kieffer 1913
G. Bryophaenocladius Thienemann 1934
+
†Bryophaenocladius beuki Baranov et al. 2015
†Bryophaenocladius circumclusus Seredszus and Wichard 2007
G. Camptocladius van der Wulp 1874
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†Camptocladius flexuosus Meunier 1904
†Camptocladius sinuosus Meunier 1904
G. Chaetocladius Kieffer 1911
+
†Chaetocladius incurvus Seredszus and Wichard 2007
G. †Chasmatonotoides Doitteau and Nel 2007
+
†Chasmatonotoides deharvengi Doitteau and Nel 2007
†Chasmatonotoides longiantennata Doitteau and Nel 2007
G. Corynoneura Winnertz 1846
+
†Corynoneura eocenica Doitteau and Nel 2007
G. Cricotopus van der Wulp 1874
+
†Cricotopus abiegnus Meunier 1904
†Cricotopus alluvionis Meunier 1904
†Cricotopus ambiguus Meunier 1904
†Cricotopus amniculus Meunier 1904
†Cricotopus antiquus Meunier 1904
†Cricotopus coniferus Meunier 1904
†Cricotopus crassicornis Meunier 1904
†Cricotopus delicatus Meunier 1904
†Cricotopus dilapsus Meunier 1904
†Cricotopus exstinctus Meunier 1904
†Cricotopus insolitus Meunier 1904
†Cricotopus minutissimus Meunier 1904
†Cricotopus minutulus Meunier 1916
†Cricotopus minutus Meunier 1904
†Cricotopus nemorivagus Meunier 1904
†Cricotopus paganus Meunier 1904
†Cricotopus parvulus Meunier 1916
†Cricotopus permutabilis Meunier 1904
†Cricotopus pulchellus Meunier 1904
†Cricotopus pygmaeus Meunier 1904
†Cricotopus robustus Meunier 1904
†Cricotopus saltuosus Meunier 1904
†Cricotopus variabilis Meunier 1904
G. †Flexicrus Seredszus and Wichard 2009
+
†Flexicrus palaeobalticus Seredszus and Wichard 2009
G. Heterotrissocladius Spark 1923
+
†Heterotrissocladius naibuchi Baranov et al. 2015
†Heterotrissocladius paleolacustris Seredszus and Wichard 2007
G. Hydrobaenus Fries 1830
+
†Hydrobaenus torpidus Seredszus and Wichard 2007
G. Hydrosmittia Ferrington and Sæther 2011
G. Krenosmittia Thienemann and Kruger 1939
+
†Krenosmittia paleofontana Seredszus and Wichard 2007
G. †Lebanorthocladius Veltz et al. 2007
+
†Lebanorthocladius furcatus Veltz et al. 2007
G. Limnophyes Eaton 1875
G. Metriocnemus van der Wulp 1874
+
†Metriocnemus cretatus Boesel 1937
G. †Myanmaro Giłka et al. 2019
+
†Myanmaro primus Giłka et al. 2019
G. Orthocladius van der Wulp 1874
G. Parachaetocladius Wulker 1959
+
†Parachaetocladius balticus Seredszus and Wichard 2007
G. †Parachasmatonotus Doitteau and Nel 2007
+
†Parachasmatonotus oesiensis Doitteau and Nel 2007
G. Parametriocnemus Goetghebuer 1932
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†Parametriocnemus electri Seredszus and Wichard 2007
G. Paraphaenocladius Thienemann 1924
+
†Paraphaenocladius evenhuisi Ashe and O'Connor 2014
+
Invalid names: Paraphaenocladius pusillus Seredszus and Wichard 2007 [replaced]
†Paraphaenocladius nadezhdae Baranov et al. 2015
G. †Ploegia Doitteau and Nel 2007
+
†Ploegia occulata Doitteau and Nel 2007
G. Psectrocladius Kieffer 1906
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†Psectrocladius trigonatus Seredszus and Wichard 2007
G. †Pseudochasmatonotus Doitteau and Nel 2007
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†Pseudochasmatonotus furcatus Doitteau and Nel 2007
G. Pseudorthocladius Goetghebuer 1932
