| Basic info | Taxonomic history | Classification | Included Taxa |
| Morphology | Ecology and taphonomy | External Literature Search | Age range and collections |
Coleoptera (beetle)
Taxonomy
Coleoptera was named by Linnaeus (1758). It is extant.
It was assigned to Oligometabola by Geyer and Kleber (1987); to Endopterygota by Carpenter (1992); to Insecta by Linnaeus (1758), Schreber (1759), Linnaeus (1761), Geoffroy (1762), Scopoli (1763), Müller (1764), Schaeffer (1766), Linnaeus (1767), Strøm (1768), Drury (1770), Forster (1771), Müller (1776), Goeze (1777), Latreille (1796), Cockerell (1921), Bode (1953), Zhang (1989), Shoshani et al. (1989), Lawrence and Newton (1995); and to Coleopterida by Kukalová-Peck and Lawrence (2004), Beutel et al. (2011), Peters et al. (2014), Misof et al. (2014), Engel (2022).
It was assigned to Oligometabola by Geyer and Kleber (1987); to Endopterygota by Carpenter (1992); to Insecta by Linnaeus (1758), Schreber (1759), Linnaeus (1761), Geoffroy (1762), Scopoli (1763), Müller (1764), Schaeffer (1766), Linnaeus (1767), Strøm (1768), Drury (1770), Forster (1771), Müller (1776), Goeze (1777), Latreille (1796), Cockerell (1921), Bode (1953), Zhang (1989), Shoshani et al. (1989), Lawrence and Newton (1995); and to Coleopterida by Kukalová-Peck and Lawrence (2004), Beutel et al. (2011), Peters et al. (2014), Misof et al. (2014), Engel (2022).
Subtaxa
Acmaeodera perreaui, Adelidium, Adikia, Adynasia, Allognosis, Allopliosilpha, Alphacoleoptera, Anancaeon, Anapiptus, Ancylocheira liasina, Anepismus, Angelinella, Anhuistoma, Anhydrophilus, Anomerus, Anypostatus, Apheloodes, Apicasia, Apioderes, Apiopyrenides, Apistotes, Apsychus, Aptilotitus (syn. Aptilotus), Archosyne, Auchenophorites (syn. Auchenophorus), Babuskaya, Bareus, Bathygerus, Bellingera, Bellingeropsis, Biadelater, Blaptoides, Bothroptera, Brachylaimon, Brachytrachelites, Brevispinnatus, Brochocoleus indibili, Brodiola, Bucklandula, Buprestium, Campopsis, Carabicina, Carabiques, Carabocera, Cardioides, Cardiosyne, Cebrionates, Cerylonopsis, Chimaerocoleites, Chlaeniopsis, Chrysomelidium, Chrysomelinae, Chrysomelophana, Cicindeletae, Cistela, Cistelites, Clathropenna, Clerii, Clinomerus (syn. Catomerus), Coccinellophana, Coleopterites, Coleopteron, Coleopteron depressus, Coleopteron interpunctatus, Coleopteron laevigatus, Coleopteron scabriusculus, Colymbetopsis, Coprophagi, Cretaceites, Cretofalselaterium, Cretogermen, Cretohelophorus, Criocerina pristina, Critoderma (syn. Cycloderma), Cryptauchenia, Curculidium, Curculionites liasinus, Curculiopsidae, Curculium, Cydnopsis nigromembranacea, Cyphospheron, Darbiopelta, Diachoristes, Diaperialae, Diaperidium, Diatarastus, Diatrypamene, Dicyphelus (syn. Dicyphus), Diphymation, Diplocelides, Diplothece, Doggeria, Dysarestus, Ecthlimma, Elateridopsidae, Elateridopsis, Elaterina, Elytrulum, Enamma, Entomocantharus, Entomophagi, Eocoleopteron, Eodromeus viriosus, Episcepes, Epomenus, Erotilenae, Erotylites, Etheridgea, Euchroma arvernica, Eucoleoptera, Eurynotellus (syn. Eurynotus), Eurysphinctus, Eusarcantarus, Explansimarginis, Fengningia, Flichea, Fuscicupes, Gastrodelus, Gastroratus, Gemelina, Glaphoptera, Glaphyropterites, Glaphyropterula, Grahamelytron, Grasselites, Gurvanocoleus, Gyrinites, Gyrinulopsis, Halticophana, Harpalidium, Heeriaopsis (syn. Heeriella), Helophoropsis, Helophorus exilis, Helophorus magnus, Helopidium, Helopii, Helopium, Helops atticus, Hubeicoleus, Hydrobius godeti, Hydrobius obsoletus, Hydrocanthares, Hydroicetes, Hydrophilites, Hydrophilus stenopterus, Hydroporopsis, Hydrous ebeninus, Hylesinus lineatus, Hyperomima, Ipidae, Isomira aurora, Kamaroma, Katapiptus, Keleusticus, Kibdelia, Kryptophagus, Laimocenos, Lamiites, Lamiophanes, Laodiscis, Latridiites, Leioodes, Leptilostis (syn. Tetillopsis), Leptomites, Leptopodocoleidae, Leptosolenophorus, Longimaxilla, Loxocamarotus, Macrogastri, Macrotrachelites, Malacodermi, Malacodermidae, Masselytron, Megacentrus, Megachorites, Megopis lineolatus, Melanophilites, Melanophilopsis, Menephiloides, Mesocoleoptera, Mesoncus (syn. Loxoncus), Mesorhynchophora, Mesotricupes, Mesotylites, Metanastes, Metaxycoleoptera, Micrelaterium, Microcoleopteron, Microentomus, Mimelater, Nannoodes, Nebrioides, Necrophagi, Necticus minutus, Nitidulariae, Nubili, Omogongylus, Onkedodimus, Onthophagus luteus, Ooidellus (syn. Ooides), Ooperiglyptus, Ooperioristus, Oropseini, Oxycephalites, Oxytoroptera, Pachycoleon, Pachypleurites, Paklapwania, Palaeotrachys, Palaeotylinae, Pallax, Palpatores, Pantodapus, Parabuprestium, Paracurculium, Paragrilium, Paragyrinus, Parasilphites, Parnidium, Parnosoma, Pedinosoma, Peltocoleops, Periboloptera, Peridosoma (syn. Perosoma), Petalocupes, Petrorophus, Phaulogyrinus, Pholipheron, Phytoplesion, Pimeliodes, Plastobuprestites, Plastonebria, Platyprosopi, Pleuralocista, Polypamon, Praedodromeus, Prionophana, Prochrysomela, Proheuristes, Prophasis, Prosthecogmus, Prosthenostictus, Protocoleoptera, Protocuneus, Prototoma, Pseudobuprestites, Pseudocarabites, Pseudocurculionites, Pseudocymindis, Pseudocyphon, Pseudoelateropsis (syn. Elateropsis), Pseudohydrophilites, Pseudoprionites, Pseudotelephorus, Pseudotenebrio, Pseudus, Rhomaleus, Rhynchophori, Saperda valdensis, Saprophagi, Scalopoides, Semiglobus, Sideriosemion, Silphidium (syn. Silphites), Sitonites, Smodicoptera, Sojanocoleidae, Sphaericites, Staphyliniae, Stenelytron, Sternoxi, Stictulus, Strongylites, Strullendorfia, Tachypus, Telephorium, Tenebrionites, Tetragonides, Thurmannia, Toksunius, Tolype, Triadocoleopteron, Trialarva, Trichelepturgetes, Tridigitati, Tripsalis, Trixagites, Trochiscites, Trochmalus, Trogossitarii, Trypocoleus, Tshekardocoleidae (syn. Moravocoleidae, Uralocoleidae), Tychon, Ulomites, Velenovskya, Versicolores, Wollastonia, Xenogyrinus, Xinbinia, Xylophagi, Xylotupia, Yanqingia, Zacoleoptera, Zetemenos
Synonyms
|
Synonymy list
| Year | Name and author |
|---|---|
