PBDB Taxon

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Testudines (turtle)

Reptilia - Testudines

Taxonomy

Testudines was named by Batsch (1788) [used as the plural of Testudo by Linnaeus and not as a proper higher order name: see Dundee 1989]. It is extant. It was considered monophyletic by Joyce (2007), Danilov and Parham (2008), Pérez-García and Murelaga (2013).

It was synonymized subjectively with Testudinata by Hay (1930); it was reranked as the unranked clade Testudines by Laurin and Reisz (1995), Li et al. (2008); it was reranked as the subclass Testudines by Benton (1985), Agnolín et al. (2023).

It was assigned to Chelonomorpha by Kuhn (1966), Haubold (1971); to Chelonomorpha by Haubold (1974); to Amniota by Benton (1985); to Testudinomorpha by Laurin and Reisz (1995); to Anapsida by Gauthier et al. (1989), Peng et al. (2001); to Testudinata by Li et al. (2008); to Reptilia by Hay (1902), Holman and Case (1988), Shoshani et al. (1989), Frank and Ramus (1996), Weems and George (2001), Pearson et al. (2002), Meyer and Zardoya (2003), Parham and Pyenson (2010), Garcia et al. (2016), Turvey et al. (2017); and to Testudinata by Joyce (2007), Danilov and Parham (2008), Sterli (2008), Pérez-García and Murelaga (2013), Perea et al. (2014), Agnolín et al. (2023).

Year	Name and author
1788	Testudines Batsch
1800	Chelonii Latreille
1842	Chelone obovata Owen p. 170
1847	Apholidemys Pomel
1854	Chelonia Mantell p. 646
1859	Chelonia Owen p. 153
1860	Chelonia Owen p. 281
1861	Chelonia Owen p. 314
1865	Chelone sopita Leidy p. 104
1866	Chelonia Haeckel p. CXXXVIII
1868	Osteopygis sopita Cope p. 147
1870	Propleura sopita Cope p. 140
1873	Chelonia Leidy p. 339
1882	Chelonii Quenstedt p. 145
1884	Apholidemys Cope
1886	Chelonia Günther p. 445
1895	Podocnemis lata Ristori
1898	Chelonia Gadow p. 20
1898	Chelonia Woodward p. 141
1902	Testudines Hay p. 434
1902	Osteopygis sopitus Hay p. 441
1904	Chelonia Case p. 60
1906	Chelonia Andrews p. 275
1908	Osteopygis sopitus Hay p. 149 figs. figs. 181-184
1908	Chelonia Lambe
1909	Platychelone Dollo p. 105
1909	Platychelone emarginata Dollo p. 105
1911	Chelonii Jaekel p. 186
1913	Chelonia Broom p. 364
1917	Chelonia Williston p. 420
1928	Chelonia Perrier pp. 3111, 3118
1930	Osteopygis sopitus Hay p. 76
1934	Apertotemporalis Stromer
1934	Apertotemporalis baharijensis Stromer
1934	Chelonia Swinton p. 70
1947	Chelonia Olson pp. 44, 45
1954	Chelonia Haughton and Brink p. 3
1955	Euclastochelys Bergounioux
1955	Euclastochelys interrupta Bergounioux
1958	Temnoclemmys cordiformis Bergounioux
1958	Pleurosternum obovata Delair p. 49
1966	Senonemys suemegensis Bohn
1966	Testudines Kuhn p. 20
1971	Testudines Haubold p. 89
1973	Trionyx tshelkarensis Chkhikvadze
1974	Testudines Haubold p. 49
1974	Trionyx fuchienensis Yeh
1979	Sokotochelys Halstead
1979	Sokotochelys lawanbungudui Halstead
1979	Sokotochelys umarumohammedi Halstead
1981	Chelonia Thurmond and Jones p. 114
1982	Chelonia Gardiner p. 228
1985	Testudines Benton p. 153
1988	Apertotemporalis Carroll
1988	Chelonia Carroll
1988	Sokotochelys Carroll
1988	Padiochelys Carroll p. 616
1988	Testudines Holman and Case p. 328
1989	Testudines Gauthier et al. p. 343
1989	Chelonia Gauthier et al. p. 344
1989	Testudines Shoshani et al. p. 436
1993	Chelonia Cvancara and Hoganson p. 15
1995	Testudines Laurin and Reisz p. 172
1995	Chelonia Lee pp. 508, 515 fig. 22
1996	Testudines Frank and Ramus
2001	Testudines Peng et al. p. 23
2001	Testudines Weems and George p. 572
2002	Testudines Pearson et al. p. 154
2002	Chelonia Sepkoski
2003	Testudines Meyer and Zardoya
2004	Chelonia Martinelli and Forasiepi p. 260
2007	Testudines Joyce
2008	Testudines Danilov and Parham
2008	Testudines Li et al. p. 500
2008	Chelonii Previtera and González Riga
2008	Testudines Sterli p. 286
2010	Testudines Parham and Pyenson p. 232
2010	Meiolania damelipi White et al. p. 1
2013	Testudines Pérez-García and Murelaga
2014	Testudines Perea et al.
2015	Meiolania damelipi Rhodin et al.
2016	Testudines Garcia et al.
2017	Testudines Turvey et al.
2023	Testudines Agnolín et al.

Or. Testudines Batsch 1788 [turtle]

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Subor. †Amphichelydia Lydekker 1889

Superfm. †Pleurosternoidea Hay 1930

Fm. †Kallokibotiidae Nopcsa 1923

G. †Kallokibotion Nopcsa 1923

Invalid names: Apertotemporalidae [empty]

Superfm. †Proganochelyoidea Baur 1887

Invalid names: Archaeochelydiidae Bergounioux 1938 [empty]

G. †Asturichelys Bergounioux 1957

G. †Chelonipus von Lilienstern 1939

†Chelonipus cuneiformis von Lilienstern 1939

†Chelonipus plieningeri Haubold 1971

†Chelonipus torquatus von Lilienstern 1939

Subor. Cryptodira Cope 1868

Fm. †Bashuchelyidae Tong et al. 2012

G. †Bashuchelys Tong et al. 2012

†Bashuchelys youngi Tong et al. 2012

†Bashuchelys zigongensis Yeh 1982

G. †Chuannanchelys Tong et al. 2012

†Chuannanchelys dashanpuensis Fang 1987

Fm. Chelonemydidae Rütimeyer 1872

Invalid names: Propleuridae Cope 1870 [synonym]

Superfm. Chelonioidea Baur 1893 [sea turtle]

Fm. Cheloniidae Bonaparte 1832

Subfm. †Allopleuroninae

G. †Allopleuron Baur 1888

G. †Glyptochelone Dollo 1903

G. †Protosphargis Capellini 1884

G. †Ashleychelys Weems and Sanders 2014

†Ashleychelys palmeri Weems and Sanders 2014

G. †Cabindachelys Myers et al. 2018

†Cabindachelys landanensis Myers et al. 2018

Subfm. Carettinae Gray 1825

G. Caretta Rafinesque 1814 [loggerhead]

Invalid names: Caouana Cocteau and Bibron 1838 [synonym], Halichelys Fitzinger 1843 [synonym], Thalassochelys Fitzinger 1835 [synonym]

G. Eretmochelys Fitzinger 1843

Invalid names: Herpysmostes Gistel 1868 [synonym], Onychochelys Gray 1873 [synonym]

G. Chelone Linnaeus 1758

†Chelone costata Owen 1853

†Chelone girundica Delfortrie 1869

†Chelone jessoni Lydekker 1889

†Chelone vanbenedeni Smets 1886

Subfm. Cheloniinae Oppel 1811

G. Chelonia Brongniart 1800 [green sea turtle]

Invalid names: Chelona Brongniart 1805 [synonym], Euchelys Girard 1858 [synonym], Megemys Gistel 1848 [synonym], Mydasea Gervais 1843 [synonym]

Tr. Cheloniini Zangerl and Turnball 1955

G. Natator McCulloch 1908

Tr. †Rupelchelyini Karl and Tichy 1999

Invalid names: Carettini Zangerl and Turnbull 1955 [empty]

G. †Chelyopsis Smets 1887

†Chelyopsis halleri Karl 1989

†Chelyopsis holsaticus Dames 1894

†Chelyopsis littoreus Van Beneden 1886

†Chelyopsis raabei Karl and Schleich 1994

G. †Dollochelys Zangerl 1971

†Dollochelys rogovichi Averianov 2002

G. †Eochelone Dollo 1903

†Eochelone athersuchi Moody 1980

†Eochelone brabantica Dollo 1903

†Eochelone monstigris Grant-Mackie et al. 2011

†Eochelone voltregana de Lapparent de Broin et al. 2018

G. †Erquelinnesia Dollo 1887

†Erquelinnesia gosseleti Dollo 1886

†Erquelinnesia planimenta Owen 1841

G. †Gigantatypus Kaddumi 2006

†Gigantatypus salahi Kaddumi 2006

G. †Itilochelys Danilov et al. 2010

†Itilochelys rasstrigin Danilov et al. 2010

G. †Lembonax Cope 1870

†Lembonax polemicus Cope 1870

†Lembonax propylaeus Cope 1872

G. Lepidochelys Fitzinger 1843 [Ridley sea turtle]

Lepidochelys kempii Garman 1880 [Kemp's Ridley turtle]

Invalid names: Testudo viridisquamosa Lacepède 1788 [synonym]

Lepidochelys olivacea Eschscholtz 1829

Invalid names: Caretta remivaga Hay 1908 [synonym], Cephalochelys oceanica Gray 1873 [synonym], Chelonia dussumierii Dumeril and Bibron 1835 [synonym], Thalassochelys controversa Philippi 1899 [synonym], Thalassochelys tarapacona Philippi 1887 [synonym]

Invalid names: Caouana ruppellii Gray 1844 [nomen nudum], Cephalochelys Gray 1873 [synonym], Chelonia dubia Bleeker in Gray 1864 [nomen nudum], Chelonia polyaspis Bleeker 1857 [nomen nudum], Testudo bomarii Meyer 1790 [nomen oblitum]

G. †Lusochelys Pérez-García and Antunes 2024

†Lusochelys emilianoi Pérez-García and Antunes 2024

G. †Lytoloma Cope 1870

†Lytoloma bruxelliensis Dollo 1909

†Lytoloma cantabrigiense Lydekker 1889

†Lytoloma crassa Bergounioux 1952

†Lytoloma elegans Bergounioux 1952

Lytoloma jeanesii Cope 1870

†Lytoloma wemmeliensis Dollo 1909

G. †Miocaretta Deraniyagala 1967

†Miocaretta lankae Deraniyagala 1967

G. †Osonachelus de Lapparent de Broin et al. 2014

†Osonachelus decorata de Lapparent de Broin et al. 2014

G. †Pacifichelys Parham and Pyenson 2010

†Pacifichelys hutchisoni Lynch and Parham 2003

†Pacifichelys urbinai Parham and Pyenson 2010

G. †Procolpochelys Hay 1908

†Procolpochelys charlestonensis Weems and Sanders 2014

†Procolpochelys grandaeva Leidy 1861

†Procolpochelys susensis Shikama and Suyama 1976

G. †Protrachyaspis Zvonok et al. 2024

†Protrachyaspis shorymensis Zvonok et al. 2024

G. †Puppigerus Cope 1870

†Puppigerus breviceps Owen 1841

†Puppigerus camperi Gray 1831

Invalid names: Lytoloma trigoniceps Owen 1849 [synonym], Puppigerus longiceps Owen 1841 [synonym], Puppigerus nessovi Averianov 2005 [synonym]

†Puppigerus crassicostata Owen 1849

Invalid names: Puppigerus latiscutatus Owen 1841 [synonym]

†Puppigerus grandaevus Leidy 1861

†Puppigerus parvisecta Cope 1870

Invalid names: Eochelys Moody 1968 [objective synonym]

Subfm. †Syllominae

G. †Bryochelys van Beneden 1871

G. †Kurobechelys Shikama 1956

G. †Syllomus Cope 1896

†Syllomus aegyptiacus Lydekker 1899

Invalid names: Chelonia marylandica Collins and Lynn 1936 [synonym], Peritresius virginianus Berry and Lynn 1936 [synonym]

†Syllomus crispatus Cope 1896

G. †Tasbacka Nesov 1986

†Tasbacka aldabergeni Nessov 1987

†Tasbacka danica Karl and Madsen 2012

†Tasbacka ouledabdounensis Tong and Hirayama 2002

†Tasbacka ruhoffi Weems 1988

†Tasbacka salisburgensis Karl 1996

G. †Trachyaspis von Meyer 1843

†Trachyaspis aegyptiaca Lydekker 1889

Trachyaspis bellunensis Dal Piaz 1916

†Trachyaspis hantoniensis Lydekker 1889

†Trachyaspis lardyi Meyer 1873

Invalid names: Testudo varicosa Costa 1851 [synonym], Trionyx bellunensis Misuri 1911 [synonym]

Invalid names: Trionyx melitensis Lydekker 1891 [nomen dubium]

G. Zangerlchelys Hirayama 2006

†Zangerlchelys arkansaw Schmidt 1944

Invalid names: Chelone acuticostata Bergounioux 1954 [nomen vanum], Chelone bellunensis Bergounioux 1954 [nomen vanum], Chelone gastaldii Portis 1879 [nomen vanum], Chelone murua de Vis 1905 [nomen dubium], Chelone simonellii Porta 1898 [nomen vanum], Chelone sismondai Portis 1879 [nomen vanum], Chelone sordellii Porta 1898 [nomen vanum], Chelone strobeli Porta 1898 [nomen vanum], Eocheloniinae [empty], Eochelyinae Moody 1967 [empty], Eretmochelyinae Deranyiagala 1953 [empty], Pliochelys derelicta Portis 1890 [nomen vanum], Progonosaurus pertinax Portis 1890 [nomen vanum]

G. †Cimochelys Zangerl 1960

†Cimochelys benstedi Mantell 1841

Fm. Dermochelyidae Lydekker 1889 [leatherback]

G. †Arabemys Tong et al. 1999

†Arabemys crassiscutata Tong et al. 1999

Subfm. †Dermochelyinae

G. †Cosmochelys Andrews 1920

G. Dermochelys de Blainville 1816 [Leatherback sea turtle]

Dermochelys coriacea Vandelli 1761 [Leatherback sea turtle]

Invalid names: Chelonias lutaria Rafinesque 1814 [synonym], Dermatochelys porcata Wagler 1830 [synonym], Sphargis angusta Philippi 1899 [synonym], Sphargis mercurialis Merrem 1820 [synonym], Testudo arcuata Catesby 1771 [synonym], Testudo lyra Lacepede 1788 [synonym], Testudo tuberculata Pennant in Schoepff 1801 [synonym]

Invalid names: Chelonias Rafinesque 1814 [nomen oblitum], Chelyra Rafinesque 1815 [nomen nudum], Chelyra Rafinesque 1832 [synonym], Coriudo Fleming 1822 [objective synonym], Dermatochelys Wagler 1830 [synonym], Dermochelis Cuvier 1829 [synonym], Dermochelis atlantica LeSueur in Cuvier 1829 [nomen nudum], Scytina Wagler 1828 [synonym], Sphargis Merrem [synonym]

G. †Egyptemys Wood et al. 1996

†Egyptemys eocaenus Andrews 1901

†Egyptemys oregonensis Packard 1940

G. †Eosphargis Lydekker 1889

†Eosphargis breineri Nielsen 1959

†Eosphargis gigas Owen 1880

Invalid names: Anglocetus beatsoni Tarlo 1963 [synonym]

†Eosphargis insularis Cope 1872

Invalid names: Anglocetus Tarlo 1963 [synonym]

G. †Maorichelys Karl and Tichy 2007

†Maorichelys wiffeni Karl and Tichy 2007

G. †Mesodermochelys Hirayama and Chitoku 1996

†Mesodermochelys undulatus Hirayama and Chitoku 1996

G. †Natemys Wood et al. 1996

†Natemys peruvianus Wood et al. 1996

G. †Psephophorus von Meyer 1847

†Psephophorus polygonus von Meyer 1847

Invalid names: Psephophorus calvertensis Palmer 1909 [synonym]

