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Aparaglossata
Taxonomy
Aparaglossata was named by Peters et al. (2014). It is extant.
It was assigned to Holometabola by Peters et al. (2014), Misof et al. (2014), Engel (2022).
It was assigned to Holometabola by Peters et al. (2014), Misof et al. (2014), Engel (2022).
Subtaxa
Synonymy list
| Year | Name and author |
|---|---|
| 2014 | Aparaglossata Misof et al. |
| 2014 | Aparaglossata Peters et al. |
| 2022 | Aparaglossata Engel |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Unr. Aparaglossata Peters et al. 2014
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Unr. Neuropteriformia Engel 2022
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Unr. Coleopterida Kukalová-Peck and Lawrence 2004
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Or. Coleoptera Linnaeus 1758 [beetle]
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G. †Allognosis Handlirsch 1906
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†Allognosis nitens Geinitz 1894
†Allognosis striatus Ponomarenko 1985
Subor. †Alphacoleoptera Engel et al. 2022
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Superfm. †Coleopsoidea Kirejtshuk and Nel 2016
Superfm. †Permocupedoidea Martynov 1933
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Invalid names: Tshekardocoleoidea Rodendorf 1944 [synonym]
Superfm. †Permosynoidea Tillyard 1924
G. †Aptilotitus Carpenter 1986
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†Aptilotitus capitecarens Bode 1953
Invalid names: Aptilotus Bode 1953 [replaced]
G. †Auchenophorites Carpenter 1986
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†Auchenophorites sculpturatus Bode 1953
Invalid names: Auchenophorus Bode 1953 [replaced]
G. †Cardioides Bode 1953
G. †Cardiosyne Martins-Neto and Gallego 2006
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†Cardiosyne elegans Martins-Neto and Gallego 2006
†Cardiosyne obesa Martins-Neto and Gallego 2006
G. †Chimaerocoleites Ponomarenko 1985
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†Chimaerocoleites gyrinoides Ponomarenko 1985
†Chimaerocoleites parvulus Ponomarenko 1985
†Chimaerocoleites punctatus Ponomarenko 1985
†Chimaerocoleites seriatus Ponomarenko 1985
G. †Cistelites Heer 1864
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†Cistelites insignis Heer 1864
†Cistelites longipes Hong 1985
†Cistelites minor Heer 1874
†Cistelites punctulatus Heer 1869
†Cistelites sachalinensis Heer 1878
†Cistelites spectabilis Heer 1869
G. †Clinomerus Carpenter 1986
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†Clinomerus laticollis Handlirsch 1939
Invalid names: Catomerus Handlirsch 1939 [replaced]
G. †Coleopteron Handlirsch 1906
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†Coleopteron antiquus Weyenbergh 1869
†Coleopteron aquaticus Brodie 1845
†Coleopteron beyrichi Giebel 1856
†Coleopteron bractoides Blake 1876
†Coleopteron lentissimus Weyenbergh 1874
†Coleopteron rugosostriatus Giebel 1856
†Coleopteron semicrematus Jarzembowski 2003
†Coleopteron sibiricus Heer 1876
†Coleopteron striatus Oppenheim 1888
†Coleopteron studeri Giebel 1856
†Coleopteron vetustus Giebel 1856
†Coleopteron wittsi Brodie 1845
†Coleopteron zekelii Giebel 1856
Invalid names: Coleopteron harpalinus Heer 1864 [nomen nudum], Coleopteron spectabilis Heer 1864 [nomen nudum], Coleopteron troglodytes Heer 1864 [nomen nudum]
G. †Cretohelophorus Ponomarenko 1987
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†Cretohelophorus cupedoides Ponomarenko 1987
†Cretohelophorus yanensis Ren 1995
G. †Critoderma Carpenter 1986
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†Critoderma deplanatum Heer 1864
Invalid names: Cycloderma Heer 1864 [replaced]
G. †Cryptauchenia Bode 1953
G. †Curculidium Handlirsch 1907
G. †Cyphospheron Bode 1953
G. †Darbiopelta Ponomarenko 1986
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†Darbiopelta crassa Ponomarenko 1986
†Darbiopelta dubia Ponomarenko 1986
G. †Dicyphelus Carpenter 1986
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†Dicyphelus concameratus Bode 1953
Invalid names: Dicyphus Bode 1953 [replaced]
G. †Doggeria Handlirsch 1906
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†Doggeria bucklandi Mantell 1844