| 1758 | Coleoptera Linnaeus p. 341 |
| 1759 | Coleoptera Schreber p. 7 |
| 1761 | Coleoptera Linnaeus p. 133 |
| 1762 | Coleoptera Geoffroy |
| 1762 | Cistela Geoffroy p. 115 |
| 1763 | Coleoptera Scopoli p. 1 |
| 1764 | Coleoptera Müller p. 1 |
| 1766 | Coleoptera Schaeffer |
| 1767 | Coleoptera Linnaeus p. 537 |
| 1768 | Coleoptera Strøm p. 313 |
| 1770 | Coleoptera Drury |
| 1771 | Coleoptera Forster p. 1 |
| 1771 | Cistela Forster p. 12 |
| 1775 | Eleuterata Fabricius p. 1 |
| 1776 | Coleoptera Müller p. 52 |
| 1777 | Coleoptera Goeze p. 1 |
| 1777 | Cistela Goeze p. 151 |
| 1796 | Coleoptera Latreille p. 1 |
| 1798 | Eleuterata Fabricius p. 1 |
| 1798 | Eleutherata Illiger p. 497 |
| 1813 | Criocerina pristina Germar p. 14 |
| 1852 | Curculionites liasinus Heer p. 15 fig. 39,40 |
| 1856 | Necticus minutus Aymard p. 234 |
| 1856 | Ancylocheira liasina Giebel p. 82 |
| 1856 | Hydrobius obsoletus Heer p. 18 figs. Pl 1, fig 19 |
| 1862 | Hydrophilus stenopterus Heer p. 65 figs. Pl 4, fig 13 |
| 1862 | Helophorus magnus Heer p. 70 figs. Pl 5, figs 22,22b |
| 1862 | Hydrobius godeti Heer p. 70 figs. Pl 5, figs 21,21b |
| 1862 | Helophorus exilis Heer p. 71 figs. Pl 5, fig 20 |
| 1864 | Hydrophilites interpunctatus Heer p. 64 |
| 1864 | Strongylites laevigatus Heer p. 89 |
| 1864 | Saperda valdensis Heer p. 376 fig. 254 |
| 1867 | Helops atticus Redtenbacher p. 33 figs. Pl XVI fig 18 |
| 1874 | Hydrobius obsoletus Oustalet p. 128 figs. Pl 6, fig 18 |
| 1874 | Onthophagus luteus Oustalet p. 194 figs. Pl 2, fig 17 |
| 1879 | Carabites depressus Heer p. 91 |
| 1879 | Buprestites scabriusculus Heer p. 92 |
| 1888 | Criocerina pristina von Schlechtendal p. 478 |
| 1891 | Hylesinus lineatus Förster p. 401 figs. Pl 12, fig 16 |
| 1907 | Crioceris pristina Handlirsch p. 792 |
| 1914 | Isomira aurora Wickham p. 268 figs. Pl 5, fig 6 |
| 1921 | Coleoptera Cockerell |
| 1929 | Elateridopsidae Zalessky |
| 1932 | Elateridopsidae Zalessky p. 201 |
| 1939 | Hydrous ebeninus Statz p. 64 figs. Pl. 18, fig. 81 |
| 1940 | Euchroma arvernica Piton pp. 181-182 figs. Pl XVII fig 7 |
| 1940 | Acmaeodera perreaui Piton pp. 193-194 figs. Pl XVII fig 5 |
| 1940 | Megopis lineolatus Piton p. 209 fig. 76 |
| 1953 | Coleoptera Bode |
| 1967 | Cydnopsis nigromembranacea Jordan p. 86 figs. Pl 3, fig 13 |
| 1982 | Leptopodocoleidae Hong p. 118 |
| 1987 | Coleoptera Geyer and Kleber p. 342 |
| 1989 | Coleoptera Shoshani et al. p. 435 |
| 1989 | Coleoptera Zhang p. 420 |
| 1992 | Coleoptera Carpenter |
| 1992 | Hylesinus lineatus Wood and Bright p. 81 |
| 1995 | Coleoptera Lawrence and Newton, Jr. |
| 1997 | Eodromeus viriosus Zhang p. 88 figs. Pl 3, fig 3; text-fig 10 |
| 2004 | Coleoptera Kukalová-Peck and Lawrence |
| 2006 | Brochocoleus indibili Soriano and Delclos |
| 2008 | Helops atticus Kirejtshuk et al. p. 135 |
| 2011 | Coleoptera Beutel et al. |
| 2014 | Coleoptera Misof et al. |
| 2014 | Coleoptera Peters et al. |
| 2018 | Isomira aurora Bousquet et al. p. 381 |
| 2022 | Eucoleoptera Cai et al. |
| 2022 | Metaxycoleoptera Cai et al. |
| 2022 | Zacoleoptera Cai et al. |
| 2022 | Coleoptera Engel |
| 2022 | Eucoleoptera Engel et al. |
| 2022 | Metaxycoleoptera Engel et al. |
| 2022 | Zacoleoptera Engel et al. |
Is something missing? Join the Paleobiology Database and enter the data
|
|
If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Or. Coleoptera Linnaeus 1758 [beetle]
show all | hide all
G. †Allognosis Handlirsch 1906
+
†Allognosis nitens Geinitz 1894
†Allognosis striatus Ponomarenko 1985
Subor. †Alphacoleoptera Engel et al. 2022
+
Superfm. †Permocupedoidea Martynov 1933
+
Fm. †Labradorocoleidae Ponomarenko 1969
+
Fm. †Oborocoleidae Kukalová 1968
+
Invalid names: Tshekardocoleoidea Rodendorf 1944 [synonym]
Superfm. †Permosynoidea Tillyard 1924
+
Fm. †Permosynidae Tillyard 1924
+
G. †Alveolicupes Lin 1983
G. †Artematopodites Ponomarenko 1990
+
†Artematopodites crispulus Yan 2010
†Artematopodites latissimus Ponomarenko 2014
†Artematopodites latus Ponomarenko 1990
†Artematopodites lepidus Yan 2010
†Artematopodites leskoviensis Yan and Zhang 2010
+
Invalid names: Artematopodites longus Ponomarenko 1990 [replaced]
†Artematopodites longus Hong 1983
†Artematopodites lozowskii Ponomarenko 2015
†Artematopodites major Ponomarenko 1990
†Artematopodites maximus Yan 2010
†Artematopodites prolixus Zhang 1997
†Artematopodites propinquus Zhang 1997
†Artematopodites shaanbeiensis Hong 1995
G. †Delpuentesyne Martins-Neto and Gallego 2007
+
†Delpuentesyne menendezi Martins-Neto and Gallego 2007
G. †Diarcuipenna Lin 1986
+
†Diarcuipenna bennettitophila Ponomarenko 2011
†Diarcuipenna heterosa Lin 1986
G. †Dinoharpalus Handlirsch 1906
+
†Dinoharpalus coptoclavoides Ponomarenko 2015
†Dinoharpalus latus Ponomarenko 2015
†Dinoharpalus liasinus Giebel 1856
†Dinoharpalus martynovi Ponomarenko 2018
†Dinoharpalus rugosus Ponomarenko 1985
G. †Dzeregia Ponomarenko 1985
+
†Dzeregia ampla Ponomarenko 1990
†Dzeregia byrrhoides Ponomarenko 1990
†Dzeregia crassa Ponomarenko 1990
†Dzeregia juxta Lin 1986
†Dzeregia lata Ponomarenko 1985
†Dzeregia longa Ponomarenko 1985
†Dzeregia pilula Ponomarenko 1990
†Dzeregia platis Ponomarenko 2014
†Dzeregia striata Ponomarenko 1985
G. †Hydrobiites Heer 1864
+
†Hydrobiites anglicus Handlirsch 1906
†Hydrobiites bellus Geinitz 1894
†Hydrobiites convexus Ponomarenko 1985
†Hydrobiites crassus Ponomarenko 1985
†Hydrobiites dobbertinensis Handlirsch 1939
†Hydrobiites handlirschi Ponomarenko 2011
†Hydrobiites liasinus Giebel 1856
†Hydrobiites longus Ponomarenko 1990
†Hydrobiites minor Ponomarenko 2014
†Hydrobiites minutissimus Ponomarenko 1985
†Hydrobiites mongolicus Ponomarenko 2014
†Hydrobiites permianus Ponomarenko 2018
†Hydrobiites punctatostriatus Handlirsch 1939
†Hydrobiites purbeccensis Giebel 1856
†Hydrobiites sulcatus Ponomarenko 1986
†Hydrobiites tillyardi Ponomarenko 2011
†Hydrobiites veteranus Heer 1852
†Hydrobiites vladimiri Ponomarenko 2011
G. †Ischichucasyne Martins-Neto and Gallego 2005