†Psephophorus pseudostracion Gervais 1849

†Psephophorus rupeliensis Van Beneden 1883

†Psephophorus scaldii van Beneden 1871

†Psephophorus terrypratchetti Köhler 1995

G. †Turgaiscapha Averianov 2002

†Turgaiscapha kushmurunica Averianov 2002

Invalid names: Chelonia californiensis Gilmore 1937 [invalid subgroup], Psephophoridae Goette 1899 [synonym], Sphargidae Gray 1825 [synonym]

Unr. †Dermochelyoidae Williams 1950

G. †Alienochelys de Lapparent de Broin et al. 2014

†Alienochelys selloumi de Lapparent de Broin et al. 2014

G. †Corsochelys Zangerl 1960

†Corsochelys bentleyi Schwimmer et al. 2015

†Corsochelys halinches Zangerl 1960

G. †Ocepechelon Bardet et al. 2013

†Ocepechelon bouyai Bardet et al. 2013

Fm. †Desmatochelyidae Williston 1894

G. †Atlantochelys Agassiz 1849

†Atlantochelys mortoni Agassiz 1849

Invalid names: Protostega neptunia Cope 1872 [objective synonym]

G. †Neptunochelys Wieland 1900

†Neptunochelys tuberosa Cope 1872

G. †Kimurachelys Brinkman et al. 2015

†Kimurachelys slobodae Brinkman et al. 2015

G. †Leviathanochelys Castillo-Visa et al. 2022

†Leviathanochelys aenigmatica Castillo-Visa et al. 2022

G. †Nichollsemys Brinkman et al. 2006

†Nichollsemys baieri Brinkman et al. 2006

Fm. †Protostegidae Cope 1873

G. †Archelon Wieland 1896

†Archelon ischyros Wieland 1896

†Archelon marshii Wieland 1900

G. †Bouliachelys Kear and Lee 2006

†Bouliachelys suteri Kear and Lee 2006

Subfm. †Chelospharginae Zangerl 1953

G. †Teguliscapha Nessov

G. †Chelosphargis Zangerl 1953

†Chelosphargis advena Hay 1908

G. †Cratochelone Longman 1915

†Cratochelone berneyi Longman 1915

G. †Desmatochelys Williston 1894

†Desmatochelys lowii Williston 1894

†Desmatochelys padillai Cadena and Parham 2015

G. †Notochelone Lydekker 1889

†Notochelone costata Owen 1882

Invalid names: Notochelys Owen 1882 [replaced]

G. †Pneumatoarthrus Cope 1870

†Pneumatoarthrus peloreus Cope 1870

G. †Protostega Cope 1871

†Protostega eaglefordensis Zangerl 1953

†Protostega gigas Cope 1871

Invalid names: Protostega dixie Zangerl 1953 [synonym], Protostega potens Hay 1908 [synonym]

Subfm. †Protosteginae Wieland

G. †Calcarichelys Zangerl 1953

G. †Microstega Hooks 1998

G. †Rhinochelys Seeley 1869

†Rhinochelys amaberti Moret 1935

†Rhinochelys cantabrigiensis Lydekker 1889

Invalid names: Rhinochelys cardiocephalus Seeley 1869 [synonym], Rhinochelys dayi Seeley 1869 [synonym], Rhinochelys eurycephalus Seeley 1869 [synonym], Rhinochelys jessoni Lydekker 1889 [synonym], Rhinochelys platyrhinus Seeley 1869 [synonym], Rhinochelys rheporhinus Seeley 1869 [synonym], Rhinochelys sphenicephalus Seeley 1869 [synonym]

†Rhinochelys elegans Lydekker 1889

Invalid names: Rhinochelys brachyrhina Lydekker 1889 [synonym], Rhinochelys macrorhina Lydekker 1889 [synonym], Rhinochelys mastocephalus Seeley 1869 [synonym], Rhinochelys stenicephalus Seeley 1869 [synonym]

†Rhinochelys nammourensis Tong et al. 2006

†Rhinochelys pulchriceps Owen 1842

Invalid names: Rhinochelys dacognathus Seeley 1869 [synonym]

G. †Santanachelys Hirayama 1998

†Santanachelys gaffneyi Hirayama 1998

G. †Terlinguachelys Lehman and Tomlinson 2004

†Terlinguachelys fischbecki Lehman and Tomlinson 2004

Fm. †Toxochelyidae Baur 1895

G. †Cynocercus Cope 1872

†Cynocercus incisus Cope 1872

Subfm. †Lophochelyinae Zangerl 1953

G. †Lophochelys Zangerl 1953

G. †Neurochelys Moody 1980

G. †Oertelia Karl et al. 2012

Subfm. †Macrobaeninae Nessov 1984

G. †Toxocheloides Nessov and Khozatsky 1981

Subfm. †Osteopyginae Zangerl 1953

G. †Propleura Cope 1870

G. †Rhetechelys Hay 1908

Invalid names: Lytolominae [synonym]

G. †Porthochelys Williston 1901

†Porthochelys laticeps Williston 1901

Subfm. †Toxochelyinae Zangerl 1953

G. †Thinochelys Zangerl 1953

Invalid names: Glyptops belviderensis Cragin 1902 [nomen dubium]

Invalid names: Pandermochelys Joyce 2004 [empty], Pliochelys Portis 1890 [nomen vanum], Progonosaurus Portis 1890 [nomen vanum]

Fm. Chelydridae Swainson 1839

G. Chelydra Schweigger 1812 [Snapping turtle]

Chelydra acutirostris Peters 1862 [South American snapping turtle]

Chelydra rossignonii Bocourt 1868 [Central American snapping turtle]

Chelydra serpentina Linnaeus 1758 [Common snapping turtle]

Invalid names: Chelydra emarginata Agassiz 1857 [synonym], Chelydra lacertina Schweigger 1812 [synonym], Chelydra laticarinata Hay 1916 [synonym], Chelydra osceola Stejneger 1918 [synonym], Chelydra sculpta Hay 1916 [synonym], Devisia mythodes Ogilby 1905 [synonym], Macrochelys floridana Hay 1907 [synonym], Testudo longicauda Shaw in Gray 1831 [synonym], Testudo serpentaria Wiedemann 1802 [synonym], Testudo serrata Pennant in Gray 1830 [synonym]

Invalid names: Cheliurus Rafinesque 1815 [nomen nudum], Chelonura Fleming 1822 [synonym], Devisia Ogilby 1905 [synonym], Emysaurus Dumeril and Bibron 1835 [synonym], Ophichelone Jarocki 1822 [synonym], Rapara Gray 1825 [synonym], Saurochelys Latreille 1825 [synonym]

G. †Chelydrops Peters 1868

Subfm. †Chelydropsinae Mlynarski 1980

G. †Chelydrasia Ckhikvadze 1999

G. †Chelydropsis Peters 1868

†Chelydropsis apellanizi Murelaga et al. 1999

†Chelydropsis aubasi Joyce et al. 2022

†Chelydropsis decheni Meyer 1852

Invalid names: Chelydropsis sanctihenrici Broin 1977 [synonym]

†Chelydropsis heweneggensis Pappa et al. 2024

†Chelydropsis kusnetzovi Chkhikvadze 1985

†Chelydropsis manuascensis Bergounioux 1936

†Chelydropsis murchisoni Bell 1836

Invalid names: Chelydra allingensis Fuchs 1939 [synonym], Chelydropsis carinata Peters 1868 [synonym], Chelydropsis murchisoni staeschei Mlynarski 1980 [synonym], Chelydropsis staeschei Mlynarski 1980 [synonym], Trionyx sansaniensis Bergounioux 1935 [synonym]

†Chelydropsis pontica Pidoplichko and Tarashchuk 1960

†Chelydropsis sansaniensis Bergounioux 1935

Invalid names: Macrocephalochelys Pidoplichko and Tarashchuk 1960 [synonym]

G. Macrochelys Gray 1856 [alligator snapping turtle]

†Macrochelys auffenbergi Dobie 1968 [Auffenberg's snapping turtle]

†Macrochelys schmidti Zangerl 1945

†Macrochelys stricta Matthew 1924

Macrochelys suwanniensis Thomas et al. 2014 [Suwannee alligator snapping turtle]

Macrochelys temminckii Troost 1835 [Alligator snapping turtle]

Invalid names: Gypochelys lacertina Agassiz 1857 [synonym], Macrochelys apalachicolae Thomas et al. 2014 [synonym]

Invalid names: Gypochelys Agassiz 1857 [synonym]

G. †Planiplastron Ckhikvadze 1971

Invalid names: Broilia Bergounioux and Crouzel 1965 [nomen dubium], Broilia robusta Bergounioux and Crouzel 1965 [nomen dubium], Chelydra argillarum Laube 1900 [nomen dubium], Chelydra strausi Schmidt 1966 [nomen dubium], Chelydrinae Lydekker 1889 [empty], Chelydropsis minax Ckhikvadze 1971 [nomen dubium], Emys sansaniensis Lartet 1851 [nomen dubium], Emysaurus meilheuratiae Pomel 1846 [nomen dubium], Hydraspis oeningensis Fitzinger 1835 [nomen nudum], Leptochelys Bergounioux and Crouzel 1965 [nomen dubium], Leptochelys braneti Bergounioux and Crouzel 1965 [nomen dubium], Protochelydrinae Gaffney 1975 [empty], Testudo grandis Macarovici and Vancea 1959 [nomen dubium]

Unr. Chelydroidea Baur 1893

Unr. †Panchelydridae Joyce et al. 2004

G. †Denverus Hutchison and Holroyd 2003

†Denverus middletoni Hutchison and Holroyd 2003

G. †Protochelydra Erickson 1973

†Protochelydra zangerli Erickson 1973

Invalid names: Hoplochelys caelata Hay 1909 [nomen dubium]

Unr. †Pankinosternoidea Joyce et al. 2004

G. †Emarginachelys Whetstone 1978

†Emarginachelys cretacea Whetstone 1978

G. †Lutemys Lyson et al. 2017

†Lutemys warreni Lyson et al. 2017

G. †Planetochelys Weems 1988

†Planetochelys dithyros Hutchison 2013

†Planetochelys savoiei Weems 1988

G. †Tullochelys Hutchison 2013

†Tullochelys montana Hutchison 2013

G. †Chitracephalus Dollo 1884

†Chitracephalus dumonii Dollo 1885

Invalid names: Salasemys pulcherrima Fuentes et al. 2003 [synonym]

Invalid names: Salasemys Fuentes et al. 2003 [synonym]

Superfm. †Dermochelyoidea

G. †Pygmaeochelys Laube 1896

Invalid names: Chelone regularis Fritsch 1905 [nomen dubium], Pygmaeochelys michelobana Laube 1896 [nomen dubium]

Unr. Durocryptodira Danilov and Parham 2008

G. †Toxochelys Cope 1873

†Toxochelys elkader Hay 1908

†Toxochelys latiremis Cope 1873

Invalid names: Porthochelys browni Hay 1905 [synonym], Toxochelys barberi Schmidt 1944 [synonym], Toxochelys brachyrhinus Case 1898 [synonym], Toxochelys serrifer Cope 1875 [synonym], Toxochelys weeksi Collins 1951 [synonym]

†Toxochelys moorevillensis Zangerl 1953

†Toxochelys procax Hay 1905

Unr. Eucryptodira Gaffney 1975

G. †Borealochelys Brinkman and Tarduno 2005

†Borealochelys axelheibergensis Brinkman and Tarduno 2005

G. †Enaliochelys Seeley 1869

†Enaliochelys chelonia Seeley 1869

G. †Hoyasemys Pérez-García et al. 2012

†Hoyasemys jimenezi Pérez-García et al. 2012

Fm. †Sandownidae Tong and Meylan 2013

G. †Angolachelys Mateus et al. 2009

†Angolachelys mbaxi Mateus et al. 2009

G. †Brachyopsemys Tong and Meylan 2013

†Brachyopsemys tingitana Tong and Meylan 2013

G. †Leyvachelys Cadena 2015

†Leyvachelys cipadi Cadena 2015

G. †Sandownia Meylan et al. 2000

†Sandownia harrisi Meylan et al. 2000

Fm. †Sinemydidae Yeh 1963

G. †Dracochelys Gaffney and Ye 1992

†Dracochelys bicuspis Gaffney and Ye 1992

G. †Hongkongochelys Ye 1999

†Hongkongochelys yehi Ye 1999

G. †Jeholochelys Shao et al. 2018

†Jeholochelys lingyuanensis Shao et al. 2018

G. †Liaochelys Zhou 2010

†Liaochelys jianchangensis Zhou 2010

G. †Manchurochelys Endo and Shikama 1942

†Manchurochelys manchoukuoensis Endo and Shikama 1942

G. †Ordosemys Brinkman and Peng 1993

†Ordosemys brinkmania Danilov and Parham 2007

†Ordosemys donhai Ma 1986

†Ordosemys leios Brinkman and Peng 1993

†Ordosemys liaoxiensis Ji 1995

†Ordosemys perforata Sukhanov and Narmandakh 2006

G. †Sinemys Wiman 1930

†Sinemys brevispinus Tong and Brinkman 2012

†Sinemys chabuensis Ji and Chen 2018

†Sinemys gamera Brinkman and Peng 1993

†Sinemys lens Wiman 1930

G. †Wuguia Matzke et al. 2004

†Wuguia efremovi Khozatsky 1996

Invalid names: Dracochelys wimani Maisch et al. 2003 [synonym]

†Wuguia hutubeiensis Matzke et al. 2004

G. †Xiaochelys Zhou and Rabi 2015

†Xiaochelys ningchengensis Zhou and Rabi 2015

G. †Yumenemys Bohlin 1953

†Yumenemys inflatus Bohlin 1953

Invalid names: Kirgizemyinae Nessov and Khozatskiy 1973 [empty]

Fm. †Sinochelyidae Chkhikvadze 1970

G. †Sinochelys Wiman 1930

†Sinochelys applanata Wiman 1930

Invalid names: Heishanemys imperfectus Bohlin 1953 [synonym], Peishanemys latipons Bohlin 1953 [synonym], Scutemys tecta Wiman 1930 [synonym]

Invalid names: Heishanemys Bohlin 1953 [synonym], Peishanemys Bohlin 1953 [synonym], Scutemys Wiman 1930 [synonym]

Invalid names: Peishanemydidae Nessov 1981 [synonym]

G. †Tienfucheloides Nessov 1978

†Tienfucheloides undatus Nessov 1978

Fm. †Xinjiangchelyidae Nessov 1990

G. †Annemys Sukhanov and Narmandakh 2006

†Annemys latiens Sukhanov and Narmandakh 2006

†Annemys levensis Sukhanov and Narmandakh 2006

†Annemys wusu Rabi et al. 2013

Invalid names: Annemys Sukhanov and Narmandakh 2000 [replaced], Annemys levensis Sukhanov and Narmandakh 2000 [nomen nudum]

G. †Brodiechelys Nopcsa 1928

†Brodiechelys brodiei Lydekker 1889

Invalid names: Plesiochelys valdensis Lydekker 1889 [synonym], Plesiochelys vectensis Hooley 1990 [synonym]

†Brodiechelys royoi Pérez-García et al. 2014

G. †Camerochelys Pérez-García and Murelaga 2013

†Camerochelys vilanovai Pérez-García and Murelaga 2013

G. †Kalasinemys Tong et al. 2019

†Kalasinemys prasarttongosothi Tong et al. 2019

G. †Phunoichelys Tong et al. 2014

†Phunoichelys thirakhupti Tong et al. 2014

G. †Protoxinjiangchelys Tong et al. 2012

†Protoxinjiangchelys salis Tong et al. 2012

G. †Shartegemys Sukhanov and Narmandakh 2006

†Shartegemys laticentralis Sukhanov and Narmandakh 2006

Invalid names: Shartegemys laticentralis Sukhanov and Narmandakh 2000 [nomen nudum]