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†Pseudorthocladius perantiquus Seredszus and Wichard 2007
†Pseudorthocladius zherikhini Baranov and Perkovsky 2013
G. Pseudosmittia Goetghebuer 1932
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†Pseudosmittia kodrulae Baranov et al. 2015
G. Pterosis Sublette and Wirth 1980
+
†Pterosis extinctus Baranov 2025
G. Rheosmittia Brundin 1986
+
†Rheosmittia pertenuis Seredszus and Wichard 2007
†Rheosmittia seni Doitteau and Nel 2007
G. Smittia Holmgren 1869
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†Smittia sukachevae Baranov et al. 2015
†Smittia veta Boesel 1937
G. Spaniotoma Philippi 1865
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†Spaniotoma conservata Boesel 1937
G. †Spinorthocladius Doitteau and Nel 2007
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†Spinorthocladius enigmaticus Doitteau and Nel 2007
G. Tokunagaia Goetghebuer 1932
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†Tokunagaia azari Doitteau and Nel 2007
G. Tokyobrillia Kobayashi and Sasa 1991
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†Tokyobrillia succinea Seredszus and Wichard 2009
G. Zalutschia Lipina 1939
+
†Zalutschia electra Seredszus and Wichard 2009
Invalid names: Metriocnemini Goetghebuer 1940 [empty]
G. †Orusa Lin 1980
+
†Orusa barba Lin 1980
G. Pelopia Latreille 1802
Subfm. Podonominae Thienemann and Edwards 1837
+
G. †Burmochlus Giłka et al. 2019
+
†Burmochlus madmaxi Giłka et al. 2019
G. †Gilkania Lukashevich 2020
+
†Gilkania transbaikalica Lukashevich 2020
G. †Glushkovella Kalugina 1993
+
†Glushkovella pallida Kalugina 1993
G. Lasiodiamesa Kieffer 1924
+
†Lasiodiamesa eocenica Seredszus and Wichard 2009
†Lasiodiamesa sinefoliis Seredszus and Wichard 2007
G. †Luanpingites Zhang 1986
+
†Luanpingites flavus Zhang 1986
G. †Oryctochlus Kalugina 1985
+
†Oryctochlus affinis Kalugina 1985
†Oryctochlus brundini Lukashevich 2012
†Oryctochlus contiguus Zhang 1991
†Oryctochlus kaluginae Lukashevich 2012
†Oryctochlus longilobus Kalugina 1985
†Oryctochlus minor Kalugina 1985
†Oryctochlus minutus Kalugina 1985
†Oryctochlus mirificus Kalugina 1993
†Oryctochlus placidus Kalugina 1993
†Oryctochlus toarciensis Ansorge 1996
†Oryctochlus vulcanus Kalugina 1985
G. †Palaeoboreochlus Baranov and Andersen 2014
+
†Palaeoboreochlus inornatus Baranov and Andersen 2014
G. Paraboreochlus Thienemann 1939
+
†Paraboreochlus bisaetiger Seredszus and Wichard 2007
G. †Podonomius Kalugina 1985
+
†Podonomius blepharis Lukashevich and Przhiboro 2011
†Podonomius macromastix Lukashevich and Przhiboro 2011
†Podonomius minimus Kalugina 1985
†Podonomius robustus Lukashevich and Przhiboro 2011
†Podonomius rotundatus Kalugina 1985
†Podonomius simplex Kalugina 1985
†Podonomius splendidus Kalugina 1985
†Podonomius tugnuicus Kalugina 1985
†Podonomius tumidus Ansorge 1996
†Podonomius undulatus Kalugina 1985
Invalid names: Luanpingitinae Zhang 1986 [synonym]
G. Polypedilum Kieffer 1912
Subfm. Prodiamesinae Saether 1976
+
G. †Cretadiamesa Veltz et al. 2007
+
†Cretadiamesa arieli Veltz et al. 2007
G. †Libanodiamesa Veltz et al. 2007
+
†Libanodiamesa deploegi Veltz et al. 2007
†Libanodiamesa simpsoni Baranov et al. 2018
G. Monodiamesa Kieffer 1922
+
†Monodiamesa guglielmia Doitteau and Nel 2007