+
†Ischichucasyne cladocosta Martins-Neto and Gallego 2009
†Ischichucasyne santajuanaensis Martins-Neto and Gallego 2005
G. †Palademosyne Rohdendorf 1961
+
†Palademosyne martynovae Rohdendorf 1961
†Palademosyne natalensis Ponomarenko 2005
G. †Permosyne Tillyard 1924
+
†Permosyne affinis Tillyard 1924
†Permosyne belmontensis Tillyard 1924
†Permosyne dentata Ponomarenko 2003
†Permosyne elongata Ponomarenko 2005
†Permosyne mitchelli Tillyard 1924
†Permosyne rasnitsyni Ponomarenko 2011
G. †Platycrossos Dunstan 1923
+
†Platycrossos caroli Ponomarenko 2011
†Platycrossos elongatus Ponomarenko 1985
†Platycrossos latus Ponomarenko 2014
†Platycrossos ligulatus Dunstan 1923
†Platycrossos longus Ponomarenko 2014
†Platycrossos loxonicus Ponomarenko 2014
†Platycrossos mongolicus Ponomarenko 2014
†Platycrossos ovum Ponomarenko 2014
†Platycrossos petalus Ponomarenko 1985
†Platycrossos punctatus Ponomarenko 1985
†Platycrossos subtumidus Dunstan 1923
†Platycrossos tumidus Tillyard 1916
G. †Tychtocoleus Rohdendorf 1961
+
†Tychtocoleus europeus Ponomarenko 2003
†Tychtocoleus neuburgae Rohdendorf 1961
†Tychtocoleus popovi Ponomarenko 2018
G. †Aptilotitus Carpenter 1986
+
†Aptilotitus capitecarens Bode 1953
Invalid names: Aptilotus Bode 1953 [replaced]
G. †Auchenophorites Carpenter 1986
+
†Auchenophorites sculpturatus Bode 1953
Invalid names: Auchenophorus Bode 1953 [replaced]
G. †Cardioides Bode 1953
G. †Cardiosyne Martins-Neto and Gallego 2006
+
†Cardiosyne elegans Martins-Neto and Gallego 2006
†Cardiosyne obesa Martins-Neto and Gallego 2006
G. †Chimaerocoleites Ponomarenko 1985
+
†Chimaerocoleites gyrinoides Ponomarenko 1985
†Chimaerocoleites parvulus Ponomarenko 1985
†Chimaerocoleites punctatus Ponomarenko 1985
†Chimaerocoleites seriatus Ponomarenko 1985
G. †Cistelites Heer 1864
+
†Cistelites insignis Heer 1864
†Cistelites longipes Hong 1985
†Cistelites minor Heer 1874
†Cistelites punctulatus Heer 1869
†Cistelites sachalinensis Heer 1878
†Cistelites spectabilis Heer 1869
G. †Clinomerus Carpenter 1986
+
†Clinomerus laticollis Handlirsch 1939
Invalid names: Catomerus Handlirsch 1939 [replaced]
G. †Coleopteron Handlirsch 1906
+
†Coleopteron antiquus Weyenbergh 1869
†Coleopteron aquaticus Brodie 1845
†Coleopteron beyrichi Giebel 1856
†Coleopteron bractoides Blake 1876
†Coleopteron lentissimus Weyenbergh 1874
†Coleopteron rugosostriatus Giebel 1856
†Coleopteron semicrematus Jarzembowski 2003
†Coleopteron sibiricus Heer 1876
†Coleopteron striatus Oppenheim 1888
†Coleopteron studeri Giebel 1856
†Coleopteron vetustus Giebel 1856
†Coleopteron wittsi Brodie 1845
†Coleopteron zekelii Giebel 1856
Invalid names: Coleopteron harpalinus Heer 1864 [nomen nudum], Coleopteron spectabilis Heer 1864 [nomen nudum], Coleopteron troglodytes Heer 1864 [nomen nudum]
G. †Cretohelophorus Ponomarenko 1987
+
†Cretohelophorus cupedoides Ponomarenko 1987
†Cretohelophorus yanensis Ren 1995
G. †Critoderma Carpenter 1986
+
†Critoderma deplanatum Heer 1864
Invalid names: Cycloderma Heer 1864 [replaced]
G. †Cryptauchenia Bode 1953
G. †Curculidium Handlirsch 1907
G. †Cyphospheron Bode 1953
G. †Darbiopelta Ponomarenko 1986
+
†Darbiopelta crassa Ponomarenko 1986
†Darbiopelta dubia Ponomarenko 1986
G. †Dicyphelus Carpenter 1986
+
†Dicyphelus concameratus Bode 1953
Invalid names: Dicyphus Bode 1953 [replaced]
G. †Doggeria Handlirsch 1906
+
†Doggeria bucklandi Mantell 1844
†Doggeria murchisoni Handlirsch 1906
†Doggeria sibirica Handlirsch 1906
G. †Etheridgea Handlirsch 1906
+
†Etheridgea australis Handlirsch 1906
†Etheridgea petrica Tillyard 1916
G. †Eurynotellus Carpenter 1986
+
†Eurynotellus brevicollis Bode 1953
Invalid names: Eurynotus Bode 1953 [replaced]
G. †Flichea Handlirsch 1906
+
†Flichea elongata Ponomarenko 1985
†Flichea lotharingiaca Fliche 1901
†Flichea sibirica Ponomarenko 1985
G. †Glaphyropterula Handlirsch 1906
+
†Glaphyropterula gracilis Heer 1852
†Glaphyroptula anglica Cockerell 1915
†Glaphyroptula liasina Giebel 1856
G. †Heeriaopsis Carpenter 1986
+
†Heeriaopsis laevigatum Heer 1878
Invalid names: Heeriella Handlirsch 1906 [replaced]
G. †Helopidium Westwood 1854
+
†Helopidium brodiei Giebel 1856
†Helopidium dubium Handlirsch 1906
†Helopidium dunkeri Giebel 1856
†Helopidium neoridas Westwood 1854
†Helopidium rugosum Handlirsch 1906
†Helopidium westwoodi Giebel 1856
G. †Hydrophilites Heer 1864
+
†Hydrophilites acherontis Heer 1852
†Hydrophilites naujatensis Heer 1883
†Hydrophilites stygius Heer 1864
G. †Keleusticus Handlirsch 1906
+
†Keleusticus acutus Ponomarenko 1985
†Keleusticus duplicatus Ponomarenko 1985
†Keleusticus zirkeli Geinitz 1894
G. Kryptophagus Herbst 1792
G. †Leptilostis Ponting 2024
+
†Leptilostis parvula Hong 1982
Invalid names: Tetillopsis Hong 1982 [replaced]
Unr. Mesocoleoptera Boudinot et al. 2022
+
Unr. Metacoleoptera Boudinot et al. 2022
+
Fm. †Ademosynidae Ponomarenko 1968
+
G. †Ademosyne Handlirsch 1906
+
†Ademosyne adunca Dunstan 1923
†Ademosyne arcucciae Martins-Neto and Gallego 2006
†Ademosyne australiensis Tillyard 1916
†Ademosyne bacca Ponomarenko 1969
†Ademosyne brevis Dunstan 1923
†Ademosyne cameroni Tillyard 1916
†Ademosyne congener Tillyard 1916
†Ademosyne curvata Dunstan 1923
†Ademosyne elliptica Ponomarenko 1969
†Ademosyne elongatus Martins-Neto and Gallego 2006
†Ademosyne hexacostata Martins-Neto and Gallego 2006
†Ademosyne intermedia Dunstan 1923
†Ademosyne kirghizica Ponomarenko 1969
†Ademosyne lata Dunstan 1923
†Ademosyne llantenesensis Brauckmann et al. 2010
†Ademosyne major Handlirsch 1906
†Ademosyne olliffi Handlirsch 1906
†Ademosyne parva Dunstan 1923
†Ademosyne polyzetete Ponomarenko 2011
†Ademosyne prisca Riek 1974
†Ademosyne punctata Tillyard 1916
†Ademosyne punctuada Martins-Neto and Gallego 2006
†Ademosyne ramocostata Dunstan 1923
†Ademosyne reducta Riek 1976
†Ademosyne rosenfeldi Brauckmann et al. 2010
†Ademosyne rugulosa Dunstan 1923
†Ademosyne sibirica Rohdendorf 1961
†Ademosyne speciosa Riek 1974
†Ademosyne umutu Martins-Neto and Gallego 2009