G. †Tienfuchelys Young and Chow 1953

†Tienfuchelys chungkingensis Young and Chow 1953

Invalid names: Xinjiangchelys chungkingensis Young and Chow 1953 [synonym]

†Tienfuchelys tzuyangensis Young and Chow 1953

†Tienfuchelys zigongensis Peng et al. 2005

G. †Undjulemys Sukhanov and Narmandakh 2006

†Undjulemys platensis Sukhanov and Narmandakh 2006

Invalid names: Undjulemys platensis Sukhanov 1997 [nomen nudum]

G. †Xinjiangchelys Yeh 1986

†Xinjiangchelys chowi Matzke et al. 2005

†Xinjiangchelys jingyanensis Yeh and Fang 1982

†Xinjiangchelys junggarensis Yeh 1986

†Xinjiangchelys oshanensis Yeh 1973

†Xinjiangchelys qiguensis Matzke et al. 2004

†Xinjiangchelys radiplicatoides Brinkman et al. 2013

†Xinjiangchelys radiplicatus Young and Chow 1953

†Xinjiangchelys tianshanensis Nessov 1995

Invalid names: Sinemys wuerhoensis Yeh 1973 [nomen dubium], Toxocheloides naryensis Nessov and Kaznyshkin 1985 [nomen dubium]

Invalid names: Shartegemys Sukhanov and Narmandakh 2000 [nomen nudum], Undjulemys Sukhanov and Narmandakh 2000 [nomen nudum]

Invalid names: Angolachelonia Mateus et al. 2009 [empty], Centrocryptodira Gaffney and Meylan 1988 [empty], Polycryptodira Gaffney and Meylan 1988 [empty]

G. †Galvechelone Pérez-García and Murelaga 2012

†Galvechelone lopezmartinezae Pérez-García and Murelaga 2012

Fm. Kinosternoidea Hay 1892 [mud turtle]

G. Aromochelys Gray

G. Cinosternum von Spix 1824 [mud turtle]

Fm. Kinosternidae Hay 1892

G. †Baltemys Hutchison 1991

Unr. Kinosterninae Baur 1893

G. Kinosternon von Spix 1824 [mud turtle]

Invalid names: Platythyra Agassiz 1857 [synonym]

G. †Leiochelys Brinkman et al. 2023

Subfm. Staurotypinae Gray 1869

G. Staurotypus Wagler 1830 [Mexican musk turtle]

G. †Xenochelys Hay 1906

G. †Yelmochelys Brinkman et al. 2016

Unr. †Pandermatemys Joyce et al. 2004

G. †Agomphus Cope 1871

Invalid names: Amphiemys Cope 1877 [synonym]

G. †Baptemys Leidy 1870

Invalid names: Gomphochelys Bourque et al. 2015 [synonym], Notomorpha Cope 1872 [synonym]

Fm. Dermatemydidae Baur 1888

G. †Hoplochelys Hay 1908

G. Sternotherus Gray 1825 [musk turtle]

Invalid names: Aromochelys Gray [valid], Cinosternum von Spix 1824 [valid], Kinosternidae Hay 1892 [valid], Pandermatemys Joyce et al. 2004 [valid], Sternotherus Gray 1825 [valid]

Fm. †Nanhsiungchelyidae Yeh 1966

G. †Anomalochelys Hirayama et al. 2001

†Anomalochelys angulata Hirayama et al. 2001

G. †Basilemys Hay 1902

†Basilemys gaffneyi Sullivan et al. 2012

†Basilemys imbricarius Cope 1876

†Basilemys morrinensis Mallon and Brinkman 2018

†Basilemys nobilis Hay 1911

†Basilemys ogmius Cope 1875

†Basilemys praeclara Hay 1911

†Basilemys sinuosa Riggs 1906

†Basilemys variolosa Cope 1876

G. †Bulganemys Sukhanov and Narmandakh 2006

Invalid names: Bulganemys jaganchobili Sukhanov and Narmandakh 2000 [nomen nudum]

G. †Hanbogdemys Sukhanov and Narmandakh 2006

†Hanbogdemys jaganchobili Sukhanov and Narmandakh 2006

†Hanbogdemys orientalis Sukhanov and Narmandakh 1975

G. †Jiangxichelys Tong and Mo 2010

†Jiangxichelys ganzhouensis Tong and Mo 2010

†Jiangxichelys neimongolensis Brinkman and Peng 1996

G. †Kharakhutulia Sukhanov et al. 2008

†Kharakhutulia kalandadzei Sukhanov et al. 2008

G. †Nanhsiungchelys Yeh 1966

†Nanhsiungchelys wuchingensis Yeh 1966

†Nanhsiungchelys yangi Ke et al. 2023

G. †Xianyuechelys Ke et al. 2024

†Xianyuechelys yingliangi Ke et al. 2024

G. †Yuchelys Tong et al. 2012

†Yuchelys nanyangensis Tong et al. 2012

G. †Zangerlia Mlynarski 1972

†Zangerlia dzamynchondi Sukhanov and Narmandakh 2006

†Zangerlia testudinimorpha Mlynarski 1972

†Zangerlia ukhaachelys Joyce and Norell 2005

Invalid names: Zangerlia dzamynchondi Sukhanov and Narmandakh 2000 [nomen nudum]

Invalid names: Bulganemys Sukhanov and Narmandakh 2000 [nomen nudum], Charitonyx Chkhikvadze 1980 [nomen dubium], Hanbogdemysa Sukhanov 2000 [nomen nudum]

Unr. Panchelonioidea Joyce et al. 2004

G. †Mexichelys Parham and Pyenson 2010

Unr. Pancheloniidae Joyce et al. 2004

G. †Argillochelys Lydekker 1889

†Argillochelys africana Tong and Hirayama 2008

†Argillochelys antiqua König 1825

†Argillochelys brachyceps Owen 1850

†Argillochelys convexa Owen 1841

Invalid names: Chelone declivis Owen 1849 [synonym]

†Argillochelys cuneiceps Owen 1849

†Argillochelys subcristata Owen 1841

Invalid names: Chelone subcarinata Owen 1849 [synonym]

G. †Australobaena Karl and Tichy 2002

†Australobaena chilensis Karl and Tichy 2002

G. †Carolinochelys Hay 1923

†Carolinochelys wilsoni Hay 1923

Fm. †Ctenochelyidae Gentry 2018

G. †Asmodochelys Gentry et al. 2019

G. †Ctenochelys Zangerl 1953

G. †Euclastes Cope 1870

†Euclastes acutirostris Jalil et al. 2009

†Euclastes coahuilaensis Brinkman et al. 2009

†Euclastes melii Misuri 1910

†Euclastes meridionalis de la Fuente and Casadio 2000

†Euclastes platyops Cope 1870

†Euclastes wielandi Hay 1908

Invalid names: Catapleura ponderosa Cope 1871 [synonym], Erquelinnesia molaria Hay 1908 [synonym], Euclastes priscus Lynch and Parham 2003 [synonym], Osteopygis repandus Cope 1868 [synonym], Osteopygis roundsi Weems 1988 [synonym], Osteopygoides priscus Karl et al. 1998 [synonym]

Invalid names: Catapleura Cope 1870 [synonym], Glossochelys Seeley 1871 [synonym], Osteopygoides Karl et al. 1998 [synonym], Pampaemys de la Fuente and Casadio 2000 [synonym]

G. †Peritresius Cope 1870

†Peritresius martini Gentry et al. 2018

†Peritresius ornatus Leidy 1856

G. †Prionochelys Zangerl 1953

†Prionochelys matutina Zangerl 1953

Invalid names: Prionochelys galeotergum Zangerl 1953 [synonym]

†Prionochelys nauta Zangerl 1953

Invalid names: Sphenophyllum colombianum Huertas 2003 [nomen vanum]

Unr. †Pantestudinoidea Joyce et al. 2004

Superfm. Testudinoidea Gray 1825

G. †Bergouniouxchelys Hervet 2004

†Bergouniouxchelys vallisnerii Bergounioux 1954

Invalid names: Paralichelys complanata Bergounioux 1954 [synonym]

Invalid names: Anthracochelys Bergounioux 1953 [nomen nudum], Anthracochelys vallisnerii Bergounioux 1953 [nomen nudum], Cyclochelys Bergounioux 1953 [nomen nudum], Cyclochelys complanata Bergounioux 1953 [nomen nudum], Cyclochelys perexpansa Bergounioux 1953 [nomen nudum]

G. †Borkenia Schleich 1994

†Borkenia eckfeldense Karl 2018

†Borkenia germanica Hummel 1935

†Borkenia oschkinisi Schleich 1994

†Borkenia philippeni Karl 2018

G. †Cucullemys Hervet 2004

†Cucullemys crocheti Hervet 2004

G. †Cuvierichelys Hervet 2004

†Cuvierichelys crassa Owen 1849

†Cuvierichelys iberica Bergounioux 1958

Invalid names: Chrysemys elegans Bergounioux 1958 [synonym]

†Cuvierichelys parisiensis Gray 1831

Fm. Emydidae Rafinesque 1815

Subfm. Batagurinae Gray 1870

G. Callagur Schlegel and Muller 1844

G. †Chelonemys Jourdan 1862

G. Chelopus Rafinesque

Subfm. Deirochelyinae Agassiz 1857

Subfm. Emydinae Rafinesque 1815

G. Emydoidea Gray 1870 [Blanding's turtle]

G. Emys Dumeril 1806 [pond turtle]

G. †Epiemys Yeh 1963

G. Glyptemys Agassiz 1857

Invalid names: Calemys Agassiz 1857 [synonym]

G. †Grayemys Chkhikvadze 1970

G. †Gyremys Hay 1908

G. Hieremys Boulenger 1903 [Yellow-headed temple turtles]

G. †Hybemys Leidy 1871

G. †Lyssemys

G. Malacoclemmys Gray 1844

G. †Palaeotheca Cope 1872

G. †Polyechmatemys Ping 1930

G. Pseudemys Gray 1856 [cooter]

Invalid names: Nectemys Agassiz 1857 [synonym], Ptychemys Agassiz 1857 [synonym]

G. †Pseudochrysemys Sukhanov and Narmandakh 1976

G. †Pseudograptemys Hutchison 1996

G. Pyxidea Gray 1862 [Keeled box turtles]

G. †Sakya Carroll 1988

G. †Senryuemys Shikama 1953

G. †Shansiemys Yeh 1963

G. †Sharemys Gilmore 1931

G. Terrapene Merrem 1820 [box turtle]

Invalid names: Cistudo Dumeril and Bibron 1835 [synonym], Toxaspis Cope 1889 [synonym]

Invalid names: Chrysemys limnodytes Galbreath 1948 [nomen dubium], Cistudinidae Cope 1882 [synonym], Clemmys hutchensorum Bourque 2016 [nomen dubium], Clemmys saxea Hay 1903 [nomen dubium], Deirochelys floridana Hay 1908 [nomen dubium], Emydoidea hutchisoni Holman 1995 [nomen dubium], Pseudemys bisornatus Cope 1877 [nomen dubium], Pseudemys extincta Hay 1908 [nomen dubium], Pseudemys hillii Cope 1878 [nomen dubium]

G. †Euroemys Hervet 2004

†Euroemys kehreri Staesche 1928

†Euroemys vidalenci Hervet 2004

G. †Francellia Hervet 2004

†Francellia messeliana Staesche 1928

†Francellia salouagmirae Hervet 2004

Fm. Geoemydidae Theobald 1868

G. †Bridgeremys Hutchison 2006

G. †Clemmydopsis Boda 1927

G. Cuora Gray 1855 [Asian box turtle]

G. Cyclemys Bell 1834 [Asian leaf turtle]

G. †Echmatemys Hay 1906

G. †Geiselemys Khosatzky and Mlynarski 1966

G. Geoemyda Gray 1834

Subfm. Geoemydinae Theobald 1868

G. †Guangdongemys Claude et al. 2012

G. †Isometremys Chow and Yeh 1962

G. Mauremys Gray 1870 [pond turtle]

Invalid names: Paralichelys Bergounioux 1935 [synonym]

G. Ocadia Gray 1870

G. Orlitia Gray 1873

G. †Palaeochelys Meyer 1847

Unr. Palatochelydia Joyce and Lyson 2010

Subfm. †Ptychogasterinae De Stefano 1903

Tr. †Ptychogastrini De Stefano 1903

Subfm. Rhinoclemmydinae Gray 1873

G. Rhinoclemmys Fitzinger 1835

G. Sacalia Gray 1870 [eyed turtle]

G. Siebenrockiella Lindholm 1929 [black marsh turtle]

Invalid names: Bellia Gray 1869 [replaced]

Invalid names: Palaeochelyinae Karl and Wettlaufer 2011 [empty]

G. †Landreatchelys Hervet 2004

†Landreatchelys oweni Lydekker 1889

Invalid names: Clemmys vidali Bergounioux 1932 [synonym]

Fm. †Lindholmemydidae Chkhikvadze 1970

G. †Amuremys Danilov et al. 2002

G. †Elkemys Chkhikvadze 1976

G. †Gravemys Sukhanov and Narmandakh 1983

G. †Hokouchelys Yeh 1974

G. †Hongilemys Sukhanov and Narmandakh 2006

G. †Jastmelchyi Chkhikvadze 1987

G. †Khodzhakulemys Danilov 1999

G. †Lindholmemys Riabinin 1935

G. †Mongolemys Khosatzky and Mlynarski 1971

G. †Paragravemys Sukhanov et al. 1999

G. †Paramongolemys Danilov and Sukhanov 2013

G. †Shandongemys Li et al. 2013

G. †Tsaotanemys Bohlin 1953

Invalid names: Hongilemys Sukhanov 2000 [nomen nudum]

G. †Owenemys Hervet 2004

†Owenemys corroyi Bergounioux 1933

†Owenemys testudiniformis Owen 1842

Invalid names: Chrysemys bicarinata Owen and Bell 1849 [synonym]

G. †Palaeomauremys Hervet 2004

†Palaeomauremys mlynarskii Hervet and de Broin 2000

†Palaeomauremys tuberculata Portis 1882

Unr. †Panemydidae Joyce et al. 2004

G. †Acherontemys Hay 1899

G. †Psilosemys Hutchison 2013

Unr. †Pangeoemydidae Joyce 2007

G. †Banhxeochelys Garbin et al. 2019

Unr. †Pantestudinidae Joyce et al. 2004

G. †Cymatholcus Clark 1932

G. †Hadrianus Cope 1872

G. †Oligopherus Hutchison 1996

G. †Promalacoclemmys Reinach 1900

†Promalacoclemmys laharpi Pictet and Humbert 1861

Invalid names: Emys charpentieri Pictet and Humbert 1861 [synonym], Emys renevieri Portis 1882 [synonym], Emys sulcata Portis 1882 [synonym]

†Promalacoclemmys protogaea Meyer 1852

Invalid names: Emys gessneri Meyer 1851 [synonym], Ocadia hessleriana Reinach 1900 [synonym], Promalacoclemmys boulengeri Reinach 1900 [synonym]

G. †Provencemys Hervet 2004

†Provencemys lachati Sauvage 1873

Invalid names: Clemmys manuascensis Bergounioux 1934 [synonym]

†Provencemys laurenti Bergounioux 1934

Fm. Testudinidae Batsch 1788

G. †Achilemys Hay 1908

G. Aldabrachelys Loveridge and Williams 1957

Invalid names: Dipsochelys Bour 1982 [synonym]

G. †Anhuichelys Yeh 1979

G. Astrochelys Gray 1873

G. †Barnesia Karl 2018

G. Cachuga Gray 1855

G. †Cautleya Lydekker 1885

G. †Cylindraspis Fitzinger 1835

G. Damonia Gray 1869

G. †Eochersina Karl 2018

G. †Eochersis Karl 2018

G. †Floridemys Williams 1950

Invalid names: Bystra Hay 1916 [replaced]