G. †Paicheleria Azar and Nel 2010
+
†Paicheleria magnifica Azar and Nel 2010
G. Prodiamesa Kieffer 1906
+
†Prodiamesa indeserta Seredszus and Wichard 2009
G. †Sendelia Meunier 1899
+
†Sendelia mirabilis Evenhuis 1994
G. Sergentia Kieffer 1922
G. †Sinoryctochlus Zhang 1991
+
G. Stictochironomus Kieffer 1919
Subfm. Tanypodinae Skuse 1889
+
Tr. Anatopyniini Fittkau 1962
+
G. †Wadelius Veltz et al. 2007
Tr. Clinotanypodini Lipina 1928
+
G. Clinotanypus Kieffer 1913
G. Coelotanypus Kieffer 1913
Invalid names: Coelotanypodini Fittkau 1962 [synonym]
G. †Electroneura Amaral et al. 2024
+
†Electroneura pinhoi Amaral et al. 2024
G. †Gurvanomyia Kalugina 1986
+
†Gurvanomyia magna Kalugina 1986
†Gurvanomyia moderata Kalugina 1986
G. †Juropelopia Evenhuis 1994
+
†Juropelopia fittkaui Kalugina 1985
Invalid names: Mesopelopia Kalugina 1985 [replaced]
G. †Libanopelopia Veltz et al. 2007
+
†Libanopelopia cretacica Veltz et al. 2007
Tr. Macropelopiini Zavrel 1929
+
G. Apsectrotanypus Fittkau 1962
G. Brundiniella Roback 1978
G. Macropelopia Thienemann 1916
G. Psectrotanypus Kieffer 1909
G. †Mailonia Kalugina 1985
+
†Mailonia pallida Kalugina 1985
Tr. Pentaneurini Hennig 1950
+
G. Ablabesmyia Johannsen 1905
G. Conchapelopia Fittkau 1957
G. †Cretapelopia Veltz et al. 2007
G. Nilotanypus Kieffer 1923
Tr. Procladiini Roback 1971
+
G. Djalmabatista Fittkau 1968
G. †Eoprocladius Szadziewski et al. 2018
G. Procladius Skuse 1889
G. †Shinlustia Kalugina 1986
+
†Shinlustia irae Kalugina 1986
Tr. Tanypodini Skuse 1889
+
G. Tanypus Meigen 1803
G. †Tanypodites Kalugina 1985
+
†Tanypodites gorchonensis Kalugina 1985
G. †Tophocladius Kalugina 1985
+
†Tophocladius stygialis Kalugina 1985
G. †Ziadeus Azar and Nel 2010
+
†Ziadeus kamili Azar and Nel 2010
Invalid names: Pelopiinae [synonym]
G. †Tinactum Lin 1980
+
†Tinactum solusum Lin 1980
Subfm. †Ulaiinae Kalugina 1993
+
G. †Ulaia Kalugina 1985
+
†Ulaia communis Kalugina 1993
†Ulaia kangilica Kalugina 1993
†Ulaia magna Kalugina 1985
†Ulaia montana Kalugina 1985
†Ulaia reducta Kalugina 1985
G. †Ulaimailia Kalugina 1985
+
†Ulaimailia vetula Kalugina 1985
G. †Ulaimailonia Kalugina 1985
+
†Ulaimailonia mixta Kalugina 1985
G. †Ulaimailoniella Kalugina 1985
+
†Ulaimailoniella fusiformis Kalugina 1985
G. †Viduata Lin 1980
+
†Viduata otiosa Lin 1980
Invalid names: Protobibionidae Rohdendorf 1946 [synonym], Tendipedidae [synonym]
Superfm. Culicoidea Meigen 1818
hide
Fm. Chaoboridae Edwards 1920 [phantom midge]
+
G. †Chachotosha Lukashevich 1996
+
†Chachotosha perplexus Lukashevich 1996
†Chachotosha probatus Lukashevich 1996
Subfm. Chaoborinae Edwards 1920
+
G. Chaoborus Lichtenstein 1800
+
†Chaoborus abundans Baranov et al. 2024
†Chaoborus ciliatus Meunier 1904
†Chaoborus kishenehnicus Baranov and Haug 2021
†Chaoborus rarus Seredszus and Wichard 2009
†Chaoborus tertiarius von Heyden 1862
Invalid names: Corethra Meigen 1803 [synonym], Culicites von Heyden 1862 [synonym]
G. Mochlonyx Loew 1844
+
†Mochlonyx secundus Seredszus and Wichard 2009
†Mochlonyx sepultus Loew 1844
Invalid names: Mochlonyx atavus Loew 1861 [nomen nudum]
G. †Chaoborites Kalugina 1985
+
†Chaoborites argillaceus Kalugina 1985
G. †Chaoburmus Lukashevich 2000
+
†Chaoburmus amphilogos Lukashevich 2011