†Ademosyne vittamargina Dunstan 1923
†Ademosyne wianamattensis Tillyard 1918
G. †Dolichosyne Ponomarenko 1969
+
†Dolichosyne confragosa Ponomarenko 1969
†Dolichosyne rostrata Ponomarenko 1969
†Dolichosyne sulcata Ponomarenko 1969
Subor. Adephaga Clairville 1806
+
G. †Amphoxyne Bode 1953
+
†Amphoxyne darbiensis Ponomarenko 1986
†Amphoxyne minuta Bode 1953
+
Invalid names: Coilotrachelus lineatus Bode 1953 [synonym], Cricotrachelites rotundatus Bode 1953 [synonym], Loxoncus procerus Bode 1953 [synonym]
Invalid names: Coilotrachelus Bode 1953 [synonym], Cricotrachelites Bode 1953 [synonym], Loxostelidotus Bode 1953 [synonym]
Fm. Carabidae Latreille 1802 [ground beetle]
+
G. †Antemnacrassa Gomez Pallerola 1979
G. Apotomopterus Hope 1838
G. †Ataktosites Haupt 1956
G. Bembidium Latreille 1802
Subfm. Brachininae Bonelli 1810
G. †Brachinites Fritsch 1882
Subfm. Broscinae Hope 1838
G. †Carabidium Westwood 1854
Subfm. Carabinae Latreille 1802
G. †Carabites Heer 1852
G. †Carabites Ponomarenko 1977
G. †Carabopteron Martynov 1926
G. †Cavicarabus Hong 1985
Unr. Celeripedes Latreille 1802
G. †Cicindelites Meunier 1898
Subfm. †Conjunctiinae Ponomarenko 1977
Subfm. Elaphrinae Latreille 1802
G. †Elaphrotites Haupt 1956
G. †Fangshania Hong 1981
G. †Feronites Frič 1884
G. †Fonsecacarabus Martins-Neto and Mendes 2002
Subfm. Gehringiinae Darlington 1933
G. †Glenopterus Heer 1847
G. †Hadrocephalus Handlirsch 1906
G. †Harpalomimes Handlirsch 1906
G. †Hebeicarabus Hong 1984
G. †Lithocantharis Lin 1997
Subfm. Loricerinae Bonelli 1810
G. †Meileyingia Hong 1987
G. †Metagrilium Handlirsch 1906
Subfm. Migadopinae Chaudoir 1861
Subfm. Nebriinae Laporte 1834
G. †Nebrorabus Ponomarenko 1989
G. †Neothanes Scudder 1890
G. †Notokistus Handlirsch 1906
Subfm. Omophroninae Bonelli 1810
Subfm. Platyninae Bonelli 1810
G. †Procarabites Handlirsch 1906
Subfm. Protopaussinae Gestro 1892
Subfm. †Protorabinae Ponomarenko 1977
G. †Rhopalochlaenius Zhang et al. 1994
G. †Sinocarabus Hong 1982
G. Stereocerus Kirby 1837
G. †Sunocarabus Hong 1987
G. †Trechoides Motschulsky 1856
G. †Tremembecarabus Martins-Neto 1998
G. †Wongyekokia Lin 1997
G. †Wuchangicarabus Hong 1985
G. †Yunnanocarabus Lin 1977
Invalid names: Cratacanthus florissantensis Wickham 1917 [nomen dubium], Fossores Latreille 1802 [empty], Loricera exita Scudder 1900 [nomen dubium], Rhysodinae Laporte 1840 [empty], Siagoninae Bonelli 1813 [empty], Trechus fractus Wickham 1912 [nomen dubium]
Fm. Cicindelidae Latreille 1802 [tiger beetle]
+
Tr. Cicindelini Latreille 1802
G. †Cretotetracha Zhao et al. 2019
G. †Goriresina Matalin et al. 2021
G. †Hujia Song et al. 2022
Tr. Megacephalini Laporte 1834
G. †Opsis Handlirsch 1906
G. †Oxycheilopsis Cassola and Werner 2004
G. †Palaeopronyssiformia Wiesner et al. 2017
Invalid names: Oxycheilini Thomson 1857 [empty]
Fm. †Coptoclavidae Ponomarenko 1961
+
G. †Agrascapha Lin 1992
G. †Amblycephalonius Bode 1953
+
Invalid names: Barocephalus Carpenter 1986 [synonym], Barycephalus Bode 1953 [synonym], Tricyrtus Bode 1953 [synonym]
Subfm. †Charonoscaphinae Ponomarenko 1977
G. †Coptoclavia Ping 1928
Subfm. †Coptoclavinae Ponomarenko 1961
Subfm. †Coptoclaviscinae Soriano et al. 2007
G. †Euroscapha Lin 1992
Subfm. †Hispanoclavinae Soriano et al. 2007
Fm. Dytiscidae Leach 1815 [predaceous diving beetle]
+
Subfm. Agabinae Thomson 1867
G. †Angaragabus Ponomarenko 1963
Subfm. Colymbetinae Erichson 1837
Subfm. Copelatinae Van den Branden 1885
Subfm. Coptotominae Van den Branden 1885
G. †Cretodytes Ponomarenko 1977
Subfm. Cybisterinae Sharp 1880
Subfm. Dytiscinae Leach 1815
Subfm. Hydroporinae Aubé 1836
Subfm. †Liadytiscinae Prokin and Ren 2010
G. †Palaeodytes Ponomarenko 1987
G. †Sinoporus Prokin and Ren 2010
Invalid names: Aubehydrinae Guignot 1942 [empty], Necticus palustris Aymard 1856 [nomen nudum]
Fm. Gyrinidae Latreille 1810 [whirligig beetle]
+
G. †Anagyrinus Handlirsch 1906
G. †Coptogyrinus Handlirsch 1906
Subfm. Gyrininae Latreille 1810
G. †Gyrinopsis Handlirsch 1906
G. †Protogyrininus Hatch 1927
G. †Tunguskagyrus Yan et al. 2018
Invalid names: Spanglerogyrinae Folkerts 1979 [empty]
Subfm. Haliplidae Aubé 1836 [crawling water beetle]
+
G. †Cretihaliplus Ren et al. 1995
G. Haliplus Latreille 1802
G. Peltodytes Regimbart 1878
G. †Memptus Handlirsch 1906
+
†Memptus ganglbaueri Ponomarenko 1985
†Memptus guttula Ponomarenko 1985
†Memptus handlirschi Ponomarenko 1985
Fm. Noteridae Thomson 1860 [burrowing water beetle]
+
Subfm. Noterinae Thomson 1860
Subfm. Phreatodytinae Ueno 1957
Fm. Trachypachidae Thomson 1857 [false ground beetle]
+
G. †Apermunda Kirejtshuk and Ansorge 2022
G. †Denudirabus Ren 1995
G. †Dundorabus Ponomarenko 1989
Subfm. †Eodromeinae Ponomarenko 1977
G. †Eodromeites Ponomarenko 1985
Subfm. Trachypachinae Thomson 1857
Invalid names: Karadromeus xiangfanggouensis Hong 1995 [nomen dubium]
Fm. †Triaplidae Ponomarenko 1977
+
G. †Avocatinus Ponomarenko 1969
G. †Catinius Ponomarenko 1968
G. †Catinoides Ponomarenko 1969
G. †Clypeus Hong 1983
G. †Macrocatinius Ponomarenko 1969
G. †Tomiaplus Ponomarenko 2021
G. †Triaplus Ponomarenko 1977
G. †Triassocatinius Ponomarenko 1969
Invalid names: Catiniidae Ponomarenko 1968 [replaced]
Invalid names: Amphizoidae Leconte 1853 [empty], Dytiscoidea Leach 1815 [invalid subgroup], Pseudomorphinae Hope 1838 [empty], Tritarsusidae Hong 2002 [invalid subgroup]
Subor. Archostemata Kolbe 1908
+
G. †Argentinocupes Martins-Neto and Gallego 2006
+
†Argentinocupes abdalai Martins-Neto and Gallego 2006
†Argentinocupes pulcher Martins-Neto and Gallego 2006
†Argentinocupes sara Martins-Neto and Gallego 2009
G. †Brachilatus Strelnikova and Yan 2023
+
†Brachilatus caducus Ponomarenko 1969
†Brachilatus dundulaensis Ponomarenko 1994
†Brachilatus foersteri Ponomarenko 1971
+
Invalid names: Notocupes reticulatus Ponomarenko 1968 [replaced]
†Brachilatus longicoxa Soriano and Delclos 2006
†Brachilatus nigrimonticola Ponomarenko 1968
†Brachilatus oculatus Soriano and Delclos 2006