†Doggeria murchisoni Handlirsch 1906
†Doggeria sibirica Handlirsch 1906
G. †Etheridgea Handlirsch 1906
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†Etheridgea australis Handlirsch 1906
†Etheridgea petrica Tillyard 1916
G. †Eurynotellus Carpenter 1986
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†Eurynotellus brevicollis Bode 1953
Invalid names: Eurynotus Bode 1953 [replaced]
G. †Flichea Handlirsch 1906
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†Flichea elongata Ponomarenko 1985
†Flichea lotharingiaca Fliche 1901
†Flichea sibirica Ponomarenko 1985
G. †Glaphyropterula Handlirsch 1906
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†Glaphyropterula gracilis Heer 1852
†Glaphyroptula anglica Cockerell 1915
†Glaphyroptula liasina Giebel 1856
G. †Heeriaopsis Carpenter 1986
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†Heeriaopsis laevigatum Heer 1878
Invalid names: Heeriella Handlirsch 1906 [replaced]
G. †Helopidium Westwood 1854
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†Helopidium brodiei Giebel 1856
†Helopidium dubium Handlirsch 1906
†Helopidium dunkeri Giebel 1856
†Helopidium neoridas Westwood 1854
†Helopidium rugosum Handlirsch 1906
†Helopidium westwoodi Giebel 1856
G. †Hydrophilites Heer 1864
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†Hydrophilites acherontis Heer 1852
†Hydrophilites naujatensis Heer 1883
†Hydrophilites stygius Heer 1864
G. †Keleusticus Handlirsch 1906
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†Keleusticus acutus Ponomarenko 1985
†Keleusticus duplicatus Ponomarenko 1985
†Keleusticus zirkeli Geinitz 1894
G. Kryptophagus Herbst 1792
G. †Leptilostis Ponting 2024
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†Leptilostis parvula Hong 1982
Invalid names: Tetillopsis Hong 1982 [replaced]
G. †Mesoncus Carpenter 1986
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†Mesoncus striatulus Bode 1953
Invalid names: Loxoncus Bode 1953 [replaced]
G. †Mesorhynchophora Tillyard 1916
G. †Microcoleopteron Handlirsch 1906
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†Microcoleopteron heydeni Weyenbergh 1869
†Microcoleopteron jurassicum Weyenbergh 1869
†Microcoleopteron lithographicum Weyenbergh 1869
†Microcoleopteron minimum Oppenheim 1888
Unr. Nubili Latreille 1804
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Invalid names: Herbicolae Latreille 1804 [empty], Sabulosi Latreille 1804 [empty]
G. †Ooidellus Carpenter 1986
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†Ooidellus denudatus Bode 1953
Invalid names: Ooides Bode 1953 [replaced]
Tr. †Oropseini Legalov and Kirejtshuk 2016
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G. †Libanorhinus Kuschel and Poinar 1993
G. †Oropsis Legalov and Kirejtshuk 2016
G. †Oxytoroptera Handlirsch 1939
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†Oxytoroptera mediocris Handlirsch 1939
†Oxytoroptera punctata Ponomarenko 1985
G. †Pantodapus Handlirsch 1906
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†Pantodapus ewaldi Giebel 1856
†Pantodapus knorri Giebel 1856
†Pantodapus westwoodi Handlirsch 1906
G. †Parabuprestium Handlirsch 1906
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†Parabuprestium pseudocarabus Handlirsch 1906
†Parabuprestium teleas Westwood 1854
G. †Pedinosoma Bode 1953
G. †Peridosoma Carpenter 1986
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†Peridosoma praecisum Bode 1953
Invalid names: Perosoma Bode 1953 [replaced]
G. †Pseudoelateropsis Handlirsch 1906
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†Pseudoelateropsis infraliassica Roemer 1876
Invalid names: Elateropsis Roemer 1876 [replaced]
G. †Pseudotelephorus Handlirsch 1906
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†Pseudotelephorus grandis Cockerell 1915
†Pseudotelephorus haueri Giebel 1856
†Pseudotelephorus punctulatus Cockerell 1915
G. †Pseudotenebrio Handlirsch 1906
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†Pseudotenebrio innominatus Weyenbergh 1869
†Pseudotenebrio relictus Weyenbergh 1869
G. †Semiglobus Handlirsch 1906
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†Semiglobus chrysomelinus Handlirsch 1906
†Semiglobus jurassicus Handlirsch 1906
†Semiglobus neptuni Giebel 1856
G. †Silphidium Handlirsch 1907
