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Coleoptera (beetle)

Insecta - Coleoptera

Taxonomy
Coleoptera was named by Linnaeus (1758). It is extant.

It was assigned to Oligometabola by Geyer and Kleber (1987); to Endopterygota by Carpenter (1992); to Insecta by Linnaeus (1758), Schreber (1759), Linnaeus (1761), Geoffroy (1762), Scopoli (1763), Müller (1764), Schaeffer (1766), Linnaeus (1767), Strøm (1768), Drury (1770), Forster (1771), Müller (1776), Goeze (1777), Latreille (1796), Cockerell (1921), Bode (1953), Zhang (1989), Shoshani et al. (1989) and Lawrence and Newton (1995); and to Coleopterida by Kukalová-Peck and Lawrence (2004), Beutel et al. (2011), Peters et al. (2014), Misof et al. (2014) and Engel (2022).

Subtaxa
Acmaeodera perreaui, Adelidium, Adikia, Adynasia, Allognosis, Allopliosilpha, Alphacoleoptera, Anancaeon, Anapiptus, Ancylocheira liasina, Anepismus, Angelinella, Anhuistoma, Anhydrophilus, Anomerus, Anypostatus, Apheloodes, Apicasia, Apioderes, Apiopyrenides, Apistotes, Apsychus, Aptilotitus (syn. Aptilotus), Archosyne, Auchenophorites (syn. Auchenophorus), Babuskaya, Bareus, Bathygerus, Bellingera, Bellingeropsis, Biadelater, Blaptoides, Bothroptera, Brachylaimon, Brachytrachelites, Brevispinnatus, Brochocoleus indibili, Brodiola, Bucklandula, Buprestium, Campopsis, Carabicina, Carabiques, Carabocera, Cardioides, Cardiosyne, Cebrionates, Cerylonopsis, Chimaerocoleites, Chlaeniopsis, Chrysomelidium, Chrysomelinae, Chrysomelophana, Cicindeletae, Cistela, Cistelites, Clathropenna, Clerii, Clinomerus (syn. Catomerus), Coccinellophana, Coleopterites, Coleopteron, Coleopteron depressus, Coleopteron interpunctatus, Coleopteron laevigatus, Coleopteron scabriusculus, Colymbetopsis, Coprophagi, Cretaceites, Cretofalselaterium, Cretogermen, Cretohelophorus, Criocerina pristina, Critoderma (syn. Cycloderma), Cryptauchenia, Curculidium, Curculionites liasinus, Curculiopsidae, Curculium, Cydnopsis nigromembranacea, Cyphospheron, Darbiopelta, Diachoristes, Diaperialae, Diaperidium, Diatarastus, Diatrypamene, Dicyphelus (syn. Dicyphus), Diphymation, Diplocelides, Diplothece, Doggeria, Dysarestus, Ecthlimma, Elateridopsidae, Elateridopsis, Elaterina, Elytrulum, Enamma, Entomocantharus, Entomophagi, Eocoleopteron, Eodromeus viriosus, Episcepes, Epomenus, Erotilenae, Erotylites, Etheridgea, Euchroma arvernica, Eucoleoptera, Eurynotellus (syn. Eurynotus), Eurysphinctus, Eusarcantarus, Explansimarginis, Fengningia, Flichea, Fuscicupes, Gastrodelus, Gastroratus, Gemelina, Glaphoptera, Glaphyropterites, Glaphyropterula, Grahamelytron, Grasselites, Gurvanocoleus, Gyrinites, Gyrinulopsis, Halticophana, Harpalidium, Heeriaopsis (syn. Heeriella), Helophoropsis, Helophorus exilis, Helophorus magnus, Helopidium, Helopii, Helopium, Helops atticus, Hubeicoleus, Hydrobius godeti, Hydrobius obsoletus, Hydrocanthares, Hydroicetes, Hydrophilites, Hydrophilus stenopterus, Hydroporopsis, Hydrous ebeninus, Hylesinus lineatus, Hyperomima, Ipidae, Isomira aurora, Kamaroma, Katapiptus, Keleusticus, Kibdelia, Kryptophagus, Laimocenos, Lamiites, Lamiophanes, Laodiscis, Latridiites, Leioodes, Leptilostis (syn. Tetillopsis), Leptomites, Leptopodocoleidae, Leptosolenophorus, Longimaxilla, Loxocamarotus, Macrogastri, Macrotrachelites, Malacodermi, Malacodermidae, Masselytron, Megacentrus, Megachorites, Megopis lineolatus, Melanophilites, Melanophilopsis, Menephiloides, Mesocoleoptera, Mesoncus (syn. Loxoncus), Mesorhynchophora, Mesotricupes, Mesotylites, Metanastes, Metaxycoleoptera, Micrelaterium, Microcoleopteron, Microentomus, Mimelater, Nannoodes, Nebrioides, Necrophagi, Necticus minutus, Nitidulariae, Nubili, Omogongylus, Onkedodimus, Onthophagus luteus, Ooidellus (syn. Ooides), Ooperiglyptus, Ooperioristus, Oropseini, Oxycephalites, Oxytoroptera, Pachycoleon, Pachypleurites, Paklapwania, Palaeotrachys, Palaeotylinae, Pallax, Palpatores, Pantodapus, Parabuprestium, Paracurculium, Paragrilium, Paragyrinus, Parasilphites, Parnidium, Parnosoma, Pedinosoma, Peltocoleops, Periboloptera, Peridosoma (syn. Perosoma), Petalocupes, Petrorophus, Phaulogyrinus, Pholipheron, Phytoplesion, Pimeliodes, Plastobuprestites, Plastonebria, Platyprosopi, Pleuralocista, Polypamon, Praedodromeus, Prionophana, Prochrysomela, Proheuristes, Prophasis, Prosthecogmus, Prosthenostictus, Protocoleoptera, Protocuneus, Prototoma, Pseudobuprestites, Pseudocarabites, Pseudocurculionites, Pseudocymindis, Pseudocyphon, Pseudoelateropsis (syn. Elateropsis), Pseudohydrophilites, Pseudoprionites, Pseudotelephorus, Pseudotenebrio, Pseudus, Rhomaleus, Rhynchophori, Saperda valdensis, Saprophagi, Scalopoides, Semiglobus, Sideriosemion, Silphidium (syn. Silphites), Sitonites, Smodicoptera, Sojanocoleidae, Sphaericites, Staphyliniae, Stenelytron, Sternoxi, Stictulus, Strongylites, Strullendorfia, Tachypus, Telephorium, Tenebrionites, Tetragonides, Thurmannia, Toksunius, Tolype, Triadocoleopteron, Trialarva, Trichelepturgetes, Tridigitati, Tripsalis, Trixagites, Trochiscites, Trochmalus, Trogossitarii, Trypocoleus, Tshekardocoleidae (syn. Moravocoleidae, Uralocoleidae), Tychon, Ulomites, Velenovskya, Versicolores, Wollastonia, Xenogyrinus, Xinbinia, Xylophagi, Xylotupia, Yanqingia, Zacoleoptera, Zetemenos