G. †Fontainechelon Pérez-García et al. 2016

Tr. Geochelonini Swingland and Klemens 1989

Subfm. Gopherinae Swingland and Klemens 1989

G. Gopherus Rafinesque 1832 [gopher tortoise]

Invalid names: Bysmachelys Johnston 1937 [synonym], Scaptochelys Bramble 1982 [synonym], Xerobates Agassiz 1857 [synonym]

G. Homopus Duméril and Bibron 1834 [cape tortoise]

G. †Juvemys Hervet 2003

G. †Kansuchelys Yeh 1963

G. Malacochersus Lindholm 1929

G. Manouria Gray 1852

G. †Megalochelys Falconer and Cautley 1837

G. †Miotestudo Deraniyagala 1967

G. †Nicoria Gray 1855

G. †Palaeoemys Schleich 1994

G. †Pelorochelon Pérez-García et al. 2016

Subfm. Platysterninae Gray 1869

G. †Protestudo Chkhikvadze 1970

G. Psammobates Fitzinger 1835

G. Pyxis Bell 1827

G. †Sinohadrianus Ping 1929

G. †Solitudo Valenti et al. 2022

Subfm. Testudininae Gray 1825

Subfm. †Xerobatinae Agassiz 1857

Invalid names: Caryoderma Cope 1886 [nomen dubium], Chelys patagonica Wieland 1923 [nomen dubium], Emys sombrerensis Leidy 1868 [nomen dubium], Geochelone (Hesperotestudo) equicomes Hay 1917 [nomen dubium], Geocheloninae Swingland and Klemens 1989 [empty], Gopherus pertenius Cope 1892 [nomen dubium], Gopherus praecedens Hay 1916 [nomen dubium], Stylemys calaverensis Sinclair 1903 [nomen dubium], Testudininei de Broin and de la Fuente 1993 [empty], Testudinites sellovii Weiss 1830 [nomen dubium], Testudo annae Hay 1923 [nomen dubium], Testudo atascosae Hay 1902 [nomen dubium], Testudo castrensis Bergounioux 1935 [nomen dubium], Testudo cultrata Cope 1873 [nomen dubium], Testudo exornata Lambe 1906 [nomen dubium], Testudo formosa Moreno 1889 [nomen dubium], Testudo francisi Hay 1923 [nomen dubium], Testudo hayi Sellards 1916 [nomen dubium], Testudo klettiana Cope 1875 [nomen dubium], Testudo laticaudata Cope 1893 [nomen dubium], Testudo louisekressmani Wark 1929 [nomen dubium], Testudo luciae Hay 1916 [nomen dubium], Testudo ocalana Hay 1916 [nomen dubium], Testudo paranensis Scalabrini 1887 [nomen nudum], Testudo peragrans Hay 1906 [nomen dubium], Testudo quadratus Cope 1884 [nomen dubium], Testudo semenensis Bergounioux 1955 [nomen dubium], Testudo undata Cope 1875 [nomen dubium]

G. †Wutuchelys Tong et al. 2017

†Wutuchelys eocenica Tong et al. 2017

Invalid names: Chrysemys (Pseudemys) hibbardi Preston 1979 [nomen dubium], Clemmys hesperia Hay 1903 [nomen dubium], Clemmys percrassus Cope 1899 [nomen dubium], Emys euglypha Leidy 1889 [nomen dubium], Emys euthnetus Cope 1873 [nomen dubium], Emys latilabiatus Cope 1871 [nomen dubium], Emys megaulax Cope 1873 [nomen dubium], Emys pachylomus Cope 1873 [nomen dubium], Emys petrolei Leidy 1868 [nomen dubium], Eupachemys obtusus Leidy 1877 [nomen dubium], Hadrianus allabiatus Cope 1871 [nomen dubium], Hybemys arenarius Leidy 1871 [nomen dubium], Notomorpha testudinea Cope 1872 [nomen dubium], Rhinoclemmys nicoyana Acuña-Mesen and Laurito-Mora 1996 [nomen dubium], Testudo culbratus Cope 1873 [nomen dubium], Testudo distans Hay 1916 [nomen dubium], Testudo isis Andrews 1906 [nomen dubium], Testudo munda Hay 1920 [nomen dubium], Trachemys delicata Hay 1916 [nomen dubium], Trachemys nuchocarinata Hay 1916 [nomen dubium]

G. †Polysternum Bergounioux 1935

Unr. †Selmacryptodira Gaffney et al.

Invalid names: Kallokibotionidae Gaffney and Meylan 1992 [empty]

Subor. Trionychia

Unr. Carettochelyidae Gill 1889

G. †Akrochelys Bergounioux 1935

†Akrochelys neuvillei Bergounioux 1935

G. †Anosteira Leidy 1871

†Anosteira crowcreekensis Hutchison and Westgate 2024

†Anosteira manchuriana Zangerl 1947

†Anosteira maomingensis Chow and Liu 1955

†Anosteira mongoliensis Gilmore 1931

Invalid names: Anosteira shantungensis Cheng 1961 [synonym]

†Anosteira ornata Leidy 1871

†Anosteira pulchra Clark 1932

†Anosteira radulina Cope 1872

†Anosteira shuvalovi Chkhikvadze 1979

Subfm. †Anosteirinae Lydekker 1889

G. †Kyzilkumemys Nesov 1977

G. †Pseudanosteira Clark 1932

G. †Pseudotrionyx Dollo 1886

Unr. †Carettochelyinae Boulenger 1887

Tr. †Allaeochelyini Godinot et al. 2018

G. †Burmemys Hutchison et al. 2004

G. †Hemichelys Lydekker 1887

G. Carettochelys Ramsay 1886 [pig-nose turtle]

Carettochelys insculpta Ramsay 1886 [Pignose turtle]

†Carettochelys niahensis White et al. 2023

G. †Kizylkumemys Nessov 1977

†Kizylkumemys khoratensis Tong et al. 2005

†Kizylkumemys schultzi Nessov 1977

G. †Pseudanosteira Clark 1932

Invalid names: Allaeochelyinae Bergounioux 1952 [empty], Anosteira radulina Cope 1872 [nomen dubium], Apholidemys granosa Pomel 1847 [nomen nudum], Apholidemys sublaevis Pomel 1847 [nomen nudum], Chorlakkichelys de Broin 1987 [nomen dubium], Chorlakkichelys shahi de Broin 1987 [nomen dubium], Kizylkumemys schultzi mirabilis Nessov 1981 [nomen dubium]

G. †Kappachelys Hirayama et al. 2013

†Kappachelys okurai Hirayama et al. 2013

Unr. †Pantrionychidae Joyce et al. 2004

G. †Drazinderetes Head et al. 1999

†Drazinderetes tethyensis Head et al. 1999

G. †Khunnuchelys Brinkman et al. 1993

†Khunnuchelys erinhotensis Brinkman et al. 1993

†Khunnuchelys kizylkumensis Brinkman et al. 1993

†Khunnuchelys lophorhothon Danilov et al. 2015

G. †Murgonemys White 2001

†Murgonemys braithwaitei White 2001

Unr. †Plastomenidae Hay 1902

G. †Aspideretoides Gardner et al. 1995

G. †Atoposemys Hutchison 2013

G. †Gilmoremys Joyce and Lyson 2011

G. †Jimemys Edgar et al. 2022

Unr. †Plastomenini Jasinski et al. 2022

Invalid names: Amyda egregia Hay 1908 [nomen dubium], Amyda salebrosa Hay 1908 [nomen dubium], Anostira molopina Cope 1872 [nomen dubium], Anostira oedemia Cope 1872 [nomen dubium], Anostira trionychoides Cope 1872 [nomen dubium], Aspideretes amnigenus Hay 1910 [nomen dubium], Aspideretes perplexus Gilmore 1919 [nomen dubium], Plastomenus catenatus Cope 1875 [nomen dubium], Plastomenus coalescens Cope 1875 [nomen dubium], Plastomenus communis Cope 1875 [nomen dubium], Plastomenus corrugatus Cope 1875 [nomen dubium], Plastomenus costatus Cope 1875 [nomen dubium], Plastomenus fractus Cope 1875 [nomen dubium], Plastomenus insignis Cope 1874 [nomen dubium], Plastomenus lachrymalis Cope 1875 [nomen dubium], Plastomenus multifoveatus Cope 1873 [nomen dubium], Plastomenus punctulatus Cope 1874 [nomen dubium], Plastomenus serialis Cope 1877 [nomen dubium], Plastomenus torrejonensis Gilmore 1919 [nomen dubium], Platypeltis amnicola Hay 1907 [nomen dubium], Platypeltis antiqua Hay 1907 [nomen dubium], Platypeltis trepida Hay 1907 [nomen dubium], Procyclanorbis sardus Portis 1901 [nomen dubium], Trionyx buiei Cope 1870 [nomen dubium], Trionyx cariosus Cope 1875 [nomen dubium], Trionyx cellulosus Cope 1868 [nomen dubium], Trionyx concentricus Cope 1872 [nomen dubium], Trionyx guttatus Leidy 1869 [nomen dubium], Trionyx halophilus Cope 1869 [nomen dubium], Trionyx heteroglyptus Cope 1873 [nomen dubium], Trionyx leptomitus Cope 1875 [nomen dubium], Trionyx pennatus Cope 1869 [nomen dubium], Trionyx priscus Cope 1869 [nomen dubium], Trionyx punctiger Cope 1891 [nomen dubium], Trionyx quinni Holman 1982 [nomen dubium], Trionyx radulus Cope 1875 [nomen dubium], Trionyx vagans Cope 1874 [nomen dubium], Trionyx ventricosus Cope 1877 [nomen dubium], Trionyx virginiana Clark 1895 [nomen dubium]

Fm. Trionychidae Gray 1825 [softshell turtle]

G. Apalone Rafinesque 1832 [softshell turtle]

Subg. Apalone (Platypeltis) Fitzinger 1835

Platypeltis ferox Schneider 1783 [Florida softshell turtle]

†Platypeltis sternbergi Wiman 1933

†Apalone amorense Valdes et al. 2017

Apalone ferox Schneider 1783 [Florida softshell turtle]

†Apalone lima Cope 1869

Apalone mutica LeSueur 1827 [smooth softshell turtle]

Apalone spinifera LeSueur 1827 [spiny softshell turtle]

†Apalone trionychoides Cope 1872

G. Aspideretes Hay 1904

Aspideretes gangeticus Cuvier 1825 [Indian softshell]

Aspideretes hurum Gray 1831 [Indian peacock softshell]

Aspideretes leithii Gray 1872 [Leith's softshell]

Aspideretes nigricans Anderson 1875 [Black softshell]

†Aspideretes sinuosus Chow and Yeh 1958

†Trionyx (Aspideretes) fuchienensis Yeh 1974

G. Aspidonectes Wagler 1830

Subfm. †Chitrainae Jasinski et al. 2022

G. Chitra Gray 1844

Chitra chitra Nutaphand 1986

Invalid names: Chitra selenkae Jaekel 1911 [replaced]

Chitra indica Gray 1831

Invalid names: Trionyx indicus Gray 1831 [synonym]

Chitra vandijki McCord and Pritchard 2003 [Burmese narrow-headed softshell turtle]

G. Pelochelys Gray 1864

Pelochelys bibroni Owen 1853 [Asian giant softshell]

Pelochelys cantorii Gray 1864

Pelochelys signifera Webb 2003

G. †Conchochelys Hay 1905

G. †Crassithecachelys Chkhikvadze 2000

Subfm. Cyclanorbinae Hummel 1929

G. Cyclanorbis Gray 1854

Cyclanorbis elegans Gray 1869

Cyclanorbis senegalensis Dumeril and Bibron 1835

†Cyclanorbis turkanensis Meylan et al. 1990

G. Cycloderma Peters 1854

Cycloderma aubryi Joyce and Lyson 2010

†Cycloderma debroinae Meylan et al. 1990

Cycloderma frenatum Peters 1854

†Cycloderma victoriae Andrews 1914

G. Lissemys Smith 1931

Lissemys ceylonensis Gray 1856 [Sri Lankan flapshell turtle]

Lissemys punctata Bonnaterre 1789

Lissemys scutata Peters 1868 [Burmese flapshell turtle]

Invalid names: Emyda lineata Lydekker 1885 [nomen dubium], Emyda palaeindica Lydekker 1885 [nomen dubium], Emyda sivalensis Lydekker 1885 [nomen dubium], Lissemys piramensis Prasad 1974 [nomen dubium], Lissemys punctata sinhaleyus Deraniyagala 1953 [nomen dubium]

G. †Nemegtemys Danilov et al. 2014

†Nemegtemys conflata Danilov et al. 2014

G. †Eugenechelys Chkhikvadze 2000

G. †Hummelichelys Chkhikvadze 2000

Subfm. †Kuhnemydinae Jasinski et al. 2022

G. †Kuhnemys Chkhikvadze 1999

†Kuhnemys breviplastra Danilov et al. 2014

†Kuhnemys maortuensis Yeh 1965

Invalid names: Aspideretes alashanensis Yeh 1965 [synonym], Trionyx alashanensis Yeh 1965 [synonym]

†Kuhnemys orlovi Khosatzky 1976

†Kuhnemys palaeocenica Danilov et al. 2015

G. †Perochelys Li et al. 2015

†Perochelys hengshanensis Brinkman et al. 2017

†Perochelys lamadongensis Li et al. 2015

G. †Petrochelys Vitek et al. 2017

†Petrochelys kyrgyzensis Nessov 1995

G. †Paleotrionyx Schmidt 1945

G. †Striatochelys Massonne et al. 2023

†Striatochelys baba Massonne et al. 2023

†Striatochelys impressa Yeh 1963

G. †Temnotrionyx Hay 1908

Subfm. Trionychinae Gray 1825 [softshell turtle]

G. †Aulacochelys Lydekker 1889

G. †Axestemys Hay 1899

†Axestemys byssina Cope 1872

†Axestemys byssinus Cope 1872

Invalid names: Eugenechelys robertemryi Chkhikvadze 2008 [synonym], Temnotrionyx manducans Hay 1908 [synonym]

†Axestemys cerevisia Vitek 2012

†Axestemys erquelinnensis Perez-Garcia and Smith 2021

†Axestemys infernalis Joyce et al. 2019

†Axestemys montinsana Vitek 2012

†Axestemys quinni Schmidt 1945

†Axestemys splendidus Hay 1908

Invalid names: Aspideretes granifer Hay 1909 [synonym], Aspideretes planus Parks 1933 [synonym]

†Axestemys vittata Pomel 1847

Invalid names: Eurycephalochelys fowleri Moody and Walker 1970 [synonym], Trionyx levalensis Dollo 1909 [synonym]

†Rafetus (Axestemys) exquisitus Hay 1908

†Rafetus (Axestemys) nelsoni Cope 1972

Invalid names: Axestus Cope 1872 [synonym], Eurycephalochelys Moody and Walker 1970 [synonym]

Tr. Chitrini Gray 1870

G. Amyda Geoffroy Saint-Hilaire 1809

G. †Palaeoamyda Cadena 2016

G. Dogania Gray 1844

†Dogania maortuensis Yeh 1965

Dogania subplana Geoffrey Saint-Hilaire 1809 [Malayan softshell]

G. Nilssonia Gray 1869 [Burmese peacock softshells]

Nilssonia formosa Gray 1869 [Burmese peacock softshell]

Nilssonia gangetica Cuvier 1825 [Indian softshell turtle]

Nilssonia hurum Gray 1830

Invalid names: Trionyx hurum Gray 1831 [synonym], Trionyx hurum sivalensis Lydekker 1889 [synonym]

Nilssonia leithii Gray 1872 [Leiths softshell turtle]

Nilssonia nigricans Anderson 1875 [Black softshell turtle]

G. Palea Siebenrock 1906 [Wattleneck softshells]

Palea steindachneri Siebenrock 1906 [Wattleneck softshell]