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†Silphidium priscum Fritsch 1882
Invalid names: Silphites Fritsch 1882 [replaced]
G. †Strongylites Heer 1864
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†Strongulites latus Ponomarenko 1985
†Strongylites morio Heer 1864
†Strongylites stygius Heer 1864
Fm. Tenebrionites Latreille 1802 [darkling beetle]
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Invalid names: Pimeliariae Latreille 1802 [empty], Tenebrionites Latreille 1802 [empty]
Fm. †Tshekardocoleidae Rohdendorf 1944
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G. †Avocoleus Ponomarenko 1969
G. †Boscoleus Kukalová 1968
G. †Eocoleus Kukalová 1968
G. †Moravocoleus Kukalová 1968
G. †Permocoleus Lubkin and Engel 2005
G. †Prosperocoleus Kukalová 1968
G. †Retelytron Kukalová 1965
G. †Sylvacoleodes Ponomarenko 1969
G. †Sylvacoleus Ponomarenko 1963
G. †Tshekardocoleus Rohdendorf 1944
G. †Umoricoleus Kukalová 1968
G. †Uralocoleus Zalessky 1947
G. †Votocoleus Kukalová 1968
Invalid names: Moravocoleidae Kukalová-Peck and Beutel 2012 [synonym], Uralocoleidae Zalessky 1947 [synonym], Uralocoleus ultimus Ponomarenko 2000 [nomen dubium]
Invalid names: Acmaeodera perreaui Piton 1940 [nomen dubium], Ancylocheira liasina Giebel 1856 [nomen dubium], Brochocoleus indibili Soriano and Delclos 2006 [nomen dubium], Carabiques Latreille 1802 [empty], Cebrionates Latreille 1802 [empty], Chrysomelinae Latreille 1802 [empty], Cicindeletae Latreille 1802 [empty], Cistela Geoffroy 1762 [nomen oblitum], Clerii Latreille 1802 [empty], Coleopteron depressus Heer 1879 [nomen nudum], Coleopteron interpunctatus Heer 1864 [nomen nudum], Coleopteron laevigatus Heer 1864 [nomen nudum], Coleopteron scabriusculus Heer 1879 [nomen nudum], Coprophagi Latreille 1802 [empty], Criocerina pristina Germar 1813 [nomen dubium], Curculionites liasinus Heer 1852 [nomen dubium], Curculiopsidae Martynov 1937 [empty], Cydnopsis nigromembranacea Jordan 1967 [nomen dubium], Diaperialae Latreille 1802 [empty], Elateridopsidae Zalessky 1929 [invalid subgroup], Eleuterata Fabricius 1775 [synonym], Entomophagi Latreille 1804 [empty], Eodromeus viriosus Zhang 1997 [nomen dubium], Erotilenae Latreille 1802 [empty], Euchroma arvernica Piton 1940 [nomen dubium], Eucoleoptera Engel et al. 2022 [invalid subgroup], Helophorus exilis Heer 1862 [nomen dubium], Helophorus magnus Heer 1862 [nomen dubium], Helopii Latreille 1802 [empty], Helops atticus Redtenbacher 1867 [nomen dubium], Hydrobius godeti Heer 1862 [nomen dubium], Hydrobius obsoletus Heer 1856 [nomen dubium], Hydrocanthares Latreille 1802 [empty], Hydrophilus stenopterus Heer 1862 [nomen dubium], Hydrous ebeninus Statz 1939 [nomen dubium], Hylesinus lineatus Förster 1891 [nomen dubium], Ipidae Bedel 1888 [empty], Isomira aurora Wickham 1914 [nomen dubium], Leptopodocoleidae Hong 1982 [invalid subgroup], Macrogastri Latreille 1802 [empty], Malacodermi Latreille 1802 [empty], Malacodermidae [empty], Megopis lineolatus Piton 1940 [nomen dubium], Metaxycoleoptera Engel et al. 2022 [invalid subgroup], Necrophagi Latreille 1802 [empty], Necticus minutus Aymard 1856 [nomen nudum], Nitidulariae Latreille 1802 [empty], Onthophagus luteus Oustalet 1874 [nomen dubium], Palpatores Latreille 1802 [empty], Platyprosopi Latreille 1804 [empty], Protocoleoptera Tillyard 1924 [empty], Rhynchophori Latreille 1804 [empty], Saperda valdensis Heer 1864 [nomen dubium], Saprophagi Latreille 1804 [empty], Staphyliniae Latreille 1802 [empty], Sternoxi Latreille 1802 [empty], Tridigitati Latreille 1802 [empty], Versicolores Latreille 1804 [empty], Xylophagi Latreille 1802 [empty], Zacoleoptera Engel et al. 2022 [invalid subgroup]
Or. †Skleroptera Kirejtshuk and Nel 2013
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Or. Strepsiptera Kirby 1813 [twisted-winged parasite]
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Fm. †Cretostylopidae Kathirithamby and Engel 2014
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G. †Cretostylops Grimaldi and Kathirithamby 2005
Fm. †Phthanoxenidae Engel and Huang 2016
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G. †Heterobathmilla Pohl and Beutel 2020
G. †Kinzelbachilla Pohl and Beutel 2016