†Brachilatus premeris Lee et al. 2022
†Brachilatus spinosus Li and Cai 2023
†Brachilatus viridis Soriano and Delclos 2006
G. †Conexicoxa Lin 1986
+
†Conexicoxa brachycephala Ponomarenko 1994
†Conexicoxa crassa Ponomarenko 1969
†Conexicoxa denticollis Jiang et al. 2020
†Conexicoxa epicharis Tan et al. 2005
†Conexicoxa eumeura Tan et al. 2006
†Conexicoxa exigua Ponomarenko 1994
†Conexicoxa homora Lin 1986
†Conexicoxa jurassica Hong 1983
†Conexicoxa khetanensis Ponomarenko 1993
†Conexicoxa kirghizica Ponomarenko 1969
†Conexicoxa longicollis Ponomarenko 1994
†Conexicoxa martinclosas Soriano and Delclos 2006
†Conexicoxa porrecta Tan et al. 2006
†Conexicoxa psilata Tan et al. 2005
†Conexicoxa siniestri Soriano and Delclos 2006
†Conexicoxa stabilis Tan et al. 2005
†Conexicoxa tripartita Oppenheim 1888
Superfm. Cupedoidea Laporte 1836
+
Fm. Cupedidae Laporte 1836 [reticulated beetle]
G. †Magnocoleus Hong 1998
Fm. Micromalthidae Barber 1913 [telephone pole beetle]
Fm. Ommatidae Sharp and Muir 1912
Invalid names: Crowsoniellidae Iablokoff-Khnzorian 1983 [empty], Magnocoleidae Hong 1998 [invalid subgroup]
G. †Notocupes Ponomarenko 1964
+
†Lupicupes trachylaenus Ren 1995
†Notocupes alienus Tan and Ren 2006
†Notocupes baojiatunensis Hong 1992
†Notocupes caudatus Ponomarenko 1966
†Notocupes cyclodontus Tan et al. 2006
†Notocupes daohugouensis Li and Cai 2023
†Notocupes diazromerali Soriano and Delclos 2006
†Notocupes elegans Ponomarenko 1994
†Notocupes excellens Ponomarenko 1966
†Notocupes kezuoensis Hong 1987
†Notocupes khasurtyiensis Strelnikova 2019
†Notocupes laetus Lin 1976
†Notocupes laiyangensis Hong and Wang 1990
†Notocupes lapidarius Ponomarenko 1968
†Notocupes latus Ponomarenko 1969
†Notocupes lini Ponomarenko et al. 2012
†Notocupes ludongensis Wang and Liu 1996
†Notocupes mongolicus Ponomarenko 1994
†Notocupes neli Tihelka et al. 2019
†Notocupes ohmkuhnlei Jarzembowski et al. 2019
†Notocupes picturatus Ponomarenko 1964
†Notocupes pingi Ponomarenko and Ren 2010
†Notocupes pulcher Ponomarenko 1968
†Notocupes reticulatus Oppenheim 1888
†Notocupes robustus Li and Cai 2023
†Notocupes rudis Tan et al. 2005
†Notocupes shiluoensis Hong 1984
†Notocupes tuanwangensis Hong and Wang 1990
†Notocupes validus Lin 1976
Invalid names: Amblomma Tan et al. 2005 [synonym], Chengdecupes Hong 1983 [synonym], Forticupes Hong and Wang 1990 [synonym], Lupicupes Ren 1995 [synonym], Ovatocupes Tan and Ren 2006 [synonym], Picticupes Hong and Wang 1990 [synonym], Sinocupes Lin 1976 [synonym]
G. †Pseudosilphites Zeuner 1930
+
†Pseudosilphites natalensis Zeuner 1960
†Pseudosilphites triassicus Zeuner 1930
G. †Rhabdocupes Ponomarenko 1966
+
†Rhabdocupes baculatus Ponomarenko 1969
†Rhabdocupes cellulosus Ponomarenko 1969
†Rhabdocupes dischides Zhang 1986
†Rhabdocupes issykkulensis Ponomarenko 1969
†Rhabdocupes laticella Ponomarenko 1969
†Rhabdocupes lentus Ren 1995
†Rhabdocupes longus Ponomarenko 1966
†Rhabdocupes minisculus Tan et al. 2006
†Rhabdocupes minor Ponomarenko 1966
†Rhabdocupes oxypygus Ponomarenko 1969
†Rhabdocupes protensus Tan et al. 2006
†Rhabdocupes rostratus Ponomarenko 1969
†Rhabdocupes sogutensis Ponomarenko 1969
†Rhabdocupes tenuis Ponomarenko 1969
†Rhabdocupes vitimensis Ponomarenko 1966
Superfm. †Schizocoleoidea Rohdendorf 1961
+
Fm. †Phoroschizidae Bouchard and Bousquet 2020
+
Invalid names: Schizophoridae Ponomarenko 1968 [replaced]
Fm. †Schizocoleidae Rohdendorf 1961
Invalid names: Schizophoroidea Ponomarenko 1968 [synonym]
G. †Tentyridium Westwood 1854
+
†Tentyridium dilatatum Handlirsch 1906
†Tentyridium peleus Westwood 1854
G. †Tillyardiopsis Dunstan 1923
+
†Tillyardiopsis granulata Dunstan 1923
†Tillyardiopsis tuberculata Dunstan 1923
†Tillyardiopsis variotuberculata Dunstan 1923
Invalid names: Cupediformia [invalid subgroup], Moravocoleus perditus Kukalová 1968 [nomen dubium], Notocupedini Ponomarenko 1966 [invalid subgroup], Priacma corrupta Ponomarenko 1986 [nomen dubium], Priacma longicapitis Ponomarenko 1997 [nomen dubium], Priacma oculata Ponomarenko 1997 [nomen dubium], Priacma striata Ponomarenko 2000 [nomen dubium]
Fm. †Asiocoleidae Rohdendorf 1961
+
G. †Bicoleus Ponomarenko 2013
+
†Bicoleus kuplensis Ponomarenko 2013
†Bicoleus laticella Ponomarenko 2013
G. †Loculitricoleus Tan and Ren 2009
+
†Loculitricoleus flatus Tan and Ren 2009
†Loculitricoleus mongolicus Ponomarenko 2014
†Loculitricoleus tenuatus Tan and Ren 2009
G. †Schizotaldycupes Rohdendorf 1961
+
†Schizotaldycupes ananjevi Rohdendorf 1961
†Schizotaldycupes beckermigdisovae Rohdendorf 1961
†Schizotaldycupes pubescens Ponomarenko 2013
Invalid names: Carinicupes Rohdendorf 1961 [synonym]
G. †Tetracoleus Ponomarenko 2008
+
†Tetracoleus aristovi Ponomarenko 2018
†Tetracoleus golubevi Ponomarenko 2015
†Tetracoleus meyeni Ponomarenko 2013
†Tetracoleus minimus Ponomarenko 2008
†Tetracoleus orenburgensis Ponomarenko 2013
†Tetracoleus sennikovi Ponomarenko 2011
†Tetracoleus tricoleoides Ponomarenko 2013
†Tetracoleus tshalyshevi Ponomarenko 2008
†Tetracoleus vasilenkoi Ponomarenko 2018
Subor. Myxophaga Crowson 1955
+
Superfm. Lepiceroidea Hinton 1936
+
Fm. Lepiceridae Hinton 1936
+
Invalid names: Cyathoceridae Scharp 1882 [synonym], Haplochelidae Kirejtshuk and Poinar 2006 [synonym]
Superfm. Sphaeriusoidea
+
Fm. Hydroscaphidae LeConte 1874 [skiff beetle]
Invalid names: Sphaeriidae Erichson 1845 [empty], Torridincolidae Steffan 1964 [empty]
Invalid names: Sphaeriiformia [empty]
Fm. †Peltosynidae Yan et al. 2017
+
G. †Peltosyne Ponomarenko 1977
+
†Peltosyne triassica Ponomarenko 1977
†Peltosyne varyvrosa Yan et al. 2017
Subor. Polyphaga Emery 1886
+
G. †Angarolarva Ponomarenko 1985
+
†Angarolarva aquatica Ponomarenko 1985
†Angarolarva incerta Ponomarenko 1990
Unr. Clambiformia Cai and Tihelka 2022
+
Superfm. Clamboidea Fischer von Waldheim 1821
+
Invalid names: Derodontoidea LeConte 1861 [synonym], Eucinetoidea Lacordaire 1857 [synonym]
G. †Cryptophagites Ponomarenko 1990
+
†Cryptophagites capitatus Ponomarenko 1990