G. Pelodiscus Fitzinger 1835

Pelodiscus axenaria Zhou et al. 1991 [Hunan softshell turtle]

†Pelodiscus gracilia Yeh 1963

Pelodiscus maackii Brandt 1857 [Northern Chinese softshell turtle]

Pelodiscus parviformis Tang 1997 [Lesser Chinese softshell turtle]

Pelodiscus sinensis Wiegmann 1835 [Chinese softshell]

Invalid names: Trionyx sinuosus Chow and Yeh 1958 [nomen dubium]

G. Rafetus Gray 1864

†Rafetus bohemicus Liebus 1930

Rafetus euphraticus Daudin 1802 [Euphrates softshell]

Rafetus swinhoei Gray 1873

Invalid names: Pelochelys taihuensis Zhang 1984 [synonym], Trionyx liupani Tao 1986 [synonym]

Tr. Trionychini Gray 1825

Subtr. Apalonina Meylan 1987 [softshell turtle]

G. †Gobiapalone Danilov et al. 2014

G. †Oliveremys Vitek 2011

G. †Rafetoides Karl 1998

Invalid names: Rafetoidina Karl 1998 [empty], Trionychina Gray 1825 [empty]

G. Trionyx Geoffroy Saint-Hilaire 1809

†Trionyx admirabilis Hay 1905

†Trionyx aequa Hay 1908

Invalid names: Amyda crassa Hay 1908 [synonym]

†Trionyx allani Gilmore 1923

†Trionyx austerus Hay 1908

Invalid names: Aspideretes fontanus Hay 1908 [synonym], Aspideretes vorax Hay 1908 [synonym]

†Trionyx baynshirensis Danilov et al. 2014

†Trionyx beecheri Hay 1904

†Trionyx boulengeri Reinach 1900

Invalid names: Trionyx clavatomarginatus Lörenthey 1903 [synonym]

†Trionyx capellinii Negri 1892

Invalid names: Trionyx affinis Negri 1892 [synonym], Trionyx capellinii conjugens Sacco 1894 [synonym], Trionyx capellinii gracilina Sacco 1895 [synonym], Trionyx capellinii montevialensis Negri 1892 [synonym], Trionyx capellinii perexpansa Sacco 1895 [synonym], Trionyx gemmellaroi Negri 1892 [synonym], Trionyx insolitus Bergounioux 1954 [synonym], Trionyx intermedius Bergounioux 1954 [synonym], Trionyx schaurothianus Negri 1893 [synonym]

†Trionyx dissolutus Vitek and Danilov 2014

†Trionyx ellipticus Hay 1908

Invalid names: Aspideretes annae Zangerl 1944 [synonym], Aspideretes grangeri Hay 1908 [synonym]

†Trionyx eloisae Gilmore 1919

†Trionyx foveatus Leidy 1856

Invalid names: Aspideretes maturus Lambe 1914 [synonym], Aspideretes rugosus Parks 1933 [synonym], Aspideretes subquadratus Lambe 1914 [synonym]

Trionyx gangeticus Cuvier 1824

†Trionyx gilbentuensis Danilov et al. 2014

†Trionyx gobiensis Danilov et al. 2014

†Trionyx gregarius Gilmore 1934

Invalid names: Rafetus gilmorei Chkhikvadze 1999 [synonym]

†Trionyx henrici Owen and Bell 1849

Invalid names: Trionyx barbarae Owen and Bell 1849 [synonym], Trionyx circumsulcatus Owen and Bell 1849 [synonym], Trionyx incrassatus Owen and Bell 1849 [synonym], Trionyx marginatus Owen and Bell 1849 [synonym], Trionyx planus Owen and Bell 1849 [synonym], Trionyx rivosus Owen and Bell 1849 [synonym]

†Trionyx ikoviensis Danilov et al. 2011

†Trionyx jixiensis Li et al. 2015

†Trionyx johnsoni Gilmore 1931

Invalid names: Amyda neimenguensis Yeh 1965 [synonym]

†Trionyx kansaiensis Vitek and Danilov 2010

†Trionyx latus Gilmore 1919

†Trionyx leucopotamicus Cope 1891

†Trionyx linchuensis Yeh 1962

†Trionyx miensis Okazaki and Yoshida 1977

†Trionyx minusculus Chkhikvadze 1973

†Trionyx miocaenus Matthew 1924

†Trionyx mira Hay 1908

Invalid names: Platypeltis extensa Hay 1908 [synonym], Platypeltis postera Hay 1908 [synonym]

†Trionyx ninae Chkhikvadze 1971

Invalid names: Trionyx turgaicus Kuznetsov and Chkhikvadze 1977 [synonym], Trionyx zaisanensis Chkhikvadze 1973 [synonym]

†Trionyx onomatoplokos Kuznetsov and Chkhikvadze 1987

Trionyx phayrei Theobald 1868

†Trionyx pliocenicus Fucini 1912

†Trionyx reesidei Gilmore 1919

Invalid names: Aspideretes quadratus Gilmore 1919 [synonym]

†Trionyx riabinini Kuznetsov and Chkhikvadze 1987

†Trionyx robustus Gilmore 1919

Invalid names: Aspideretes ovatus Gilmore 1935 [synonym]

†Trionyx shiluutulensis Danilov et al. 2014

†Trionyx silvestris Walker and Moody 1974

Invalid names: Trionyx michauxi Broin 1977 [synonym]

†Trionyx singularis Hay 1907

Invalid names: Aspideretes puercensis Hay 1908 [synonym], Aspideretes sagatus Hay 1908 [synonym]

Trionyx triunguis Forksal 1775 [African softshell turtle]

†Trionyx uintaensis Cope 1873

Invalid names: Amyda franciscae Hay 1908 [synonym], Amyda nelsoni Case 1927 [synonym], Aspidonectes tritor Hay 1904 [synonym]

†Trionyx vindobonensis Peters 1855

Invalid names: Trionyx brunhuberi Ammon 1911 [synonym], Trionyx gergensi Reinach 1900 [synonym], Trionyx hoernesi Heritsch 1909 [synonym], Trionyx peneckei Heritsch 1909 [synonym], Trionyx petersi Hoernes 1881 [synonym], Trionyx rostratus Arthaber 1898 [synonym], Trionyx septemcostatus Hoernes 1881 [synonym], Trionyx teiritzbergensis Gemel 2002 [synonym], Trionyx teyleri Winkler 1869 [synonym]

Invalid names: Francedebroinella Chkhikvadze 1999 [synonym], Sinamyda Chkhikvadze 2000 [synonym], Trionyx nopcsai Szalai 1934 [nomen dubium]

Invalid names: Trionyx (Amyda) subangularis Bergounioux 1954 [nomen dubium], Trionyx bambolii Ristori 1891 [nomen dubium], Trionyx blayaci Bergounioux 1933 [nomen dubium], Trionyx desmostyli Matsumoto 1918 [nomen dubium], Trionyx dieupentalensis Bergounioux 1935 [nomen dubium], Trionyx harmati Szalai 1934 [nomen dubium], Trionyx irregularis Bergounioux 1954 [nomen dubium], Trionyx pliopedemontana Sacco 1889 [nomen dubium], Trionyx stadleri Teppner 1913 [nomen dubium], Trionyx stiriacus Peters 1855 [nomen dubium]

G. †Zangerlichelys Chkhikvadze 2000

Invalid names: Altaytrionyx Chkhikvadze 2008 [nomen dubium], Altaytrionyx burtschaki Chkhikvadze 2008 [nomen dubium], Altaytrionyx devjatkini Chkhikvadze 2008 [nomen dubium], Altaytrionyx phiruzae Chkhikvadze 2008 [nomen dubium], Amyda menneri Chkhikvadze 1988 [nomen dubium], Aspideretes jaxarticus Riabinin 1938 [nomen nudum], Aspideretes muyuensis Lei and Ye 1985 [nomen dubium], Aspilus Portis 1885 [nomen dubium], Aspilus cortesii Portis 1885 [nomen dubium], Chitra minor Jaekel 1911 [nomen dubium], Lissemydinae Williams 1950 [empty], Paleotrionyx jimenezfuentesi Chkhikvadze 1995 [nomen dubium], Plastomenus gabunii Chkhikvadze 1984 [nomen dubium], Plastomenus jaxarticus Riabinin 1938 [nomen nudum], Plastomenus mlynarskii Chkhikvadze 1970 [nomen dubium], Platypeltis subcircularis Chow and Yeh 1957 [nomen dubium], Procyclanorbis Portis 1901 [nomen dubium], Rafetini Chkhikvadze 2000 [empty], Rafetus karkhualexandri Chkhikvadze 2000 [nomen nudum], Rafetus yexiangkuii Chkhikvadze 1999 [nomen dubium], Trionyx acutiformis Bergounioux 1935 [nomen dubium], Trionyx amansii Gray 1831 [nomen dubium], Trionyx anthracotheriorum Portis 1883 [nomen dubium], Trionyx aquitanicus Delfortrie 1869 [nomen dubium], Trionyx aspidiformis Laube 1900 [nomen dubium], Trionyx australiensis De Vis 1894 [nomen dubium], Trionyx austriacus Peters 1859 [nomen dubium], Trionyx borkenensis Gramann 1956 [nomen dubium], Trionyx bowerbanki Lydekker 1889 [nomen dubium], Trionyx bruxelliensis Winkler 1869 [nomen dubium], Trionyx burdigalensis Bergounioux 1935 [nomen dubium], Trionyx capellinii bulgaricus Khosatzky et al. 1983 [nomen dubium], Trionyx chaubeti Bergounioux 1935 [nomen dubium], Trionyx ciryi Bergounioux 1935 [nomen dubium], Trionyx cliftii Fitzinger 1835 [nomen nudum], Trionyx croaticus Koch 1915 [nomen dubium], Trionyx danovi Chkhikvadze 1989 [nomen dubium], Trionyx dodunii Gray 1831 [nomen nudum], Trionyx elongatus Liebus 1930 [nomen dubium], Trionyx erquelinnensis Dollo 1909 [nomen nudum], Trionyx hilberi Hoernes 1892 [nomen dubium], Trionyx ishiharaensis Miura and Uyama 1987 [nomen dubium], Trionyx italicus Schauroth 1865 [nomen dubium], Trionyx jakhimovitchae Chkhikvadze 1989 [nomen dubium], Trionyx kazusensis Otsuka 1969 [nomen dubium], Trionyx khosatzkyi Chkhikvadze 1983 [nomen dubium], Trionyx laurillardii Gray 1831 [nomen dubium], Trionyx lockardi Gray 1831 [nomen nudum], Trionyx lorioli Portis 1882 [nomen dubium], Trionyx manouri Gray 1831 [nomen dubium], Trionyx marini Hernández Sampelayo and Bataller 1944 [nomen dubium], Trionyx moldaviensis Khosatzky 1986 [nomen dubium], Trionyx mourieri Bergounioux 1935 [nomen dubium], Trionyx munzenbergensis Hummel 1927 [nomen nudum], Trionyx oligocenica Negri 1892 [nomen nudum], Trionyx oweni Reinach 1900 [nomen dubium], Trionyx parisiensis Gray 1831 [nomen dubium], Trionyx partschii Peters 1855 [nomen dubium], Trionyx pedemontana Portis 1879 [nomen dubium], Trionyx petersi trifailensis Teppner 1914 [nomen dubium], Trionyx pliocaenicus Reinach 1903 [nomen dubium], Trionyx pompignanensis Bergounioux 1933 [nomen dubium], Trionyx pontanus Laube 1895 [nomen dubium], Trionyx portisi Ristori 1891 [nomen dubium], Trionyx preschenensis Laube 1900 [nomen dubium], Trionyx propinquus Ristori 1891 [nomen dubium], Trionyx pseudovindobonensis Szalai 1934 [nomen dubium], Trionyx pustulatus Owen and Bell 1849 [nomen dubium], Trionyx ragusensis De Gregorio 1883 [nomen dubium], Trionyx reticulatus Rieppel 1979 [nomen dubium], Trionyx rocchettiana Portis 1882 [nomen dubium], Trionyx roncensis Harrassowitz 1919 [nomen dubium], Trionyx rotundiformis Bergounioux 1933 [nomen dubium], Trionyx sculptus Gilmore 1931 [nomen dubium], Trionyx senckenbergianus Reinach 1903 [nomen dubium], Trionyx senensis Ristori 1891 [nomen dubium], Trionyx siegeri Heritsch 1909 [nomen dubium], Trionyx sophiae Heritsch 1909 [nomen dubium], Trionyx stormsi Delheid 1899 [nomen dubium], Trionyx ubeensis Chitani 1925 [nomen dubium], Trionyx valdensis Portis 1882 [nomen dubium], Trionyx zakhidovi Khosatzky 1966 [nomen dubium], Zaisanonyx Chkhikvadze 2008 [nomen dubium]

Superfm. Trionychoidea Fitzinger 1826

Fm. †Adocidae Cope 1869

G. †Adocoides Sukhanov and Narmandakh 2006

G. †Adocus Cope 1868

†Adocus agilis Cope 1868

†Adocus aksary Nessov and Krasovskaya 1984

†Adocus amtgai Narmandakh 1985

†Adocus beatus Leidy 1865

Invalid names: Adocus lacer Hay 1908 [synonym], Adocus punctatus Marsh 1890 [synonym]

†Adocus bossi Gilmore 1919

†Adocus bostobensis Syromyatnikova and Danilov 2009

†Adocus dzhurtasensis Syromyatnikova and Danilov 2009

†Adocus firmus Leidy 1856

†Adocus foveatus Nesov and Khozarskii 1977

†Adocus hesperius Gilmore 1919

†Adocus inexpectatus Danilov et al. 2013

†Adocus kazachstanica Chkhikvadze 1973

†Adocus kirtlandius Gilmore 1919

†Adocus kizylkumensis Nesov 1981

†Adocus kokaku Hirayama et al. 2021

†Adocus lineolatus Cope 1874

†Adocus microglypha Cope 1871

†Adocus onerosus Gilmore 1919

†Adocus orientalis Gilmore 1931

†Adocus planus Sukhanov and Narmandakh 2006

†Adocus pravus Leidy 1856

†Adocus sengokuensis Sonoda et al. 2015

†Adocus striatula Cope 1869

†Adocus syntheticus Cope 1869

†Adocus vigoratus Hay 1910

Invalid names: Alamosemys Hay 1908 [synonym], Homorophus Cope 1869 [synonym], Shineusemys Sukhanov and Narmandakh 2006 [synonym], Zygoramma Cope 1869 [synonym]

G. †Ferganemys Nessov and Yulinen 1977

†Ferganemys itemirensis Nesov 1981

†Ferganemys verzilini Nesov and Khozatskii 1977

G. †Isanemys Tong et al. 2006

†Isanemys srisuki Tong et al. 2006

Subfm. †Shachemydinae Khosatzky 1977

G. †Protoshachemys Tong et al. 2019

Invalid names: Mlynarskiella Shuvalov and Chkhikvadze 1986 [nomen dubium], Mlynarskiella mariani Shuvalov and Ckhikvadze 1986 [nomen dubium]

G. †Shachemys Kuznetsov 1976

†Shachemys ancestralis Nessov 1984

†Shachemys baibolatica Kuznetsov 1976

†Shachemys laosiana de Lapparent de Broin 2004

G. †Shineusemys Sukhanov and Narmandakh 2006

†Sineusemys plana Sukhanov and Narmandakh 2000

Invalid names: Sineusemys plana Sukhanov and Narmandakh 2000 [valid]

Invalid names: Adocinae Cope 1869 [empty], Adocoides Narmandakh 1985 [nomen nudum], Adocus askari Nesov and Krasovskaya 1984 [nomen dubium]

G. †Basilochelys Tong et al. 2009

†Basilochelys macrobios Tong et al. 2009

Fm. Kinosternoidea Hay 1892 [mud turtle]