G. †Phthanoxenos Engel and Huang 2016
Invalid names: Kinzelbachillidae Pohl and Beutel 2016 [synonym]
Invalid names: Mengenilliformia Kinzelbach 1971 [invalid subgroup], Mengenilloidea Hofeneder 1910 [invalid subgroup], Mengeoidea Pierce 1908 [invalid subgroup]
Superor. Neuropterida
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Unr. Metaxyneuroptera Engel 2022
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Fm. †Permithonidae Tillyard 1922
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G. †Eopsychops Martynov 1932
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†Eopsychops angaridensis Martynova 1961
†Eopsychops sojanensis Martynov 1932
G. †Kunguromaritus Vilesov 1995
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†Kunguromaritus guttatus Vilesov 1995
†Kunguromaritus lacer Vilesov 1995
G. †Okolpania Vilesov 1995
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†Okolpania captiosa Vilesov 1995
†Okolpania eugeniae Makarkin and Shcherbakov 2009
†Okolpania favorabilis Novokshonov and Novokshonova 1997
†Okolpania observabilis Vilesov 1995
G. †Palaemerobius Martynov 1928
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†Palaemerobius avus Martynova 1952
†Palaemerobius latibasis Martynova 1961
†Palaemerobius proavitus Martynov 1928
G. †Permantispa Ponomarenko and Shcherbakov 2004
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†Permantispa emelyanovi Ponomarenko and Shcherbakov 2004
G. †Permegalomus Martynov 1931
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†Permegalomus maculipennis Martynov 1932
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Invalid names: Permithonopsis triramosa Martynova 1952 [synonym]
†Permegalomus regularis Martynov 1931
†Permegalomus spectabilis Martynova 1952
Invalid names: Bianchia Martynova 1952 [synonym]
G. †Permipsythone Pinto and de Ornellas 1980
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†Permipsythone panfilovi Pinto and de Ornellas 1980
†Permipsythone perantiqua Martynova 1961
G. †Permithone Tillyard 1922
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†Permithone belmontensis Tillyard 1922
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Invalid names: Permosmylus pincombeae Tillyard 1926 [synonym]
†Permithone oliarcoides Tillyard 1926
Invalid names: Permosmylus Tillyard 1926 [synonym]
G. †Permithonopsis Martynov 1932
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†Permithonopsis cellulosa Martynova 1952
†Permithonopsis dubia Vilesov and Novokshonov 1994
†Permithonopsis enormis Martynova 1952
†Permithonopsis grandis Martynova 1952
†Permithonopsis ivensis Martynov 1932
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Invalid names: Permithonopsis proxima Martynov 1932 [synonym]
†Permithonopsis kaltanensis Martynova 1961
†Permithonopsis longifurcata Martynova 1952
†Permithonopsis obscura Martynova 1952
†Permithonopsis sharovi Martynova 1961
G. †Permopsychops Tillyard 1926
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†Permopsychops karaungirensis Vilesov and Novokshonov 1994
†Permopsychops saurensis Vilesov and Novokshonov 1994
G. †Permorapisma Tillyard 1926
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†Permorapisma biserialis Tillyard 1926
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Invalid names: Permorapisma triserialis Tillyard 1926 [synonym]
†Permorapisma fragmentata Vilesov and Novokshonov 1994
†Permorapisma gori Ponomarenko and Shcherbakov 2004
G. †Permosisyra Martynov 1932
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†Permosisyra latipennis Martynov 1932
†Permosisyra paurovenosa Martynova 1952
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Invalid names: Sialidopsis sojanensis Martynova 1952 [objective synonym]
†Permosisyra punctata Martynov 1932
G. †Sauropsychops Novokshonov 1996
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†Sauropsychops kazakhstanensis Vilesov and Novokshonov 1994
†Sauropsychops tillyardi Vilesov and Novokshonov 1994
G. †Sialidopsis Zalessky 1926
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†Sialidopsis kargalensis Zalessky 1926
†Sialidopsis similis Martynov 1937
G. †Tshekardithonopsis Vilesov 1995
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†Tshekardithonopsis oblivius Vilesov 1995