†Cryptophagites elongatus Ponomarenko 1990
Infraor. Cucujiformia Lameere 1938
+
Superfm. Cleroidea Latreille 1802
Superfm. Coccinelloidea Latreille 1807
Superfm. Cucujoidea Latreille 1802
Superfm. Erotyloidea Latreille 1802
Superfm. Lymexyloidea Fleming 1821
Superfm. Nitiduloidea Latreille 1802
Fm. †Parandrexidae Kirejtshuk 1994
G. †Petratypus Kaddumi 2005
Unr. Phytophaga
Superfm. Tenebrionoidea Latreille 1802
Fm. †Wabbelidae Alekseev 2017
Infraor. Elateriformia Crowson 1960
+
Superfm. Buprestoidea Leach 1815
Superfm. Byrrhoidea Latreille 1804
Superfm. Cantharoidea
Superfm. Dascilloidea Gue'rin-Me'neville 1843
Superfm. Dryopoidea
Invalid names: Mesosperchus angulatus Ponomarenko 1985 [nomen dubium], Podabrocephalidae Pic 1930 [empty]
G. †Jingxidiscus Hong 1990
+
†Jingxidiscus lushangfenensis Hong 1981
Invalid names: Discus Hong 1981 [replaced]
Fm. Jurodidae Ponomarenko 1985
+
G. †Jurodes Ponomarenko 1985
Invalid names: Sikhotealiniidae Lafer 1996 [synonym]
G. †Larvula Ponomarenko 1985
+
†Larvula cassa Ponomarenko 1985
†Larvula triassica Prokin and Ponomarenko 2013
Fm. †Lasiosynidae Kirejtshuk et al. 2010
+
G. †Bupredactyla Kirejtshuk et al. 2010
G. †Crassisyne Yan et al. 2014
G. †Glaphyropterodes Handlirsch 1906
G. †Microsyne Yan et al. 2014
Invalid names: Lasiosyne laxa Yan et al. 2014 [nomen dubium]
G. †Plastelater Handlirsch 1906
+
†Plastelater glabratus Ponomarenko 1985
†Plastelater neptuni Giebel 1856
G. †Polysitum Dunstan 1923
+
†Polysitum elongatum Ponomarenko 1985
†Polysitum hosbayari Ponomarenko 1986
†Polysitum kuznetskiense Rohdendorf 1961
†Polysitum latum Ponomarenko 1985
†Polysitum majus Ponomarenko 1990
†Polysitum minutus Dunstan 1923
†Polysitum ovale Ponomarenko 1990
†Polysitum punctatus Dunstan 1923
†Polysitum rugosum Ponomarenko 1986
†Polysitum sharategense Ponomarenko 2014
†Polysitum wudenghaoensis Hong 1976
G. †Sphaerocantharis Bode 1953
+
†Sphaerocantharis defossa Bode 1953
†Sphaerocantharis sibirica Ponomarenko 1990
Invalid names: Mikrocarpides Bode 1953 [synonym], Theornithion Bode 1953 [synonym]
Infraor. Staphyliniformia Lameere 1900
+
Superfm. Histeroidea Gyllenhal 1808
Superfm. Hydrophiloidea Latreille 1802 [water scavenger beetle]
Superfm. Scarabaeoidea Latreille 1802 [scarab beetle]
Superfm. Staphylinoidea Latreille 1802
Invalid names: Scarabaeiformia Crowson 1960 [synonym]
Invalid names: Agyrtes primoticus Scudder 1900 [nomen dubium], Derodontiformia Le Conte 1861 [invalid subgroup], Escheria crassipunctata Förster 1891 [nomen dubium], Escheria dimidiata Förster 1891 [nomen dubium], Escheria punctulata Förster 1891 [nomen dubium], Eucinetiformia [empty], Limnius perrini Theobald 1937 [nomen dubium], Malachius vertummni Heer 1847 [nomen dubium], Mesosperchus notatus Ponomarenko 1977 [nomen dubium], Tomocoleites Hong 2002 [nomen nudum], Troglops punctatissima Théobald 1935 [nomen dubium]
Fm. †Ponomarenkiumidae Lawrence and Bouchard 2022
+
G. †Ponomarenkium Yan et al. 2018
+
†Ponomarenkium belmonthense Yan et al. 2017
Invalid names: Ponomarenkia Yan et al. 2017 [replaced]
Invalid names: Ponomarenkiidae Yan et al. 2017 [invalid subgroup]
Fm. †Taldycupedidae Rohdendorf 1961
+
G. †Afrotaldycupes Kirejtshuk 2020
+
†Afrotaldycupes africanus Ponomarenko 2005
†Afrotaldycupes lidgettoniensis Ponomarenko 2005
G. †Mesothoris Tillyard 1916
+
†Mesothoris clathrata Tillyard 1916
†Mesothoris grandis Dunstan 1923
†Mesothoris punctomarginum Dunstan 1923
G. †Proterocupes Ponomarenko 2015
+
†Proterocupes major Ponomarenko 2015
†Proterocupes nedubrovensis Ponomarenko 2015
†Proterocupes permiana Ponomarenko 2013
Invalid names: Lobanovia Kirejtshuk 2020 [objective synonym]
G. †Simmondsia Dunstan 1923
+
†Simmondsia cylindrica Dunstan 1923
†Simmondsia subpiriformis Dunstan 1923
G. †Taldycupes Rohdendorf 1961
+
†Taldycupes cellulosus Ponomarenko 2013
†Taldycupes khalfini Rohdendorf 1961
†Taldycupes moltshanovi Rohdendorf 1961
†Taldycupes pingi Ponomarenko et al. 2014
†Taldycupes reticulatus Ponomarenko 1969
†Taldycupes rosanovi Ponomarenko 2006
†Taldycupes vasjuchitshevi Rohdendorf 1961
Invalid names: Cryptocupes Rohdendorf 1961 [synonym], Taldycupidium Rohdendorf 1961 [synonym]
G. †Tecticupes Rohdendorf 1961
+
†Tecticupes heckeri Rohdendorf 1961
+
Invalid names: Stegocupes efremovi Rohdendorf 1961 [synonym], Stegocupes fedotovi Rohdendorf 1961 [synonym]
†Tecticupes indistinctus Rohdendorf 1961
+
Invalid names: Tychticupes tchihatcheffi Rohdendorf 1961 [synonym]
†Tecticupes martynovi Ponomarenko 2013
Invalid names: Stegocupes Rohdendorf 1961 [synonym]
G. †Tychticupes Rohdendorf 1961
+
†Tychticupes beljanini Rohdendorf 1961
†Tychticupes radtshenkoi Rohdendorf 1961
†Tychticupes ragozini Rohdendorf 1961
Invalid names: Tychticupes ussovi Rohdendorf 1961 [nomen dubium], Uskaticupes Rohdendorf 1961 [nomen dubium], Uskaticupes javorskyi Rohdendorf 1961 [nomen dubium]
Fm. †Triadocupedidae Ponomarenko 1966
+
G. †Cupesia Ponomarenko 1966
+
†Cupesia monilicornia Ponomarenko 1966
†Cupesia sepulta Ponomarenko 1969
†Cupesia serricornia Ponomarenko 1969
G. †Platycupes Ponomarenko 1966
+
†Platycupes dolichocerus Ponomarenko 1966
†Platycupes major Ponomarenko 1966
†Platycupes pusillus Ponomarenko 1966
†Platycupes reticulatus Ponomarenko 1966
G. †Triadocupes Ponomarenko 1966
+
†Triadocupes ellipticus Ponomarenko 1966
†Triadocupes ferghanensis Ponomarenko 1966
†Triadocupes latus Ponomarenko 1966
Fm. †Permocupedidae Martynov 1932
+
Subfm. †Permocupedinae Martynov 1932
+
G. †Afrocupes Geertsema and van den Heever 1996
+
†Afrocupes firmae Geertsema and van den Heever 1996
G. †Archicupes Rohdendorf 1961
+
†Archicupes jacobsoni Rohdendorf 1961
+
Invalid names: Palaeocupes kaltanicus Rohdendorf 1961 [synonym]
Invalid names: Palaeocupes Rohdendorf 1961 [synonym]
G. †Eocupes Rohdendorf 1961
+
†Eocupes cellulosus Ponomarenko 1969
†Eocupes lukjanovitshi Rohdendorf 1961
G. †Frankencupes Ponomarenko and Bashkuev 2018
+
†Frankencupes ultimus Ponomarenko and Bashkuev 2018
G. †Ichthyocupes Rohdendorf 1961