G. Aromochelys Gray

G. Cinosternum von Spix 1824 [mud turtle]

Fm. Kinosternidae Hay 1892

G. †Baltemys Hutchison 1991

Unr. Kinosterninae Baur 1893

G. Kinosternon von Spix 1824 [mud turtle]

Invalid names: Platythyra Agassiz 1857 [synonym]

G. †Leiochelys Brinkman et al. 2023

Subfm. Staurotypinae Gray 1869

G. Staurotypus Wagler 1830 [Mexican musk turtle]

G. †Xenochelys Hay 1906

G. †Yelmochelys Brinkman et al. 2016

Unr. †Pandermatemys Joyce et al. 2004

G. †Agomphus Cope 1871

Invalid names: Amphiemys Cope 1877 [synonym]

G. †Baptemys Leidy 1870

Invalid names: Gomphochelys Bourque et al. 2015 [synonym], Notomorpha Cope 1872 [synonym]

Fm. Dermatemydidae Baur 1888

G. †Hoplochelys Hay 1908

G. Sternotherus Gray 1825 [musk turtle]

Invalid names: Aromochelys Gray [valid], Cinosternum von Spix 1824 [valid], Kinosternidae Hay 1892 [valid], Pandermatemys Joyce et al. 2004 [valid], Sternotherus Gray 1825 [valid]

G. †Sinaspideretes Young and Chow 1953

†Sinaspideretes wimani Young and Chow 1953

Invalid names: Yehguia Danilov and Parham 2006 [synonym]

Invalid names: Peltochelyidae Seeley 1880 [empty], Planetochelyidae Hutchison 2013 [empty]

Invalid names: Adocusia Danilov and Parham 2006 [empty], Carettochelyoidea Romer 1956 [empty], Charitonyx tajanikolaevae Chkhikvadze 1980 [nomen dubium], Chersidae Zittel 1890 [empty], Cryptoderinea Vaillant 1894 [empty], Dermatochelydidae [empty], Glyptopidae [empty], Glyptopsidae Marsh 1890 [empty], Indochelyidae Datta et al. 2000 [empty], Kallokibotioninae Nopsca 1923 [empty], Kayentachelyidae Gaffney et al. 1987 [empty], Kinosternia Gaffney and Meylan 1988 [empty], Kinosternoidae Holroyd and Hutchison 2002 [empty], Meiolaniidae Lydekker 1887 [empty], Pantrionychia Joyce et al. 2004 [empty], Platysternoidea Baur 1893 [empty], Tritosternidae Lydekker 1889 [empty]

G. †Entoplastrites Huene 1956

†Entoplastrites nuertingensis Huene 1956

Fm. †Haichemydidae Sukhanov and Narmandakh 2006

G. †Haichemys Sukhanov and Narmandakh 2006

†Haichemys ulensis Sukhanov and Narmandakh 2006

G. †Larachelus Pérez-García and Murelaga 2012

†Larachelus morla Pérez-García and Murelaga 2012

G. †Odontochelys Li et al. 2008

†Odontochelys semitestacea Li et al. 2008

Unr. Pancryptodira Joyce et al. 2004

Unr. †Macrobaenidae Sukhanov 1964

G. †Anatolemys Khozatsky and Nessov 1979

†Anatolemys maximus Khozatskii and Nessov 1979

†Anatolemys oxensis Khozatskii and Nessov 1977

G. †Appalachemys Gentry et al. 2023

†Appalachemys ebersolei Gentry et al. 2023

G. †Asiachelys Sukhanov and Narmandakh 2006

Invalid names: Asiachelys perforata Sukhanov and Narmandakh 2000 [nomen nudum]

G. †Aurorachelys Vandermark et al. 2009

†Aurorachelys gaffneyi Vandermark et al. 2009

G. †Changmachelys Brinkman et al. 2013

†Changmachelys bohlini Brinkman et al. 2013

G. †Gallica Pérez-García 2020

†Gallica lapparentiana Pérez-García 2020

G. †Hangaiemys Sukhanov and Narmandakh 1974

†Hangaiemys acutus Bohlin 1953

†Hangaiemys hoburensis Sukhanov and Narmandakh 1974

†Hangaiemys kansuensis Bohlin 1953

†Hangaiemys latilimbata Bohlin 1953

†Hangaiemys leptis Sukhanov and Narmandakh 2006

Invalid names: Hangaiemys leptis Sukhanov and Narmandakh 2000 [nomen nudum]

G. †Judithemys Parham and Hutchison 2003

†Judithemys sukhanovi Parham and Hutchison 2003

G. †Kirgizemys Nessov and Khozatskiy 1973

†Kirgizemys dmitrievi Nessov and Khozatskii 1981

†Kirgizemys exaratus Nessov and Khozatskiy 1973

G. †Macrobaena Tatarinov 1959

†Macrobaena mongolica Tatarinov 1959

G. †Osteopygis Cope 1868

†Osteopygis backmani Russell 1934

†Osteopygis emarginata Cope 1868

†Osteopygis emarginatus Cope 1868

Invalid names: Amblypeza entellus Hay 1908 [synonym], Catapleura chelydrina Cope 1875 [synonym], Osteopygis borealis Wieland 1904 [synonym], Osteopygis chelydrinus Cope 1868 [synonym], Osteopygis erosus Cope 1875 [synonym], Osteopygis gibbi Wieland 1904 [synonym], Osteopygis platylomus Cope 1870 [synonym], Osteopygis robustus Hay 1908 [synonym]

†Osteopygis kranzi Weems 2014

†Osteopygis russelli Brinkman 2015

G. †Oxemys Nessov 1977

†Oxemys gutta Nessov 1977

G. †Yakemys Tong et al. 2021

†Yakemys multiporcata Tong et al. 2021

Invalid names: Asiachelys Sukhanov 2000 [nomen nudum], Batoremys Narmandakh 2000 [nomen nudum]

G. †Owadowia Szczygielski et al. 2017

†Owadowia borsukbialynickae Szczygielski et al. 2017

Fm. †Thalassemydidae Zittel 1889

G. †Changisaurus Young 1959

†Changisaurus microrhinus Young 1959

†Changisaurus wutuensis Young 1961

G. †Phyllemys Schmidt 1944

G. †Sontiochelys Stache 1905

†Sontiochelys cretacea Stache 1905

G. †Yaxartemys Riabinin 1948

†Yaxartemys longicauda Riabinin 1948

Invalid names: Acichelyidae Lydekker 1889 [synonym], Emys beaugrandi Sauvage 1872 [nomen dubium], Trionyx primoevus Bergounioux 1937 [nomen dubium]

Unr. †Panpleurodira Danilov and Parham 2008

G. †Caribemys Fuente and Iturralde-Vinet 2001

Unr. †Dortokidae Lapparent de Broin and Murelaga 1996

G. †Dortoka Lapparent de Broin and Murelaga 1996

†Dortoka botanica Gheerbrant et al. 1999

†Dortoka vasconica Lapparent de Broin and Murelaga 1996

†Dortoka vremiri Augustin et al. 2021

Invalid names: Ronella Gheerbrant et al. 1999 [synonym]

G. †Eodortoka Pérez-García et al. 2014

†Eodortoka morellana Pérez-García et al. 2014

Invalid names: Muehlbachia Vremir and Codrea 2009 [nomen nudum], Muehlbachia nopcsai Vremir and Codrea 2009 [nomen nudum]

G. Pelomedusa Wagler 1830 [African helmeted turtle]

†Pelomedusa senutpickfordina de Lapparent de Broin 2008

Pelomedusa subrufa Bonaterre 1789 [Helmeted turtle]

Unr. †Platychelyidae

G. †Notoemys Cattoi and Freiberg 1961

†Notoemys laticentralis Cattoi and Freiberg 1961

†Notoemys oxfordiensis de la Fuente and Iturralde-Vinent 2001

†Notoemys tlaxiacoensis Lopez-Conde et al. 2017

†Notoemys zapatocaensis Cadena Rueda and Gaffney 2005

G. †Platychelys Wagner 1853

†Platychelys oberndorferi Wagner 1853

Invalid names: Helemys Rutimeyer 1859 [nomen nudum], Helemys serrata Rutimeyer 1859 [nomen nudum], Platychelys courrenti Bergounioux 1935 [nomen nudum]

Subor. Pleurodira Cope 1865 [sideneck turtle]

Unr. †Bothremydidae Baur 1891

G. †Arenila Lapparent de Broin and Werner 1998

†Arenila krebsi Lapparent de Broin and Werner 1998

G. †Atolchelys Romano et al. 2014

†Atolchelys lepida Romano et al. 2014

G. †Azabbaremys Gaffney et al. 2001

†Azabbaremys moragjonesi Gaffney et al. 2001

Subfm. †Bothremydinae Gaffney 1975

G. †Paiutemys Joyce et al. 2016

Invalid names: Cearachelyodda Gaffney et al. 2006 [empty]

Tr. †Bothremydini Gaffney et al. 2006

Subtr. †Bothremydina Gaffney et al. 2006

G. †Akoranemys Pérez-Garcia 2020

G. †Algorachelus Pérez-García 2017

G. †Araiochelys Gaffney et al. 2006

G. †Bothremys Leidy 1865

Invalid names: Karkaemys Zalmout et al. 2005 [synonym]

G. †Zolhafah Lapparent de Broin and Werner 1998

Subtr. †Foxemydina Gaffney et al. 2006

G. †Foxemys Tong et al. 1998

G. †Iberoccitanemys Pérez-García et al. 2012

G. †Polysternon Portis 1882

G. †Tartaruscola Pérez-García 2016

Invalid names: Polysternon isonae Marmi et al. 2012 [nomen dubium]

G. †Khargachelys AbdelGawad et al. 2023

†Khargachelys caironensis AbdelGawad et al. 2023

G. †Palauchelys López-Conde et al. 2021

†Palauchelys montellanoi López-Conde et al. 2021

G. †Puentemys Cadena et al. 2012

†Puentemys mushaisaensis Cadena et al. 2012

Unr. †Bothremydodda Gaffney et al. 2006

G. †Abalakemys Pérez-García 2023

†Abalakemys chapmanae Pérez-García 2023

G. †Carteremys Williams 1953

†Carteremys leithii Carter 1852

Tr. †Cearachelyini Gaffney et al. 2006

G. †Elkanemys Maniel et al. 2021

†Elkanemys pritchardi Maniel et al. 2021

G. †Galianemys Gaffney et al. 2002

†Galianemys emringeri Gaffney et al. 2002

†Galianemys whitei Gaffney et al. 2002

G. †Cearachelys Gaffney et al. 2001

†Cearachelys placidoi Gaffney et al. 2001

G. †Chedighaii Gaffney et al. 2006

†Chedighaii barberi Schmidt 1940

Invalid names: Podocnemis alabamae Zangerl 1948 [synonym]

†Chedighaii hutchisoni Gaffney et al. 2006

G. †Elochelys Nopsca 1931

†Elochelys perfecta Nopsca 1931

G. †Inaechelys de Araujo Carvalho et al. 2016

Tr. †Kurmademydini Gaffney et al. 2006

G. †Jainemys Joyce and Bandyopadhyay 2020

†Jainemys pisdurensis Jain 1977

G. †Kinkonychelys Gaffney et al. 2009

†Kinkonychelys rogersi Gaffney et al. 2009

G. †Kurmademys Gaffney et al. 2001

†Kurmademys kallamedensis Gaffney et al. 2001

G. †Labrostochelys Gaffney et al. 2006

†Labrostochelys galkini Gaffney et al. 2006

G. †Nigeremys Broin 1977

†Nigeremys gigantea Bergounoux and Crouzel 1968

Invalid names: Potamochelys Fitzinger 1843 [synonym]

G. †Palemys Gray 1870

†Palemys bowerbankii Owen 1842

Invalid names: Dacochelys delabechei Bell 1849 [synonym], Emys conybearii Owen 1858 [synonym], Emys delabechii Owen and Bell 1849 [synonym], Emys laevis Owen and Bell 1849 [synonym]

Invalid names: Dacochelys Lydekker 1889 [synonym]

G. †Phosphatochelys Gaffney and Tong 2003

†Phosphatochelys tedfordi Gaffney and Tong 2003

G. †Pleurochayah Adrian et al. 2021

†Pleurochayah appalachius Adrian et al. 2021

G. †Rhothonemys Gaffney et al. 2006

†Rhothonemys brinkmani Gaffney et al. 2006

G. †Rosasia Costa 1940

†Rosasia soutoi Carrington da Costa 1940

Invalid names: Inaechelys pernambucensis de Araujo Carvalho et al. 2016 [nomen dubium]

G. †Sankuchemys Gaffney et al. 2003

†Sankuchemys sethnai Gaffney et al. 2003

G. †Sindhochelys de Lapparent de Broin et al. 2021

†Sindhochelys ragei de Lapparent de Broin et al. 2021

Tr. †Taphrosphyini Gaffney et al. 2006

G. †Chupacabrachelys Lehman and Wick 2010

†Chupacabrachelys complexus Lehman and Wick 2010

G. †Ilatardia Pérez-García 2019

†Ilatardia cetiotesta Pérez-García 2019

Subtr. †Taphrosphyina Gaffney et al. 2006

G. †Acleistochelys Gaffney et al. 2007

G. †Eotaphrosphys Pérez-García 2018

G. †Motelomama Pérez-García 2018

Invalid names: Crassachelys Moody 1972 [nomen dubium], Eusarkia Bergounioux 1952 [nomen dubium], Gafsachelys Stefano 1903 [nomen dubium], Nigeremydina Gaffney et al. 2006 [empty]

G. †Taphrosphys Cope 1869

†Taphrosphys congolensis Dollo 1912

†Taphrosphys ippolitoi Gaffney et al. 2006

†Taphrosphys nodosus Cope 1870

†Taphrosphys strenuus Cope 1870

Invalid names: Prochonias princeps Cope 1869 [synonym]

†Taphrosphys sulcatus Leidy 1856

Invalid names: Amblypeza antellius Hay 1908 [synonym], Prochonias longinuchus Cope 1870 [synonym], Taphrosphys enodis Cope 1870 [synonym], Taphrosphys leslianus Cope 1870 [synonym], Taphrosphys molops Cope 1870 [synonym]

Invalid names: Amblypeza Hay 1908 [synonym], Bantuchelys Dollo 1924 [synonym], Prochonias Cope 1869 [synonym]

G. †Ummulisani Gaffney et al. 2006

†Ummulisani rutgersensis Gaffney et al. 2006

Invalid names: Kurmademydinae Gaffney et al. 2006 [empty], Podocnemis somaliensis Walker 1966 [invalid subgroup]

Fm. Chelidae Gray 1825

G. Acanthochelys Gray 1873

†Acanthochelys cosquinensis de la Fuente 1992

Acanthochelys macrocephala Rhodin et al. 1984 [Pantanal swamp turtle]

Acanthochelys pallidipectoris Freiberg 1945 [Chaco sideneck turtle]

Acanthochelys radiolata Mikan 1820 [Radiolated Brazilian swamp turtle]

Acanthochelys spixii Duméril and Bibron 1837 [black spine-neck swamp turtle]

G. †Baalemys Sarda and Maniel 2024

†Baalemys mansillai Sarda and Maniel 2024

G. †Birlimarr Megirian and Murray 1999

†Birlimarr gaffneyi Megirian and Murray 1999

G. †Calvarichelys Oriozabala et al. 2025

†Calvarichelys coloniensis Oriozabala et al. 2025

Subfm. Chelinae Gray 1825

G. Mesoclemmys Gray 1873

Mesoclemmys dahli Zangerl and Medem 1958

Mesoclemmys gibba Schweigger 1812 [Gibba turtle]

Mesoclemmys heliostemma McCord et al. 2001 [Yellow-crowned toad-headed turtle]

Mesoclemmys hogei Mertens 1967 [Hoge's side-necked turtle]

Mesoclemmys nasuta Schweigger 1812 [Guyanan toad-headed turtle]

Invalid names: Batrachemys nasuta Schweigger 1812 [synonym]

Mesoclemmys perplexa Bour and Zaher 2005 [Cerrado side-necked turtle]