†Tshekardithonopsis pictus Vilesov 1995
†Tshekardithonopsis zalesskyi Vilesov 1995
Invalid names: Palaemerobiidae Martynov 1928 [synonym], Permegalomidae Martynova 1952 [synonym], Permopsychopsidae Riek 1953 [synonym], Permosisyridae Martynov 1932 [synonym], Sialidopsidae Zalessky 1928 [synonym]
Subor. †Priscaenigmatomorpha Engel 2002
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Fm. †Priscaenigmatidae Engel 2002
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G. †Cretohondelagia Khramov 2020
G. †Hondelagia Bode 1953
G. †Priscaenigma Whalley 1985
G. †Sukachevia Khramov 2020
Or. Raphidioptera Handlirsch 1908 [snakefly]
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G. †Caririraphidia Martins-Neto 2002
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†Caririraphidia reticulata Martins-Neto 2002
†Caririraphidia sertaneja Martins-Neto 2002
G. †Cratoraphidia Martins-Neto and Nel 1993
Fm. †Nanosialidae Shcherbakov 2013
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Subfm. †Nanosialinae Shcherbakov 2013
Subfm. †Raphisialinae Shcherbakov 2013
Fm. †Parasialidae Ponomarenko 1977
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G. †Parasialis Ponomarenko 1977
G. †Sojanasialis Ponomarenko 1977
Subor. Raphidiomorpha Engel 2002
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Fm. †Baissopteridae Martynova 1961
Unr. Raphidiformia Bechly and Wolf-Schwenninger 2011
G. †Partisaniferus Haug et al. 2020
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†Partisaniferus atrickmuelleri Haug et al. 2020
†Partisaniferus edjarzembowskii Haug and Haug 2022
Unr. Panorpida
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Superor. Amphiesmenoptera Kiriakoff 1948
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Or. †Permotrichoptera Martynova 1958
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Fm. †Microptysmatidae Martynova 1958
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G. †Kamopanorpa Martynov 1928
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Invalid names: Microptysma Martynova 1958 [synonym], Microptysmodes Martynova 1958 [synonym], Uraloptysma Ivanov 1992 [synonym]
Invalid names: Uraloptysmatidae Ivanov 1992 [synonym]
Or. †Protomeroptera Engel 2022
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Fm. †Protomeropidae Tillyard 1926
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G. †Glanomerope Poschmann and Nel 2021
G. †Permomerope Tillyard 1926
G. †Pseudomerope Kukalová-Peck and Willmann 1990
G. †Pseudomeropella Kukalová-Peck and Willmann 1990
G. †Stenomerope Kukalová-Peck and Willmann 1990
G. †Westphalomerope Nel et al. 2007
Invalid names: Marimerobiidae Willmann 1978 [synonym], Permomeropidae Riek 1953 [synonym], Platychoristidae Carpenter 1930 [synonym]
Fm. †Stereochoristidae Tillyard 1919
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Invalid names: Necrotaulius handlirschi Fischer 1962 [nomen dubium], Necrotaulius regularis Handlirsch 1939 [nomen dubium], Necrotaulius vicinus Handlirsch 1939 [nomen dubium], Papilionidea Laicharting 1781 [synonym]
Superor. Antliophora Hennig 1969
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Or. Diptera Linnaeus 1758 [fly]
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Infraor. Culicomorpha
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Superfm. Chironomoidea Macquart 1838
Superfm. Culicoidea Meigen 1818
Superfm. Simulioidea Newman 1834
Subor. Diarchineura Krzemiński 1992
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Infraor. †Grauvogeliomorpha Krzemiński et al. 1994
Infraor. †Hennigmatomorpha Krzemiński and Krzemińska 2003
Infraor. †Nadipteromorpha Krzeminski and Evenhuis 2000
Invalid names: Tanyderomorpha [empty]
G. Dicranophora Macquart 1834
Fm. Dryomyzidae Schiner 1864
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G. Dryomyza Fallen 1820
G. †Palaeotimia Meunier 1908
G. †Prodryomyza Hennig 1965
Invalid names: Dryomyzinae Schiner 1864 [empty]
Superfm. Mycetophiloidea Newman 1834
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Fm. †Eoditomyidae Ansorge 1996
Fm. †Protopleciidae Rohdendorf 1946
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Invalid names: Dyspolyneuridae Rohdendorf 1961 [synonym], Palaeopleciidae Rohdendorf 1962 [synonym], Phragmoligoneuridae Rohdendorf 1962 [synonym], Pleciodictyidae Rohdendorf 1962 [synonym], Protoligoneuridae Rohdendorf 1962 [synonym]