†Chaoburmus breviusculus Lukashevich 2000
†Chaoburmus victimaartis Lukashevich 2000
G. †Chironomaptera Ping 1928
+
†Chironomaptera bucucunica Kalugina 1985
†Chironomaptera collessi Jell and Duncan 1986
†Chironomaptera gobiensis Cockerell 1924
†Chironomaptera gregaria Grabau 1923
+
Invalid names: Chironomaptera melanura Ping 1928 [synonym]
†Chironomaptera karabonica Kalugina 1993
†Chironomaptera scobloi Kalugina 1985
†Chironomaptera vesca Kalugina 1980
+
Invalid names: Petiolatendipes shouchangensis Lin 1980 [synonym]
Invalid names: Mesotendipes Hong 1982 [objective synonym], Petiolatendipes Lin 1980 [synonym]
Subfm. †Chironomapterinae Kalugina 1974
+
G. †Astrocorethra Kalugina 1986
+
†Astrocorethra gurvanica Kalugina 1986
†Astrocorethra khukhmortica Kalugina 1986
†Astrocorethra mjangadica Kalugina 1986
G. †Baleiomyia Kalugina 1993
+
†Baleiomyia discussoria Kalugina 1993
G. †Helokrenia Kalugina 1985
+
†Helokrenia nana Kalugina 1985
G. †Hypsocorethra Kalugina 1985
+
†Hypsocorethra jurassica Kalugina 1985
†Hypsocorethra toficola Kalugina 1985
G. †Mesocorethra Kalugina 1993
+
†Mesocorethra cincta Lukashevich 2011
†Mesocorethra levis Kalugina 1993
G. †Praechaoborus Kalugina 1985
+
†Praechaoborus tugnuicus Kalugina 1985
†Praechaoborus udaensis Kalugina 1985
Invalid names: Mesotendipedinae Hong 1982 [objective synonym]
G. †Dixamima Rohdendorf 1951
+
†Dixamima villosa Rohdendorf 1951
Subfm. Eucorethrinae Edwards 1932
+
G. †Libanoculex Azar et al. 2023
+
†Libanoculex intermedius Azar et al. 2023
Invalid names: Libanoculicinae Azar et al. 2023 [synonym]
G. †Gedanoborus Szadziewski and Giłka 2007
+
†Gedanoborus kerneggeri Szadziewski and Giłka 2007
†Gedanoborus resinatus Seredszus and Wichard 2009
G. †Jordanobotomus Kaddumi 2005
+
†Jordanobotomus faigi Kaddumi 2005
G. †Libanoborus Azar et al. 2009
+
†Libanoborus lukashevichae Azar et al. 2009
G. †Mesochaoborus Zhang et al. 1986
+
†Mesochaoborus pallens Zhang et al. 1986
G. †Palaeomochlonyx Wichard et al. 2009
+
†Palaeomochlonyx aestimabilis Seredszus and Wichard 2009
G. †Rhaetomyia Rohdendorf 1962
+
†Rhaetomyia herrigi Ansorge 1996
†Rhaetomyia necopinata Rohdendorf 1962
G. †Taimyborus Lukashevich 1999
+
†Taimyborus aequiarticulatus Lukashevich 1999
G. †Triassomyia Lukashevich 2022
+
†Triassomyia shcherbakovi Lukashevich 2022
Invalid names: Corethra exita Scudder 1877 [nomen dubium], Dixamimidae Rohdendorf 1962 [synonym], Rhaetomyiidae Rohdendorf 1962 [synonym]
Fm. Corethrellidae Newman 1834 [frog biting midge]
+
G. Corethrella Coquillett 1902
+
Subg. Corethrella (Corethrella) Coquillett 1902
+
†Corethrella (Corethrella) miocaenica Szadziewski et al. 1994
†Corethrella (Corethrella) nudistyla Borkent and Szadziewski 1992
†Corethrella (Corethrella) prisca Borkent and Szadziewski 1992
†Corethrella (Corethrella) rovnoensis Baranov and Kvifte 2016
†Corethrella (Corethrella) sontagae Baranov and Kvifte 2016
Subg. †Corethrella (Fossicorethrella) Szadziewski 1995
+
†Corethrella andersoni Poinar and Szadziewski 2007
†Corethrella patula Baranov and Kvifte 2019
Fm. Culicidae Meigen 1818 [mosquito]
+
Subfm. Anophelinae Bassi 1900
+
G. Anopheles Meigen 1818
+
Subg. Anopheles (Nyssorhynchus) Blanchard 1902
†Anopheles rottensis Statz 1944