+
†Ichthyocupes kuznetskiensis Ponomarenko 1969
†Ichthyocupes skoki Rohdendorf 1961
†Ichthyocupes tyzhnovi Rohdendorf 1961
G. †Kaltanicupes Rohdendorf 1961
+
†Kaltanicupes acutus Ponomarenko 1963
†Kaltanicupes kargalensis Ponomarenko 1963
†Kaltanicupes kitjakensis Ponomarenko 1963
†Kaltanicupes richteri Rohdendorf 1961
G. †Permocupes Martynov 1932
+
†Permocupes distinctus Martynov 1932
†Permocupes grandis Ponomarenko 1963
†Permocupes latus Ponomarenko 2013
†Permocupes semenovi Martynov 1932
†Permocupes sojanensis Ponomarenko 1963
Invalid names: Permocupoides Martynov 1932 [synonym]
G. †Protocupoides Rohdendorf 1961
+
†Protocupoides acer Rohdendorf 1961
†Protocupoides carinatus Rohdendorf 1961
†Protocupoides elongatus Ponomarenko 2013
†Protocupoides esini Ponomarenko 2013
†Protocupoides plavilstshikovi Rohdendorf 1961
†Protocupoides sharovi Rohdendorf 1961
Invalid names: Protocupoides rohdendorfi Pinto 1987 [nomen dubium], Tomiocupes Rohdendorf 1961 [synonym], Tricupes Rohdendorf 1961 [synonym]
Invalid names: Protocupes Rohdendorf 1961 [nomen dubium], Taldycupes chachlovi Rohdendorf 1961 [nomen dubium]
Invalid names: Archicupes reichardti Rohdendorf 1961 [nomen dubium], Kaltanocoleidae Rohdendorf 1961 [synonym], Protocupes martynovi Rohdendorf 1961 [nomen dubium]
G. †Mesoncus Carpenter 1986
+
†Mesoncus striatulus Bode 1953
Invalid names: Loxoncus Bode 1953 [replaced]
G. †Mesorhynchophora Tillyard 1916
G. †Microcoleopteron Handlirsch 1906
+
†Microcoleopteron heydeni Weyenbergh 1869
†Microcoleopteron jurassicum Weyenbergh 1869
†Microcoleopteron lithographicum Weyenbergh 1869
†Microcoleopteron minimum Oppenheim 1888
Unr. Nubili Latreille 1804
+
Invalid names: Herbicolae Latreille 1804 [empty], Sabulosi Latreille 1804 [empty]
G. †Ooidellus Carpenter 1986
+
†Ooidellus denudatus Bode 1953
Invalid names: Ooides Bode 1953 [replaced]
Tr. †Oropseini Legalov and Kirejtshuk 2016
+
G. †Oxytoroptera Handlirsch 1939
+
†Oxytoroptera mediocris Handlirsch 1939
†Oxytoroptera punctata Ponomarenko 1985
Subfm. †Palaeotylinae Poinar et al. 2018
+
G. †Pantodapus Handlirsch 1906
+
†Pantodapus ewaldi Giebel 1856
†Pantodapus knorri Giebel 1856
†Pantodapus westwoodi Handlirsch 1906
G. †Parabuprestium Handlirsch 1906
+
†Parabuprestium pseudocarabus Handlirsch 1906
†Parabuprestium teleas Westwood 1854
G. †Pedinosoma Bode 1953
G. †Peridosoma Carpenter 1986
+
†Peridosoma praecisum Bode 1953
Invalid names: Perosoma Bode 1953 [replaced]
G. †Pseudoelateropsis Handlirsch 1906
+
†Pseudoelateropsis infraliassica Roemer 1876
Invalid names: Elateropsis Roemer 1876 [replaced]
G. †Pseudotelephorus Handlirsch 1906
+
†Pseudotelephorus grandis Cockerell 1915
†Pseudotelephorus haueri Giebel 1856
†Pseudotelephorus punctulatus Cockerell 1915
G. †Pseudotenebrio Handlirsch 1906
+
†Pseudotenebrio innominatus Weyenbergh 1869
†Pseudotenebrio relictus Weyenbergh 1869
G. †Semiglobus Handlirsch 1906
+
†Semiglobus chrysomelinus Handlirsch 1906
†Semiglobus jurassicus Handlirsch 1906
†Semiglobus neptuni Giebel 1856
G. †Silphidium Handlirsch 1907
+
†Silphidium priscum Fritsch 1882
Invalid names: Silphites Fritsch 1882 [replaced]
Fm. †Sojanocoleidae Martynov 1932
+
G. †Strongylites Heer 1864
+
†Strongulites latus Ponomarenko 1985
†Strongylites morio Heer 1864
†Strongylites stygius Heer 1864
Fm. Tenebrionites Latreille 1802 [darkling beetle]
+
Invalid names: Pimeliariae Latreille 1802 [empty], Tenebrionites Latreille 1802 [empty]
Fm. †Tshekardocoleidae Rohdendorf 1944
+
G. †Avocoleus Ponomarenko 1969
+
†Avocoleus fractus Kukalová 1968
†Avocoleus neglegens Kukalová 1968
G. †Sylvacoleus Ponomarenko 1963
+
†Sylvacoleus richteri Ponomarenko 1963
†Sylvacoleus sharovi Ponomarenko 1963
G. †Tshekardocoleus Rohdendorf 1944
+
†Tshekardocoleus magnus Rohdendorf 1944
†Tshekardocoleus minor Ponomarenko 1963
Invalid names: Moravocoleidae Kukalová-Peck and Beutel 2012 [synonym], Uralocoleidae Zalessky 1947 [synonym], Uralocoleus ultimus Ponomarenko 2000 [nomen dubium]
Invalid names: Acmaeodera perreaui Piton 1940 [nomen dubium], Ancylocheira liasina Giebel 1856 [nomen dubium], Brochocoleus indibili Soriano and Delclos 2006 [nomen dubium], Carabiques Latreille 1802 [empty], Cebrionates Latreille 1802 [empty], Chrysomelinae Latreille 1802 [empty], Cicindeletae Latreille 1802 [empty], Cistela Geoffroy 1762 [nomen oblitum], Clerii Latreille 1802 [empty], Coleopteron depressus Heer 1879 [nomen nudum], Coleopteron interpunctatus Heer 1864 [nomen nudum], Coleopteron laevigatus Heer 1864 [nomen nudum], Coleopteron scabriusculus Heer 1879 [nomen nudum], Coprophagi Latreille 1802 [empty], Criocerina pristina Germar 1813 [nomen dubium], Curculionites liasinus Heer 1852 [nomen dubium], Curculiopsidae Martynov 1937 [empty], Cydnopsis nigromembranacea Jordan 1967 [nomen dubium], Diaperialae Latreille 1802 [empty], Elateridopsidae Zalessky 1929 [invalid subgroup], Eleuterata Fabricius 1775 [synonym], Entomophagi Latreille 1804 [empty], Eodromeus viriosus Zhang 1997 [nomen dubium], Erotilenae Latreille 1802 [empty], Euchroma arvernica Piton 1940 [nomen dubium], Eucoleoptera Engel et al. 2022 [invalid subgroup], Helophorus exilis Heer 1862 [nomen dubium], Helophorus magnus Heer 1862 [nomen dubium], Helopii Latreille 1802 [empty], Helops atticus Redtenbacher 1867 [nomen dubium], Hydrobius godeti Heer 1862 [nomen dubium], Hydrobius obsoletus Heer 1856 [nomen dubium], Hydrocanthares Latreille 1802 [empty], Hydrophilus stenopterus Heer 1862 [nomen dubium], Hydrous ebeninus Statz 1939 [nomen dubium], Hylesinus lineatus Förster 1891 [nomen dubium], Ipidae Bedel 1888 [empty], Isomira aurora Wickham 1914 [nomen dubium], Leptopodocoleidae Hong 1982 [invalid subgroup], Macrogastri Latreille 1802 [empty], Malacodermi Latreille 1802 [empty], Malacodermidae [empty], Megopis lineolatus Piton 1940 [nomen dubium], Metaxycoleoptera Engel et al. 2022 [invalid subgroup], Necrophagi Latreille 1802 [empty], Necticus minutus Aymard 1856 [nomen nudum], Nitidulariae Latreille 1802 [empty], Onthophagus luteus Oustalet 1874 [nomen dubium], Palpatores Latreille 1802 [empty], Platyprosopi Latreille 1804 [empty], Protocoleoptera Tillyard 1924 [empty], Rhynchophori Latreille 1804 [empty], Saperda valdensis Heer 1864 [nomen dubium], Saprophagi Latreille 1804 [empty], Staphyliniae Latreille 1802 [empty], Sternoxi Latreille 1802 [empty], Tridigitati Latreille 1802 [empty], Versicolores Latreille 1804 [empty], Xylophagi Latreille 1802 [empty], Zacoleoptera Engel et al. 2022 [invalid subgroup]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