Invalid names: Sinoditomyiidae Hong 2002 [invalid subgroup]
Subor. Nematocera Latreille 1825
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G. †Amblylexis Bode 1953
G. †Amianta Bode 1953
G. †Amphipromeca Bode 1953
G. †Apistogrypotes Bode 1953
Fm. †Asiochaoboridae Hong and Wang 1990
G. †Asuba Giebel 1856
Infraor. Axymyiomorpha McAlpine et al. 1981
G. †Bibionites Handlirsch 1906
Infraor. Blephariceromorpha Rohdendorf 1961
G. †Clavapartus Gall and Grauvogel 1966
G. †Culiciscolex Bode 1953
G. †Cyrtomides Bode 1953
G. †Dara Giebel 1856
Fm. †Elliidae Krzemińska et al. 1993
G. †Empidocampe Bode 1953
G. †Eoculex Fritsch 1910
Fm. †Eopolyneuridae Rohdendorf 1962
Fm. †Hyperpolyneuridae Rohdendorf 1961
G. †Meconeura Zhang et al. 1986
Fm. †Mesophantasmatidae Rohdendorf 1962
Fm. †Mesosciophilidae Rohdendorf 1946
Infraor. Nymphomyiomorpha Rohdendorf 1964
G. †Palaeoerioptera Meunier 1899
G. †Paradictyodiptera Rohdendorf 1961
G. †Platyplecia Hong 1983
G. †Propexis Bode 1953
Fm. †Protendipedidae Rohdendorf 1962
Infraor. Ptychopteromorpha
G. †Rhopaloscolex Bode 1953
G. †Sama Giebel 1856
G. †Sphallonymphites Bode 1953
Fm. †Tillyardipteridae Lukashevich and Shcherbakov 1999
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Invalid names: Rhaetaniidae Krzemiński and Krzemińska 2002 [synonym]
Infraor. Tipulomorpha
Fm. †Tipulopleciidae Rohdendorf 1962
Invalid names: Axymyioidea Shannon 1921 [empty], Hongocaloneuridae Hong 2002 [invalid subgroup], Penthetria vaillantii Oustalet 1870 [nomen dubium], Plecia nigrescens Oustalet 1870 [nomen dubium]
Unr. Neodiptera Michelsen 1994
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Infraor. Bibionomorpha Hennig 1954
Subor. Brachycera Zetterstedt 1842
G. †Polyclona Menge 1856
Infraor. Psychodomorpha
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Fm. Deuterophlebiidae Edwards 1922 [mountain midge]
Superfm. Psychodoidea Newman 1834
Superfm. Scatopsoidea Newman 1834
Invalid names: Ptychopteroidea [empty], Tanyderoidea Osten-Sacken 1879 [empty]
Fm. Tipulariae Latreille 1802 [crane fly]
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G. Scathopse Geoffroy 1762
Invalid names: Styringia Berendt 1845 [nomen nudum]
Invalid names: Aemodipsus Aymard 1856 [nomen nudum], Agromyza incerta Theobald 1937 [nomen dubium], Agromyza protogaea Heer 1849 [nomen dubium], Agromyza serresi Theobald 1937 [nomen dubium], Antliata Fabricius 1775 [synonym], Aphis dubia Giebel 1856 [nomen dubium], Archidiptera Rohdendorf 1961 [empty], Asilici Latreille 1802 [empty], Atherix exigua Meunier 1910 [nomen dubium], Atherix saunieri Theobald 1937 [nomen dubium], Bibio batheri Pongrácz 1928 [nomen dubium], Bombylidae Latreille 1802 [empty], Conopsariae Latreille 1802 [empty], Coriaceae Latreille 1802 [empty], Cyrtonotum gigas Theobald 1937 [nomen dubium], Dichaneurum Aymard 1856 [nomen nudum], Eoheleinae Petrunkevitch 1957 [empty], Eudiptera [empty], Halterata Scopoli 1763 [synonym], Inflata Latreille 1802 [empty], Lasioptera recessa Scudder 1877 [nomen dubium], Muscides Latreille 1802 [empty], Mycetobia terricola Scudder 1878 [nomen dubium], Myopidae Swainson 1840 [empty], Nakridletia Vršanský et al. 2010 [synonym], Neoneura Krzemiński 1992 [empty], Oligoneura Brauer 1880 [empty], Phoromorpha [empty], Phytomyza vetusta Theobald 1937 [nomen dubium], Polyneura Brauer 1880 [empty], Rhagionides Latreille 1802 [empty], Simulium ventricum Lin 1988 [nomen dubium], Siphunculata Latreille 1802 [empty], Stratiomydae Latreille 1802 [empty], Syrphiae Latreille 1802 [empty], Tabanii Latreille 1802 [empty], Vinculomusca Townsend 1938 [nomen dubium], Vinculomusca vinculata Scudder 1877 [nomen dubium], Xylomyia shcherbakovi Mostovski 1999 [nomen dubium]
Or. Mecoptera Packard 1886 [scorpionfly]
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Fm. †Choristopsychidae Martynov 1937
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G. †Choristopsyche Martynov 1937
G. †Paristopsyche Qiao et al. 2013
Subor. †Eumecoptera Tillyard 1919
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Fm. †Mesorthophlebiidae Hong 1983
Fm. †Parachoristidae Handlirsch 1937