G. †Priscoculex Poinar et al. 2019
+
†Priscoculex burmanicus Poinar et al. 2019
Subfm. †Burmaculicinae Borkent and Grimaldi 2016
+
G. †Burmaculex Borkent and Grimaldi 2004
+
†Burmaculex antiquus Borkent and Grimaldi 2004
Subfm. Culicinae Meigen 1818
+
Tr. Aedini Neveu-Lemaire 1902
+
G. Aedes Meigen 1818
G. †Aetheapnomyia Harbach and Greenwalt 2012
G. †Eoaedes Harbach and Greenwalt 2012
G. Ochlerotatus Lynch Arribalzaga 1891
+
Invalid names: Taeniorhynchus Lynch Arribalzaga 1891 [synonym]
Tr. Culicini Meigen 1818
+
G. Culex Linnaeus 1758
Tr. Culisetini Belkin 1962
+
G. Culiseta Felt 1904
Tr. Mansoniini Belkin 1962
+
G. Coquillettidia Dyar 1905
G. Mansonia Blanchard 1901
Tr. Toxorhynchitini Theobald 1905
+
G. Toxorhynchites Theobald 1901 [elephant mosquito]
G. †Neoculicites Evenhuis 1994
+
†Neoculicites ceyx von Heyden 1870
†Neoculicites depereti Meunier 1915
Invalid names: Culicites Meunier 1915 [replaced]
G. †Paleoculicis Poinar et al. 2000
+
†Paleoculicis minutus Poinar et al. 2000
Fm. Dixidae Van der Wulp 1877 [meniscus midge]
+
G. Dixa Meigen 1818
+
†Dixa minuta Meunier 1906
†Dixa tertiaria Meunier 1915
+
Invalid names: Dixa hyalipennis Theobald 1937 [synonym]
Invalid names: Eriopterites Meunier 1915 [synonym]
G. Dixella Dyar and Shannon 1924
+
†Dixella curvistyla Greenwalt and Moulton 2016
†Dixella distans Hennig 1966
†Dixella eomarginata Greenwalt and Moulton 2016
†Dixella filiforceps Hennig 1966
†Dixella intacta Greenwalt and Moulton 2016
Dixella lirio Dyar and Shannon 1924
†Dixella priscula Cockerell 1921
†Dixella spinilobata Greenwalt and Moulton 2016
†Dixella succinea Meunier 1906
Invalid names: Paradixa Tonnoir 1924 [synonym]
G. †Eucorethrina Kalugina 1985
+
†Eucorethrina convexa Lukashevich 1996
†Eucorethrina flexa Kalugina 1985
†Eucorethrina westwoodi Lukashevich et al. 2001
G. †Syndixa Lukashevich 1996
+
†Syndixa liasina Lukashevich 1996
†Syndixa mollis Lukashevich 1996
†Syndixa sibirica Lukashevich 1996
Invalid names: Dixa cimbrica Ansorge 1992 [nomen dubium]
Superfm. Simulioidea Newman 1834
hide
Fm. †Kaluginamyiidae Lukashevich et al. 2019
+
G. †Kaluginamyia Lukashevich et al. 2019
+
†Kaluginamyia baissica Lukashevich et al. 2019
†Kaluginamyia brevirostris Kalugina 1986
†Kaluginamyia enigmatica Lukashevich et al. 2019
Fm. Simuliidae Newman 1834 [black fly]
+
Subfm. Gymnopaidinae Rubzov 1956
+
G. †Baisomyia Kalugina 1991
+
†Baisomyia incognita Kalugina 1991
G. †Gydarina Kalugina 1991
+
†Gydarina karabonica Kalugina 1991
G. Inseliellum Rubtsov 1974
Subfm. †Kovalevimyiinae Kalugina 1991
+
G. †Kovalevimyia Kalugina 1991
+
†Kovalevimyia lacrimosa Kalugina 1991
G. Nevermannia Enderlein 1921
Subfm. Simuliinae Newman 1834
+
G. Ectemnia Enderlein 1930
+
†Ectemnia cerberus Enderlein 1921
Tr. Nevermanniini Enderlein 1921
+
G. Hellichiella Rivosecchi and Cardinale 1975
Tr. Simuliini Newman 1834
+
G. †Archicnephia Currie and Grimaldi 2000
G. †Ugolyakia Perkovsky and Sukhomlin 2018
Invalid names: Ectemnia lithuanica Yankovsky and Bernotiene 2005 [nomen dubium]
Tr. Stegopterniini Enderlein 1930
+
G. Greniera Doby and David 1959
G. †Simuliites Kalugina 1986
+
†Simuliites yantardakh Perkovsky and Sukhomlin 2016
G. †Simulimima Kalugina 1985
+
†Simulimima grandis Kalugina 1985