Mesoclemmys raniceps Gray 1856 [Amazon toad-headed turtle]

Invalid names: Batrachemys raniceps Gray 1856 [synonym]

Mesoclemmys tuberculata Luederwaldt 1926 [Tuberculate toad-headed turtle]

Mesoclemmys vanderhaegei Bour 1973 [Vanderhaege's toad-headed turtle]

†Mesoclemmys vanegasorum Cadena et al. 2020

Mesoclemmys zuliae Pritchard and Trebbau 1984 [Zulia toad-headed turtle]

Invalid names: Batrachemys Stejneger 1909 [synonym], Bufocephala McCord et al. 2001 [synonym], Ranacephala McCord et al. 2001 [synonym]

G. Rhinemys Wagler 1830

Rhinemys rufipes Spix 1824 [Red side-necked turtle]

Subfm. Chelodininae Baur 1893

G. Chelodina Fitzinger 1826

Subg. Chelodina (Chelodina) Fitzinger 1826

Subg. Chelodina (Chelydera) Shea et al. 2020

Subg. Chelodina (Macrochelodina) Wells and Wellington 1985

Invalid names: Chelodina (Macrodiremys) McCord and Joseph-Ouni 2007 [objective synonym]

Chelodina longicollis Shaw 1802 [Eastern snake-necked turtle]

Chelodina mccordi Rhodin 1994 [McCord's snakeneck turtle]

Chelodina novaeguineae Boulenger 1888 [New Guinea snakeneck turtle]

Chelodina pritchardi Rhodin 1994 [Pritchard's snakeneck turtle]

Chelodina reimanni Philippen and Grossmann 1990 [Reimann's snakeneck turtle]

Chelodina steindachneri Siebenrock 1914 [Steindachner's snakeneck turtle]

G. Elusor Cann and Legler 1994

Elusor macrurus Cann and Legler 1994 [Mary River turtle]

G. Flaviemys Le et al. 2013 [Manning River sawshelled turtle]

Flaviemys purvisi Wells and Wellington 1985 [Manning River sawshelled turtle]

G. Chelus Duméril 1806

†Chelus colombianus Wood 1976 [Chelus colombiana]

Invalid names: Chelus lewisi Wood 1976 [synonym]

Chelus fimbriata Schneider 1783 [Mata mata]

Chelus fimbriatus Schneider 1783 [Mata mata]

G. Elseya Gray 1867

Elseya albagula Thomson et al. 2006 [White-throated snapping turtle]

Elseya branderhorsti Ouwens 1914 [White-bellied snapping turtle]

Elseya dentata Gray 1863 [Northern Australia snapping turtle]

Elseya irwini Cann 1997 [Irwins snapping turtle]

Elseya lavarackorum White 1994

Elseya macrurus Cann and Legler 1994 [Cann's snakeneck turtle]

†Elseya nadibajagu Thomson and Mackness 1999

Elseya novaeguineae Meyer 1874 [New Guinea snapping turtle]

Elseya oneiros Joseph-Ouni et al. 2020

Elseya schultzei Vogt 1911 [Schultzes snapping turtle]

†Elseya uberrima De Vis 1897

Invalid names: Pelecomastes De Vis 1897 [synonym]

G. Emydura Bonaparte 1836

Emydura australis Gray 1841 [Australian bighead turtle]

Emydura krefftii Gray 1871 [Krefft's river turtle]

Emydura macquarii Gray 1831 [Murray River turtle]

Emydura signata Ahl 1932 [Brisbane shortneck turtle]

Emydura subglobosa Krefft 1876 [Redbelly shortneck turtle]

Emydura tanybaraga Cann 1997 [Northern yellow-faced turtle]

Emydura victoriae Gray 1842 [Victoria shortneck turtle]

G. Hydromedusa Wagler 1830

†Hydromedusa casamayorensis de la Fuente and Bona 2002

Hydromedusa maximiliani Mikan 1820 [Brazilian snakeneck turtle]

Hydromedusa tectifera Cope 1869 [Argentine snake-necked turtle]

Invalid names: Platemys antiqua Ameghino 1882 [synonym], Platemys fossilis Ameghino 1882 [synonym], Platemys laevis Ameghino 1882 [synonym], Platemys robusta Ameghino 1882 [synonym]

Subfm. Hydromedusinae Baur 1893

G. †Iaremys Agnolín et al. 2024

†Iaremys batrachomorpha Agnolín et al. 2024

G. †Mendozachelys de la Fuente et al. 2017

†Mendozachelys wichmanni de la Fuente et al. 2017

G. Myuchelys Thomson and Georges 2009

Myuchelys bellii Gray 1844

Myuchelys georgesi Georges and Adams 1992

Myuchelys latisternum Gray 1867 [Serrated snapping turtle]

Myuchelys purvisi Wells and Wellington 1985

G. Phrynops Wagler 1830

Phrynops geoffroanus Schweigger 1812 [side-necked turtle of Geoffroy]

Phrynops hilarii Dumeril and Bibron 1835 [side-necked turtle of Saint Hillaire]

†Phrynops paranaensis Wieland 1923

Phrynops tuberosus Peters 1870 [Guianan Shield side-necked turtle]

Phrynops williamsi Rhodin and Mittermeier 1983 [Williams side-necked turtle]

Phrynops zuliae Pritchard and Trebbau 1984 [Zulia toadhead turtle]

Invalid names: Parahydraspis Wieland 1923 [synonym]

G. Platemys Wagner 1830

Platemys platycephala Schneider 1792 [Twistneck turtle]

G. †Prochelidella Lapparent de Broin and de la Fuente 2001

†Prochelidella argentinae Lapparent de Broin and de la Fuente 2001

†Prochelidella buitreraensis Maniel et al. 2020

†Prochelidella cerrobarcinae de la Fuente et al. 2011

†Prochelidella palomoi Maniel et al. 2022

†Prochelidella portezuelae de la Fuente 2003

G. Pseudemydura Siebenrock 1901

Pseudemydura umbrina Siebenrock 1901 [Western swamp turtle]

G. Rheodytes Legler and Cann 1980

†Rheodytes devisi Thomson 2000

Rheodytes leukops Legler and Cann 1980 [Fitzroy River turtle]

G. †Rionegrochelys De la Fuente et al. 2017

†Rionegrochelys caldieroi De la Fuente et al. 2017

G. †Yaminuechelys de la Fuente et al. 2001

†Yaminuechelys gasparinii de la Fuente et al. 2001

†Yaminuechelys maior Staesche 1929

Invalid names: Osteopygis sculptus Staesche 1929 [synonym]

†Yaminuechelys sulcipeculiaris Oriozabala et al. 2020

Invalid names: Chelydidae Gray 1869 [synonym], Chelymys antiqua De Vis 1897 [nomen dubium], Chelymys arata De Vis 1897 [nomen dubium], Hydraspididae Cope 1868 [synonym], Hydraspis arenarius Rusconi 1934 [nomen dubium], Naiadochelys patagonica Staesche 1929 [nomen dubium], Pelecomastes ampla De Vis 1897 [nomen dubium], Platemys bergii Ambrosetti 1893 [nomen dubium], Platemys entreriana Ambrosetti 1893 [nomen dubium], Platemys holmbergii Ambrosetti 1893 [nomen dubium], Platemys mesopotamica Ambrosetti 1893 [nomen dubium], Platemys robusta Ambrosetti 1893 [nomen dubium], Platemys torrentium Burmeister 1885 [nomen dubium], Trionyx argentina Ameghino 1889 [nomen dubium]

Unr. Eupleurodira Gaffney and Meylan 1988

Invalid names: Bothremydinura Gaffney et al. 2006 [empty], Cheloides Gray 1825 [empty]

Unr. †Panchelidae Joyce et al. 2004

G. †Bonapartemys Lapparent de Broin and de la Fuente 2001

†Bonapartemys bajobarrealis Lapparent de Broin and de la Fuente 2001

G. †Linderochelys de la Fuente et al. 2007

†Linderochelys rinconensis de la Fuente et al. 2007

G. †Lomalatachelys Lapparent de Broin and de la Fuente 2001

†Lomalatachelys neuquina Lapparent de Broin and de la Fuente 2001

G. †Palaeophrynops Lapparent de Broin and de la Fuente 2001

†Palaeophrynops patagonicus Lapparent de Broin and de la Fuente 2001

G. †Salamanchelys Bona 2006

†Salamanchelys palaeocenica Bona 2006

Invalid names: Platemys guaranitica Ameghino 1889 [nomen dubium], Platemys patagonica Ameghino 1889 [nomen dubium]

Unr. †Pelomedusoides Broin 1988

G. †Apodichelys Price 1954

†Apodichelys lucianoi Price 1954

Fm. †Araripemydidae Price 1973

G. †Araripemys Price 1973

†Araripemys barretoi Price 1973

Invalid names: Araripemys arturi Fielding et al. 2005 [synonym]

G. †Laganemys Sereno and ElShafie 2013

G. †Taquetochelys Broin 1980

†Taquetochelys decorata de Broin 1980

Invalid names: Laganemys tenerensis Sereno and ElShafie 2013 [synonym]

G. †Caririemys Oliveira and Kellner 2007

†Caririemys violetae Oliveira and Kellner 2007

G. †Francemys Pérez-García 2019

†Francemys gadoufaouaensis Pérez-García 2019

G. †Itapecuruemys Batista et al. 2021

†Itapecuruemys amazonensis Batista et al. 2021

G. †Palaeaspis Gray 1870

†Palaeaspis bowerbanki Owen 1842

Fm. †Peiropemydidae Gaffney et al. 2011

G. †Pricemys Gaffney et al. 2011

†Pricemys caiera Gaffney et al. 2011

G. †Yuraramirim Ferreira et al. 2018

†Yuraramirim montealtensis Ferreira et al. 2018

Unr. †Pelomedusera Gaffney et al. 2006

Fm. Pelomedusidae Cope 1868

G. †Duerochelys Jimenez Fuentes 1975

G. †Latisternon Auffenberg 1981

G. Pelusios Wagler 1830

Invalid names: Eusarkiidae Bergounioux 1952 [synonym], Pelomedusinae Zangerl 1948 [empty], Peltocephalidae Gray 1870 [synonym], Pelusiinae Zangerl 1948 [empty], Sternothaeridae Cope 1868 [synonym]

Unr. †Podocnemidera

Superfm. †Euraxemydoidea Gaffney et al. 2006

Fm. †Euraxemydidae Gaffney et al. 2006

G. †Teneremys Broin 1980

†Teneremys lapparenti de Broin 1980

Invalid names: Platycheloides nyasae Haughton 1928 [nomen dubium]

Superfm. Podocnemidoidae Cope 1868

G. †Amabilis Hermanson et al. 2020

†Amabilis uchoensis Hermanson et al. 2020

G. †Roxochelys Price 1953

†Roxochelys vilavilensis de Broin 1971

†Roxochelys wanderleyi Price 1953

Invalid names: Cambaremys langertoni França and Langer 2005 [synonym]

Invalid names: Roxochelys harrisi Pacheco 1913 [nomen dubium]

Invalid names: Brasilemydidae Lapparent de Broin 2000 [empty]

Superfm. †Podocnemidoidea Gaffney et al. 2006

G. †Nostimochelone Georgalis et al. 2013

†Nostimochelone lampra Georgalis et al. 2013

Fm. Podocnemididae Cope 1868

G. †Albertwoodemys Gaffney et al. 2011

†Albertwoodemys testudinum Gaffney et al. 2011

G. †Bairdemys Gaffney and Wood 2002

†Bairdemys hartsteini Gaffney and Wood 2002

†Bairdemys healeyorum Weems and Knight 2013

†Bairdemys miocenica Collins and Lynn 1936

†Bairdemys sanchezi Gaffney et al. 2008

†Bairdemys thalassica Ferreira et al. 2015

†Bairdemys venezuelensis Wood and Diaz 1971

†Bairdemys winklerae Gaffney et al. 2008

G. †Bauruemys Suarez 1969

†Bauruemys elegans Suarez 1969

Invalid names: Bauruemys brasiliensis Staesche 1937 [nomen dubium]

G. †Brontochelys Gaffney et al. 2011

†Brontochelys gaffneyi Wood 1970

G. †Cambaremys França and Langer 2005

G. †Carbonemys Cadena et al. 2012

†Carbonemys cofrinii Cadena et al. 2012

G. †Cerrejonemys Cadena et al. 2010

†Cerrejonemys wayuunaiki Cadena et al. 2010

G. †Cordichelys Gaffney et al. 2011

†Cordichelys antiqua Andrews 1903

G. †Dacquemys Williams 1954

†Dacquemys paleomorpha Williams 1954

Subfm. †Erymnochelyinae Broin 1988

G. †Andrewsemys Pérez-García 2019

†Andrewsemys libyca Andrews 1903

G. †Caninemys Meylan et al. 2009

†Caninemys tridentata Meylan et al. 2009

G. †Eocenochelus Pérez-García et al. 2017

†Eocenochelus eremberti Broin 1977

†Eocenochelus farresi Pérez-García et al. 2017

†Eocenochelus lacombianus Pérez-García et al. 2017

Tr. †Erymnochelyini de Broin 1988

G. †Apeshemys Perez-Garcia 2021

G. †Mokelemys Pérez-García 2022

G. †Shetwemys Perez-Garcia 2021

G. †Neochelys Bergounioux 1954

†Neochelys arenarum Broin 1977

†Neochelys bolcensis Bergounioux 1954

†Neochelys capellinii De Zigno 1890

†Neochelys eocaenica de Stefano 1902

†Neochelys franzeni Schleich 1993

†Neochelys laurenti Tong 1998

†Neochelys liriae Pérez-García and Lapparent de Broin 2013

†Neochelys mengaudi Bergounioux 1935

†Neochelys nicolisii Zigno 1890

†Neochelys salmanticensis Jimenez 1971

Invalid names: Duerochelys arribasi Jimenez 1975 [synonym], Podocnemis armuniensis Jimenez 1970 [synonym], Podocnemis entodermica Jimenez 1970 [synonym]

†Neochelys zamorensis Jimenez-Fuentes 1992

Invalid names: Polysternum mengaudi Bergounioux 1935 [nomen dubium]

G. †Ragechelus de Lapparent de Broin et al. 2020

†Ragechelus sahelica de Lapparent de Broin et al. 2020

G. †Stupendemys Wood 1976

†Stupendemys geographica Wood 1976

†Stupendemys souzai Bocquetin and Melo 2006

G. Erymnochelys Baur 1888

Erymnochelys madagascariensis Grandidier 1867

G. †Gestemys de la Fuente et al. 2022

†Gestemys powelli de la Fuente et al. 2022

G. †Hamadachelys Tong and Buffetaut 1996

G. †Kenyemys Wood 1983

†Kenyemys williamsi Wood 1983

G. †Lapparentemys Gaffney et al. 2011

†Lapparentemys vilavilensis de Broin 1971

G. †Latentemys Gaffney et al. 2011

†Latentemys plowdeni Gaffney et al. 2011

G. †Lemurchelys Gaffney et al. 2011

†Lemurchelys diasphax Gaffney et al. 2011

G. †Mogharemys Gaffney et al. 2011

†Mogharemys blanckenhorni Dacque 1912

G. †Papoulemys Tong 1998

G. †Peiropemys Gaffney et al. 2011

†Peiropemys mezzalirai Gaffney et al. 2011

G. Peltocephalus Duméril and Bibron 1835

Peltocephalus dumerilianus Schweigger 1812 [Bighead Amazon River turtle]

†Peltocephalus maturin Ferreira et al. 2024

Subfm. Podocnemidinae Cope 1868

Unr. †Podocnemidodda Gaffney et al. 2011

Invalid names: Erymnochelydand Gaffney et al. 2011 [empty], Podocnemidand Gaffney et al. 2011 [empty]

G. Podocnemis Wagler 1830 [river turtle]

†Podocnemis argentinensis Cattoi and Freiberg 1958

†Podocnemis bassleri Williams 1956

†Podocnemis brasiliensis Staesche 1937

Podocnemis erythrocephala Spix 1824 [Red-headed river turtle]