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Invalid names: Choristopanorpidae Willmann 1987 [synonym], Triassochoristidae Willmann 1989 [synonym]
Fm. †Kaltanidae Martynova 1958
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G. †Cyclorista Martynova 1958
G. †Kaltanochorista Martynova 1958
G. †Kamochorista Bashkuev 2008
G. †Megachorista Martynova 1958
G. †Moravochorista Kukalová-Peck and Willmann 1990
G. †Pinnachorista Martynova 1958
G. †Pseudochorista Novokshonov 1994
G. †Tomiochorista Martynova 1958
Invalid names: Cyclopteridae Martynova 1958 [synonym], Cycloristidae Martynova 1958 [synonym], Tomiochoristidae Martynova 1958 [synonym]
G. †Mecaenigma Novokshonov and Sukacheva 2001
+
†Mecaenigma suspectum Novokshonov and Sukacheva 2001
†Mecaenigma viduum Novokshonov and Sukacheva 2001
Superfm. †Mesopsychoidea Tillyard 1917
+
Fm. †Aneuretopsychidae Rasnitsyn and Kozlov 1990
Fm. †Mesopsychidae Tillyard 1917
+
Invalid names: Mesopanorpodidae Tillyard 1918 [synonym], Mesosetidae Riek 1976 [synonym]
Fm. †Nedubroviidae Bashkuev 2011
Subor. Nannomecoptera Hinton 1981
+
Fm. Nannochoristidae Tillyard 1917
Fm. †Robinjohniidae Martynova 1948
Subor. †Paratrichoptera Tillyard 1919
+
Fm. †Liassophilidae Tillyard 1933
+
Invalid names: Laurentipteridae Martynova and Willmann 1978 [synonym], Pseudodipteridae Martynova 1961 [synonym]
Fm. †Permochoristidae Tillyard 1918
+
Subfm. †Agetopanorpinae Carpenter 1930
G. †Duraznochorista Lara and Bashkuev 2020
G. †Kalochorista Lin 1992
G. †Kazakhopanorpa Novokshonov 1995
G. †Mesageta Novokshonov 1997
Subfm. †Permochoristinae Tillyard 1917
Subfm. †Pseudonannochoristinae Novokshonov 1994
G. †Qingochorista Guo and Hong 2003
Subfm. †Sylvopanorpinae Novokshonov 1997
Invalid names: Caenoptilonidae Zalessky 1933 [synonym], Eosetidae Tindale 1945 [synonym], Idelopanorpidae Zalessky 1929 [synonym], Petrochoristidae Willmann 1989 [synonym], Protochoristinae Martynova 1961 [empty], Protopanorpidae Handlirsch 1937 [synonym], Tychtodelopteridae Ponomarenko 1977 [synonym], Tychtopsychidae Martynova 1958 [synonym], Xenochoristidae Riek 1953 [synonym]
Fm. †Permopanorpidae Tillyard 1926
+
G. †Martynopanorpa Willmann 1989
G. †Neopermopanorpa Riek 1955
G. †Permopanorpa Tillyard 1926
G. †Xenopanorpa Riek 1955
Invalid names: Lithopanorpidae Carpenter 1930 [synonym], Martynopanorpidae Willmann 1989 [synonym]
Fm. †Permotanyderidae Riek 1953
+
Subfm. †Chimaeropterinae Bashkuev 2022
Subfm. †Permotanyderinae Riek 1953
Fm. †Permotipulidae Tillyard 1929
+
G. †Permila Willmann 1989
G. †Permotipula Tillyard 1929
Invalid names: Permilidae Willmann 1989 [synonym]
Subor. Pistillifera Willmann 1987
+
Infraor. Opisthogonopora Willmann 1987
Infraor. Raptipedia Willmann 1987
Fm. †Sibiriothaumatidae Sukatsheva and Novokshonov 1998
+
G. †Sibiriothauma Sukatsheva and Novokshonov 1998
Fm. †Sinoagetopanorpidae Lian et al. 2023
+
G. †Permoagetopanorpa Lian et al. 2023
G. †Raragetopanorpa Lian et al. 2023
G. †Sinoagetopanorpa Lin et al. 2010
Fm. †Thaumatomeropidae Willmann 1978
+
G. †Blattomerope Rasnitsyn 1974
G. †Thaumatomerope Rasnitsyn 1974
Invalid names: Aneuretopsychina Rasnitsyn and Kozlov 1990 [invalid subgroup], Cyclopterinidae Carpenter 1986 [empty], Eobanksiidae Cockerell 1909 [empty], Panorpata Brauer [synonym], Paramecoptera Tillyard 1919 [empty], Protodiptera Tillyard 1937 [invalid subgroup], Protomecoptera Tillyard 1917 [empty]
Or. Siphonaptera Latreille 1825 [flea]
+
Superfm. †Saurophthiroidea Ponomarenko 1986
+
Fm. †Pseudopulicidae Gao et al. 2012
Fm. †Saurophthiridae Ponomarenko 1986
+
Invalid names: Saurophthiridae Gao et al. 2013 [objective synonym]
Fm. †Tarwiniidae Huang et al. 2013
Invalid names: Hystrichopsyllidae Tiraboschi 1904 [empty]
Invalid names: Neuropteroidea [empty]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
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| Source: c = class, p = phylum | |||||
| References: Bush and Bambach 2015, Kiessling 2004 | |||||
Age range
Maximum range based only on fossils: base of the Late/Upper Eifelian to the top of the Holocene or 389.60000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 387.7 Ma
Minimum age of oldest fossil (stem group age): 387.7 Ma
Collections (3191 total)
Oldest occurrences | |||