G. Simulium Latreille 1802
+
Subg. Simulium (Inseliellum) Rubtsov 1974
+
†Simulium (Inseliellum) raivavaense Craig and Porch 2013
Simulium (Inseliellum) teruamanga Craig and Craig 1986
Fm. Thaumaleidae Bezzi 1913
+
G. †Mesothaumalea Kovalev 1989
+
†Mesothaumalea fossilis Kovalev 1989
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Bush and Bambach 2015, Kiessling 2004

Age range

Maximum range based only on fossils: base of the Aegean to the top of the Holocene or 246.70000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 244.99 Ma

Collections (399 total)

Oldest occurrences

Time interval Ma Country or state Original ID and collection number
Aegean246.7 - 244.99France Anisinodus crinitus (106916)
Ladinian241.464 - 237.0Kyrgyzstan (Osh) Triassomyia shcherbakovi (228748)
Sevatian211.18 - 205.7Germany Chironomidae indet. (216439)
Rhaetian205.7 - 201.4United Kingdom (England) Aenne triassica (115590)
Hettangian - Sinemurian201.4 - 192.9Kyrgyzstan (Tonskiy) Rhaetomyia necopinata (105086) Syndixa liasina (105084)
Early/Lower Toarcian184.2 - 182.9Germany (Mecklenburg-Vorpommern) Rhaetomyia herrigi, Podonominae indet., Oryctochlus toarciensis, Podonomius tumidus, Aenne liasina (114993)
Falciferum184.2 - 182.9Germany (Mecklenburg-Vorpommern) Aenne liasina (151350)
Toarcian184.2 - 174.7Russian Federation (Krasnoyarsk) Podonomius minimus (110819)
Toarcian - Aalenian184.2 - 170.9Russian Federation (Kemerovo) Chironomaptera bucucunica (125475)
Bajocian - Bathonian170.9 - 165.3Russian Federation (Krasnoyarsk) Eucorethrina flexa, Syndixa mollis, Syndixa sibirica, Helokrenia nana, Chaoborites argillaceus, Podonomius simplex, Podonomius undulatus, Podonomius rotundatus, Podonomius minimus (118010)
Callovian - Oxfordian165.3 - 154.8Kazakhstan (Chimkent) Dixamima villosa (122331) Eucorethrina sp. (105426) Protobibio jurassicus (115717)
Callovian - Oxfordian165.3 - 154.8China (Hebei) Luanpingites flavus (156314)
Oxfordian161.5 - 154.8Russian Federation Praechaoborus tugnuicus (123219)
Oxfordian161.5 - 154.8Russian Federation (Chita) Chaoboridae indet., Chironomidae indet. (125499)
Oxfordian161.5 - 154.8Russian Federation (Buryatia) Hypsocorethra jurassica (123195) Hypsocorethra sp. (123147) Podonominae indet. (143413) Simulimima grandis (124595)
Oxfordian161.5 - 154.8Russian Federation (Buryat) Praechaoborus tugnuicus, Podonomius tugnuicus, Podonomius splendidus, Mesopelopia fittkaui (105017)
Oxfordian - Kimmeridgian161.5 - 149.2Russian Federation (Chita) Chironomaptera scobloi, Mogsonomus tophus (114661)
Oxfordian - Kimmeridgian161.5 - 149.2Russian Federation (Buryat) Hypsocorethra toficola, Oryctochlus minutus, Oryctochlus affinis, Tanypodites gorchonensis (114614) Praechaoborus udaensis, Hypsocorethra toficola, Ulaimailia vetula, Ulaimailoniella fusiformis, Oryctochlus vulcanus, Oryctochlus minor, Oryctochlus longilobus, Oryctochlus affinis, Ulaimailonia mixta, Jurochlus rigor, Jurochlus sibiricus, Tophocladius stygialis, Mailonia pallida, Ulaia magna, Ulaia montana, Ulaia reducta (104950)
Late/Upper Jurassic161.5 - 143.1Russian Federation Chaoboridae indet., Chironomaptera bucucunica (114317)
Late/Upper Jurassic161.5 - 143.1Russian Federation (Chita) Chaoboridae indet. (143412)