Podocnemis expansa Schweigger 1812

†Podocnemis geologorum Simpson 1943

†Podocnemis harrisi Pacheco 1913

Podocnemis lewyana Dumeril 1852

†Podocnemis medemi Wood 1997

†Podocnemis negrii Carvalho et al. 2002

†Podocnemis pritchardi Wood 1997

Podocnemis sextuberculata Cornalia 1849

†Podocnemis stromeri von Reinach 1903

†Podocnemis tatacoensis Cadena and Vanegas 2023

Podocnemis unifilis Troschel 1848

Podocnemis vogli Muller 1935

Invalid names: Podocnemis bramlyi Fourtau 1918 [nomen dubium], Podocnemis freibergi Agnolin 2004 [nomen dubium]

G. †Shweboemys Swinton 1939

†Shweboemys pilgrimi Swinton 1939

Subtr. †Stereogenyina Gaffney et al. 2011

Invalid names: Bairdemydita Gaffney et al. 2011 [empty]

Tr. †Stereogenyini Gaffney et al. 2011

Unr. †Stereogenyita Gaffney et al. 2011

G. †Piramys Prasad 1974

Invalid names: Mogharemydina Gaffney et al. 2011 [empty]

G. †Stereogenys Andrews 1901

†Stereogenys cromeri Andrews 1901

Invalid names: Stereogenys podocnemoides Reinach 1903 [nomen dubium]

G. †Turkanemys Wood 2003

†Turkanemys pattersoni Wood 2003

Invalid names: Albertwoodemys Gaffney et al. 2011 [valid], Bairdemys Gaffney and Wood 2002 [valid], Bauruemydinae Gaffney et al. 2011 [empty], Bauruemydinae Gaffney et al. 2011 [empty], Bauruemys Suarez 1969 [valid], Brontochelys Gaffney et al. 2011 [valid], Cambaremys França and Langer 2005 [valid], Carbonemys Cadena et al. 2012 [valid], Cerrejonemys Cadena et al. 2010 [valid], Cordichelys Gaffney et al. 2011 [valid], Dacquemys Williams 1954 [valid], Erymnochelyinae Broin 1988 [valid], Erymnochelys Baur 1888 [valid], Gestemys de la Fuente et al. 2022 [valid], Hamadachelys Tong and Buffetaut 1996 [valid], Kenyemys Wood 1983 [valid], Lapparentemys Gaffney et al. 2011 [valid], Latentemys Gaffney et al. 2011 [valid], Lemurchelys Gaffney et al. 2011 [valid], Mogharemys Gaffney et al. 2011 [valid], Papoulemys Tong 1998 [valid], Peiropemys Gaffney et al. 2011 [valid], Peltocephalus Duméril and Bibron 1835 [valid], Podocnemidinae Cope 1868 [valid], Podocnemoidea Cope 1868 [empty], Podocnemoidea Cope 1868 [empty], Shweboemys Swinton 1939 [valid], Stereogenyina Gaffney et al. 2011 [valid], Stereogenyini Gaffney et al. 2011 [valid], Stereogenys Andrews 1901 [valid], Turkanemys Wood 2003 [valid]

Unr. †Podocnemidinura Gaffney et al. 2006

G. †Brasilemys Lapparent de Broin 2000

G. †Portezueloemys de la Fuente 2003

Unr. †Sahonachelyidae Joyce et al. 2021

G. †Sahonachelys Joyce et al. 2021

†Sahonachelys mailakavava Joyce et al. 2021

G. †Sokatra Gaffney and Krause 2011

†Sokatra antitra Gaffney and Krause 2011

Invalid names: Chrysemys montolivensis Roman 1897 [invalid subgroup], Naiadochelys Hay 1908 [nomen dubium], Panpodocnemidae Joyce et al. 2004 [empty], Taphrosphys dares Hay 1908 [nomen dubium]

Fm. Podocnemididae Cope 1868

G. †Albertwoodemys Gaffney et al. 2011

†Albertwoodemys testudinum Gaffney et al. 2011

G. †Bairdemys Gaffney and Wood 2002

†Bairdemys hartsteini Gaffney and Wood 2002

†Bairdemys healeyorum Weems and Knight 2013

†Bairdemys miocenica Collins and Lynn 1936

†Bairdemys sanchezi Gaffney et al. 2008

†Bairdemys thalassica Ferreira et al. 2015

†Bairdemys venezuelensis Wood and Diaz 1971

†Bairdemys winklerae Gaffney et al. 2008

G. †Bauruemys Suarez 1969

†Bauruemys elegans Suarez 1969

Invalid names: Bauruemys brasiliensis Staesche 1937 [nomen dubium]

G. †Brontochelys Gaffney et al. 2011

†Brontochelys gaffneyi Wood 1970

G. †Cambaremys França and Langer 2005

G. †Carbonemys Cadena et al. 2012

†Carbonemys cofrinii Cadena et al. 2012

G. †Cerrejonemys Cadena et al. 2010

†Cerrejonemys wayuunaiki Cadena et al. 2010

G. †Cordichelys Gaffney et al. 2011

†Cordichelys antiqua Andrews 1903

G. †Dacquemys Williams 1954

†Dacquemys paleomorpha Williams 1954

Subfm. †Erymnochelyinae Broin 1988

G. †Andrewsemys Pérez-García 2019

†Andrewsemys libyca Andrews 1903

G. †Caninemys Meylan et al. 2009

†Caninemys tridentata Meylan et al. 2009

G. †Eocenochelus Pérez-García et al. 2017

†Eocenochelus eremberti Broin 1977

†Eocenochelus farresi Pérez-García et al. 2017

†Eocenochelus lacombianus Pérez-García et al. 2017

Tr. †Erymnochelyini de Broin 1988

G. †Apeshemys Perez-Garcia 2021

G. †Mokelemys Pérez-García 2022

G. †Shetwemys Perez-Garcia 2021

G. †Neochelys Bergounioux 1954

†Neochelys arenarum Broin 1977

†Neochelys bolcensis Bergounioux 1954

†Neochelys capellinii De Zigno 1890

†Neochelys eocaenica de Stefano 1902

†Neochelys franzeni Schleich 1993

†Neochelys laurenti Tong 1998

†Neochelys liriae Pérez-García and Lapparent de Broin 2013

†Neochelys mengaudi Bergounioux 1935

†Neochelys nicolisii Zigno 1890

†Neochelys salmanticensis Jimenez 1971

Invalid names: Duerochelys arribasi Jimenez 1975 [synonym], Podocnemis armuniensis Jimenez 1970 [synonym], Podocnemis entodermica Jimenez 1970 [synonym]

†Neochelys zamorensis Jimenez-Fuentes 1992

Invalid names: Polysternum mengaudi Bergounioux 1935 [nomen dubium]

G. †Ragechelus de Lapparent de Broin et al. 2020

†Ragechelus sahelica de Lapparent de Broin et al. 2020

G. †Stupendemys Wood 1976

†Stupendemys geographica Wood 1976

†Stupendemys souzai Bocquetin and Melo 2006

G. Erymnochelys Baur 1888

Erymnochelys madagascariensis Grandidier 1867

G. †Gestemys de la Fuente et al. 2022

†Gestemys powelli de la Fuente et al. 2022

G. †Hamadachelys Tong and Buffetaut 1996

G. †Kenyemys Wood 1983

†Kenyemys williamsi Wood 1983

G. †Lapparentemys Gaffney et al. 2011

†Lapparentemys vilavilensis de Broin 1971

G. †Latentemys Gaffney et al. 2011

†Latentemys plowdeni Gaffney et al. 2011

G. †Lemurchelys Gaffney et al. 2011

†Lemurchelys diasphax Gaffney et al. 2011

G. †Mogharemys Gaffney et al. 2011

†Mogharemys blanckenhorni Dacque 1912

G. †Papoulemys Tong 1998

G. †Peiropemys Gaffney et al. 2011

†Peiropemys mezzalirai Gaffney et al. 2011

G. Peltocephalus Duméril and Bibron 1835

Peltocephalus dumerilianus Schweigger 1812 [Bighead Amazon River turtle]

†Peltocephalus maturin Ferreira et al. 2024

Subfm. Podocnemidinae Cope 1868

Unr. †Podocnemidodda Gaffney et al. 2011

Invalid names: Erymnochelydand Gaffney et al. 2011 [empty], Podocnemidand Gaffney et al. 2011 [empty]

G. Podocnemis Wagler 1830 [river turtle]

†Podocnemis argentinensis Cattoi and Freiberg 1958

†Podocnemis bassleri Williams 1956

†Podocnemis brasiliensis Staesche 1937

Podocnemis erythrocephala Spix 1824 [Red-headed river turtle]

Podocnemis expansa Schweigger 1812

†Podocnemis geologorum Simpson 1943

†Podocnemis harrisi Pacheco 1913

Podocnemis lewyana Dumeril 1852

†Podocnemis medemi Wood 1997

†Podocnemis negrii Carvalho et al. 2002

†Podocnemis pritchardi Wood 1997

Podocnemis sextuberculata Cornalia 1849

†Podocnemis stromeri von Reinach 1903

†Podocnemis tatacoensis Cadena and Vanegas 2023

Podocnemis unifilis Troschel 1848

Podocnemis vogli Muller 1935

Invalid names: Podocnemis bramlyi Fourtau 1918 [nomen dubium], Podocnemis freibergi Agnolin 2004 [nomen dubium]

G. †Shweboemys Swinton 1939

†Shweboemys pilgrimi Swinton 1939

Subtr. †Stereogenyina Gaffney et al. 2011

Invalid names: Bairdemydita Gaffney et al. 2011 [empty]

Tr. †Stereogenyini Gaffney et al. 2011

Unr. †Stereogenyita Gaffney et al. 2011

G. †Piramys Prasad 1974

Invalid names: Mogharemydina Gaffney et al. 2011 [empty]

G. †Stereogenys Andrews 1901

†Stereogenys cromeri Andrews 1901

Invalid names: Stereogenys podocnemoides Reinach 1903 [nomen dubium]

G. †Turkanemys Wood 2003

†Turkanemys pattersoni Wood 2003

Invalid names: Amphichelydidae Hay [empty], Megapleurodira Gaffney et al. 2006 [empty], Minipleurodira Gaffney et al. 2006 [empty], Notoemydidae Fernandez and de la Fuente 1994 [empty], Panpelomedusoides Joyce et al. 2004 [empty], Platychelira Gaffney et al. 2006 [empty]

Subor. †Proganochelydia

Fm. †Proganochelyidae

Invalid names: Triassochelyidae Abel 1919 [synonym]

G. †Tacuarembemys Perea et al. 2014

†Tacuarembemys kusterae Perea et al. 2014

G. †Triassochelys Jaekel 1918

Invalid names: Stegochelys Jaekel 1914 [replaced]

Invalid names: Apertotemporalis Stromer 1934 [nomen dubium], Apholidemys Pomel 1847 [nomen dubium], Casichelydia Gaffney 1975 [empty], Chelone obovata Owen 1842 [nomen dubium], Chelone sopita Leidy 1865 [nomen vanum], Chelonia Latreille 1800 [synonym], Chelonii Latreille 1800 [synonym], Cryptrodiramorpha Lee 1995 [empty], Euclastochelys Bergounioux 1955 [nomen dubium], Euclastochelys interrupta Bergounioux 1955 [nomen dubium], Meiolania damelipi White et al. 2010 [nomen dubium], Platychelone Dollo 1909 [nomen nudum], Platychelone emarginata Dollo 1909 [nomen nudum], Plesiochelydiae [empty], Podocnemis lata Ristori 1895 [invalid subgroup], Senonemys suemegensis Bohn 1966 [nomen dubium], Sokotochelys Halstead 1979 [nomen dubium], Temnoclemmys cordiformis Bergounioux 1958 [nomen vanum], Trionyx fuchienensis Yeh 1974 [nomen dubium], Trionyx tshelkarensis Chkhikvadze 1973 [nomen nudum]

Reference	Diagnosis
R. Owen 1860 (Chelonia)	Trunk-ribs broad, flat, suturally united, forming with vertebrae and sternum an expanded thoracic abdominal case, into which, as into a portable chamber, the head, tail, and limbs can, usually, be withdrawn. No teeth: external nostril single.
R. Owen 1861 (Chelonia)	Trunk-ribs broad, flat, suturally united, forming, with their vertebra, the sternum, and dermal bones, an expanded thoracic-abdominal case, into which the limbs, tail, and, usually, the head, can be withdrawn. No teeth: external nostril single.
E. Perrier 1928 (Chelonia)	Corps de forme arrondie, revêlu de plaques cornées ou d'un épais tégument, recouvrant d'autre part une carapace formée d'os dermiques plus ou moins développés, lesquels se soudent à leur tour aux apophyses épineuses des vertèbres, aux côtes et au sternum; ceintures scapulaire et pelviènne demeurant, sans perdre leurs caractères essentiels, à l'intérieur de la carapace. Membres courts, construits pour pousser l'animal plutôt que pour servir à une marche régulière. Pas de dents; mâchoires courtes, revêtues par une gaîne cornée rappelant celle du bec des Oiseaux.

Time interval	Ma	Country or state	Original ID and collection number
Oldest occurrences
Olenekian - Anisian	249.9 - 241.464	Germany (Thuringia)	Chelonipus cuneiformis, Chelonipus torquatus (182300)
Spathian	248.1 - 246.7	USA (Wyoming)	Chelonipus sp. (135635 135636 135637)
Tuvalian	233.6 - 227.3	China (Guizhou)	Odontochelys semitestacea (85166)
Norian	227.3 - 205.7	USA (New Mexico)	Testudines indet. (80926)
Norian	227.3 - 205.7	Germany (Bayern)	Reptilia indet. (48683)
Rhaetian	205.7 - 201.4	Italy (Lombardy)	Emydidae indet. (38083)
Hettangian - Sinemurian	201.4 - 192.9	USA (Arizona)	Testudines indet. (216632 216633 216634)
Sinemurian	199.5 - 192.9	Germany (Bavaria)	Thalassemydidae indet. (144689)
Sinemurian - Pliensbachian	199.5 - 184.2	USA (Arizona)	Cryptodira indet. (51898) Testudines indet. (51895 216645)
Sinemurian - Aalenian	199.5 - 170.9	India (Telangana)	Testudines indet. (61857)
Toarcian - Aalenian	184.2 - 170.9	China (Chongqing)	Xinjiangchelyidae indet., Protoxinjiangchelys sp. (212359)
Middle Jurassic	174.7 - 161.5	China (Yunnan)	Chelonia indet. (49774)
Bajocian	170.9 - 168.2	China (Yunnan)	Testudines indet. (70438)
Bajocian - Oxfordian	170.9 - 154.8	China (Sichuan)	Chengyuchelys dashanpuensis, Protoxinjiangchelys sp., Chengyuchelys sp., Chengyuchelyidae indet., Chengyuchelys baenoides, Chelonia indet., Chengyuchelys zigongensis (24883)
Late/Upper Bajocian - Early/Lower Bathonian	168.6 - 165.3	France (Occitanie)	Testudines indet. (58416)
Bathonian	168.2 - 165.3	Madagascar (Mahajunga)	Chelonia indet. (194795 194853 194861)
Bathonian	168.2 - 165.3	United Kingdom (Scotland)	Chelonia indet. (105872 139746)
Bathonian	168.2 - 165.3	Russian Federation (Krasnoyarsk)	Testudines indet. 1, Testudines indet. 2, Testudines indet. 3, Xinjiangchelys sp., Annemys sp. (61891)
Early/Lower Bathonian	168.2 - 165.3	United Kingdom (England)	Testudines indet. (31054)
Late/Upper Bathonian	168.2 - 165.3	United Kingdom (England)	Chelonia indet. (57139 166423) Chelonia indet., Cryptodira indet. (57124 57125) Cryptodira histomorph II (39075)