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| Late/Upper Eifelian | Germany | Schizophora blankenheimensis (235174) | |
| Westphalian A | France | Westphalomerope maryvonneae (122506) | |
| Moscovian | Czech Republic | Phryganea kolbi, Phryganea arenacea (125662) | |
| Westphalian C | Czech Republic | Phryganea solitaria (155300) | |
| Westphalian D | Germany (Lower Saxony) | Microcarbonella paradoxa (224159) | |
| Gzhelian | France | Stephanastus polinae (121979) | |
| Asselian | Germany | Glanomerope virgoferroa, Moscheloptera phantasma (112604) | |
| Asselian | Germany (Rhineland-Palatinate) | Coleopsis archaica (156248) | |
| Asselian | USA (New Mexico) | Mecoptera indet. (124386) | |
| Asselian | Germany (Thüringer Wald) | Pachypleurites nodosus, Permorhipida furcativenosa (128485) | |
| Asselian | Czech Republic | Microptysmella moravica, Protomeropidae indet., Pseudomerope havlati, Pseudomerope oborana, Pseudomerope mareki, Pseudomeropella nekvasilovae, Stenomerope spinari, Moravochorista carolina, Moravocoleus perditus, Tshekardocoleidae indet., Moravocoleus permianus, Retelytron conopeum, Votocoleus submissus, Boscoleus blandus, Eocoleus scaber, Prosperocoleus prosperus, Umoricoleus perplex, Moravocoleus neglegens, Moravocoleus fractus, Liberocoleus intactus, Oborocoleus rohdendorfi (114186) Pseudomerope gallei (114141) | |
| Asselian - Kungurian | Russian Federation | Minusinia martynovae (126338) | |
| Artinskian | USA (Oklahoma) | Permocoleus wellingtonensis (104816) Permocoleus wellingtonensis, Permoberotha villosa (104826) Permopanorpa inaequalis (107405) | |
| Artinskian | France (Languedoc) | Lodevosisyra reducta (105044) | |
| Artinskian | South Africa (Western Cape) | Afrocupes firmae (156986) | |
| Artinskian | USA (Kansas) | Permoberotha convergens (111320) Permoberotha villosa (111319) Permopanorpa raymondi (151412) Platychorista venosa, Agetopanorpa maculata, Protochorista tetraclada, Protopanorpa permiana, Permopanorpa inaequalis, Elmothone martynovae (111217) Platychorista venosa, Protomerope permiana, Anormochorista oligoclada, Protochorista tetraclada, Protochorista pentaclada, Protopanorpa permiana, Permopanorpa schucherti, Permopanorpa inaequalis, Permopanorpa sellardsi, Permopanorpa tenuis, Permopanorpa dunbari, Permopanorpa gracilis, Permopanorpa formosa, Protopanorpa pusilla, Permoberotha villosa, Dictyobiella nervosa, Permoberotha convergens (111219) | |
| Artinskian - Kungurian | Russian Federation (Kemerovo) | Mesochorista javorskyi, Asiachorista neuburgae (124614) | |
| Kungurian | Brazil (Rio Grande do Sul) | Petromantis evansi, Asiachorista beckermigdisovae, Protocupoides rohdendorfi, Kaltanicupes ponomarenkoi, Permipsythone panfilovi (116121) Petromantis rieki (116120) | |
| Kungurian | Russian Federation (Suksun) | Kamopanorpa fasciipennis, Agetochorista intermedia, Tshekardocoleus magnus (116652) Kamopanorpa fasciipennis, Marimerobius sp., Agetochorista sp., Agetochorista tillyardi, Agetochorista permiana, Sylvopanorpa carpenteri, Permochorista sylvaensis (116641) Kamopanorpa pritykinae, Kamopanorpa uralensis, Kamopanorpa incerta, Uraloptysma maculata, Kamopanorpa pallida, Protomeropidae indet., Marimerobius sp., Tshekardopanorpa biarmica, Agetopanorpa sp., Agetochorista kungurica, Agetochorista intermedia, Agetochorista sp., Agetochorista tillyardi, Agetochorista punctata, Agetochorista permiana, Phipoides pusillus, Phipoides ornatus, Protopanorpoides medius, Protopanorpoides minutus, Uraloageta archaica, Petromantis sylvaensis, Tshekardocoleidae indet., Sylvacoleus richteri, Sylvacoleus sharovi, Sylvacoleodes admirandus, Tshekardocoleus minor, Sylvaelytron latipennatum, Kunguromaritus lacer, Kunguromaritus guttatus, Uralisyra angusta, Tshekardithonopsis zalesskyi, Tshekardithonopsis pictus, Sylvamarita minor, Sylvasenex lacrimabunda, Jurla bisubcostata, Okolpania observabilis (108206) Parachorista uralensis (116642) Parachorista uralensis, Kamopanorpa fasciipennis (113379) Parachorista uralensis, Kamopanorpa fasciipennis, Protomeropidae indet., Kunguromaritus lacer (124241) Protomeropidae indet. (122864) Protomeropidae indet., Curvilarva tshekardensis (122753) | |
| Kungurian | South Africa (West Cape) | Mecoptera indet. (117279) | |