|
|
||||
|
|||||
|
|
||||
| Source: c = class, p = phylum | |||||
| References: Bush and Bambach 2015, Kiessling 2004 | |||||
Age range
Maximum range based only on fossils: base of the Asselian to the top of the Holocene or 298.90000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 293.52 Ma
Minimum age of oldest fossil (stem group age): 293.52 Ma
Collections (1997 total)
Oldest occurrences | |||
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| Asselian | Germany (Thüringer Wald) | Pachypleurites nodosus (128485) | |
| Asselian | Germany (Rhineland-Palatinate) | Coleopsis archaica (156248) | |
| Asselian | Czechia | Moravocoleus perditus, Tshekardocoleidae indet., Retelytron conopeum, Eocoleus scaber, Prosperocoleus prosperus, Moravocoleus fractus, Moravocoleus neglegens, Votocoleus submissus, Moravocoleus permianus, Boscoleus blandus, Umoricoleus perplex, Liberocoleus intactus, Oborocoleus rohdendorfi (114186) | |
| Artinskian | South Africa (Western Cape) | Afrocupes firmae (156986) | |
| Artinskian | USA (Oklahoma) | Permocoleus wellingtonensis (104816 104826) | |
| Kungurian | Russian Federation | Maricoleus valentinae (115214) Sylvacoleus richteri, Tshekardocoleus magnus (107787) Uralocoleus splendens (155819) | |
| Kungurian | Brazil (Rio Grande do Sul) | Protocupoides rohdendorfi, Kaltanicupes ponomarenkoi (116121) | |
| Kungurian | Russian Federation (Suksun) | Tshekardocoleidae indet., Sylvacoleus sharovi, Sylvacoleus richteri, Sylvacoleodes admirandus, Tshekardocoleus minor (108206) Tshekardocoleus magnus (116652) | |
| Roadian | Russian Federation (Arkangelsk) | Permocrossos elongatus (104993) Permocupes grandis, Permocupes sojanensis (116933) Permocupes semenovi (104716) Permocupes sojanensis, Uralocoleus ultimus (104992) Sojanocoleus reticulatus (116874) | |
| Roadian | Russian Federation (Kirov) | Kaltanicupes major, Kaltanicupes kitjakensis, Kaltanicupes acutus, Rossocoleus altus (105058) | |
| Roadian | Russian Federation (Kazan) | Permocupes semenovi, Permocupoides distinctus (116877) | |
| Roadian | Russian Federation | Elateridopsis permiensis (156163) Stegosyne rugosa (124611) | |
| Roadian | Russian Federation (Kemerovo) | Archicupes reichardti, Cytocupes angustus, Ichthyocupes tyzhnovi, Tomiocupes sharovi, Protocupoides plavilstshikovi, Palaeocupes kaltanicus, Kaltanocoleus pospelovi, Eocupes cellulosus, Schizotaldycupes ananjevi, Carinicupes beckermigdisovae, Asiocoleus novojilovi, Stegocupes efremovi, Uskatocoleus kaltanicus (104704) Ichthyocupes kuznetskiensis (122572) Kaltanicupes richteri (122568) Protocupes martynovi, Archicupes jacobsoni, Cytocupoides elongatus, Eocupes lukjanovitshi, Rossocoleus avus (122567) Protocupes martynovi, Kaltanocoleus pospelovi (122569) Schizocoleus kuznetskiensis (124604) | |
| Wordian | Russian Federation (Kemerovo) | Palademosyne martynovae (124612) Schizocupes obrutshevi (124600) Taldycupes chachlovi, Protocupes ogloblini, Tecticupes heckeri, Stegocupes fedotovi, Tecticupes indistinctus, Tychticupes tchihatcheffi, Uskatocoleus surijokovicus, Uskatocoleus iljinskiensis, Uskatocoleus avitus, Karakanocoleus janischewskyi, Rossocoleus latior, Rossocoleus surijokovensis, Rossocoleus latus, Rossocoleus grandis, Rossocoleus parvus (122566) Uskatocoleus convexus, Rossocoleus verrucosus (124603) | |
| Wordian | Russian Federation (Udmurtia) | Coleoptera hindwing indet., Permocupes latus, Protocupoides elongatus, Schizophoridae indet., Rossocoleus angustus, Rossocoleus novojilovi (103502) Schizotaldycupes pubescens, Rhombocoleites uralensis (104750) | |
| Wordian | South Africa | Permocupedidae indet. (226396) | |
| Wordian - Capitanian | Russian Federation | Uskatocoleus kushejakovicus (124610) | |
| Capitanian | Russian Federation (Orenburg) | Curculiopsis ellipticus (104994) Kaltanicupes kargalensis, Tecticupes martynovi, Rhombocoleites adumbratus, Kargalarva permosialis (104057) Schizocoleus major, Aenigmocoleus uralensis, Rossocoleus sakmara, Sakmaracoleus orenburgensis (152235) Tatarocupes granulatus, Tetracoleus orenburgensis, Bicoleus kuplensis, Schizocoleus longus, Schizocoleus minimus, Schizocoleus glabrus, Schizocoleus robustus, Erunakicupes brevis, Palademosyne ovum, Karakanocoleus europeus, Rhombocoleus gomankovi (104717) | |
| Capitanian | China (Anhui) | Linicupes yinpinensis, Asiocoleopsis hongi, Taldycupes pingi, Dikerocoleus divisus, Chaocoleus limnebius, Archosyne permiana (103239) | |
| Wuchiapingian | Russian Federation | Permocupoides skoki, Tomiocupes carinatus, Tricupes acer, Tychticupoides grjazevi, Taldycupes moltshanovi, Tychticupes vasjuchitshevi, Taldycupidium bergi, Cryptocupes rjabinini, Polysitum kuznetskiense, Erunakicupes schmalhauseni, Erunakicupes kryshtofovichi, Erunakicupes fomitshevi, Aenigmocoleus borissiaki, Uskatocoleus minimus, Uskatocoleus zalesskyi, Uskatocoleus lutugini, Uskatocoleus foveolatus, Karakanocoleus major, Karakanocoleodes latissimus, Erunakicupes venjukovi, Karakanocoleus lebedevi, Karakanocoleus striatus, Karakanocoleus magnus, Uskatocoleus angustus, Uskatocoleus angustior, Uskatocoleus longus, Rhombocoleus andreae, Ademosyne sibirica, Tychtocoleus neuburgae (124608) Permocupoides skoki, Tychticupes ussovi, Uskaticupes javorskyi, Taldycupes khalfini, Tychticupes ragozini, Taldycupes beljanini, Tychticupes radtshenkoi, Uskatocoleus zalesskyi, Tychtocoleus neuburgae (122565) | |