Synonyms
Synonymy list
YearName and author
1758Coleoptera Linnaeus p. 341
1759Coleoptera Schreber p. 7
1761Coleoptera Linnaeus p. 133
1762Coleoptera Geoffroy
1762Cistela Geoffroy p. 115
1763Coleoptera Scopoli p. 1
1764Coleoptera Müller p. 1
1766Coleoptera Schaeffer
1767Coleoptera Linnaeus p. 537
1768Coleoptera Strøm p. 313
1770Coleoptera Drury
1771Coleoptera Forster p. 1
1771Cistela Forster p. 12
1775Eleuterata Fabricius p. 1
1776Coleoptera Müller p. 52
1777Coleoptera Goeze p. 1
1777Cistela Goeze p. 151
1796Coleoptera Latreille p. 1
1798Eleuterata Fabricius p. 1
1798Eleutherata Illiger p. 497
1813Criocerina pristina Germar p. 14
1852Curculionites liasinus Heer p. 15 fig. 39,40
1856Necticus minutus Aymard p. 234
1856Ancylocheira liasina Giebel p. 82
1856Hydrobius obsoletus Heer p. 18 figs. Pl 1, fig 19
1862Hydrophilus stenopterus Heer p. 65 figs. Pl 4, fig 13
1862Helophorus magnus Heer p. 70 figs. Pl 5, figs 22,22b
1862Hydrobius godeti Heer p. 70 figs. Pl 5, figs 21,21b
1862Helophorus exilis Heer p. 71 figs. Pl 5, fig 20
1864Hydrophilites interpunctatus Heer p. 64
1864Strongylites laevigatus Heer p. 89
1864Saperda valdensis Heer p. 376 fig. 254
1867Helops atticus Redtenbacher p. 33 figs. Pl XVI fig 18
1874Hydrobius obsoletus Oustalet p. 128 figs. Pl 6, fig 18
1874Onthophagus luteus Oustalet p. 194 figs. Pl 2, fig 17
1879Carabites depressus Heer p. 91
1879Buprestites scabriusculus Heer p. 92
1888Criocerina pristina von Schlechtendal p. 478
1891Hylesinus lineatus Förster p. 401 figs. Pl 12, fig 16
1907Crioceris pristina Handlirsch p. 792
1914Isomira aurora Wickham p. 268 figs. Pl 5, fig 6
1921Coleoptera Cockerell
1929Elateridopsidae Zalessky
1932Elateridopsidae Zalessky p. 201
1939Hydrous ebeninus Statz p. 64 figs. Pl. 18, fig. 81
1940Euchroma arvernica Piton pp. 181-182 figs. Pl XVII fig 7
1940Acmaeodera perreaui Piton pp. 193-194 figs. Pl XVII fig 5
1940Megopis lineolatus Piton p. 209 fig. 76
1953Coleoptera Bode
1967Cydnopsis nigromembranacea Jordan p. 86 figs. Pl 3, fig 13
1982Leptopodocoleidae Hong p. 118
1987Coleoptera Geyer and Kleber p. 342
1989Coleoptera Shoshani et al. p. 435
1989Coleoptera Zhang p. 420
1992Coleoptera Carpenter
1992Hylesinus lineatus Wood and Bright p. 81
1995Coleoptera Lawrence and Newton, Jr.
1997Eodromeus viriosus Zhang p. 88 figs. Pl 3, fig 3; text-fig 10
2004Coleoptera Kukalová-Peck and Lawrence
2006Brochocoleus indibili Soriano and Delclos
2008Helops atticus Kirejtshuk et al. p. 135
2011Coleoptera Beutel et al.
2014Coleoptera Misof et al.
2014Coleoptera Peters et al.
2018Isomira aurora Bousquet et al. p. 381
2022Eucoleoptera Cai et al.
2022Metaxycoleoptera Cai et al.
2022Zacoleoptera Cai et al.
2022Coleoptera Engel
2022Eucoleoptera Engel et al.
2022Metaxycoleoptera Engel et al.
2022Zacoleoptera Engel et al.

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—Ecdysozoa
—Panarthropoda
phylumArthropodaLatreille 1829
superclassHexapodaLatreille 1825
classInsecta
—Dicondylia
RankNameAuthor
—Paranotalia
subclassPterygota()
—NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
—Holometabola
—Aparaglossata
—Neuropteriformia
—Coleopterida
orderColeoptera
orderColeoptera

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Or. Coleoptera Linnaeus 1758 [beetle]
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G. †Adelidium Tillyard 1918
+
†Adelidium cordatum Tillyard 1918
G. †Adikia Handlirsch 1906
+
†Adikia punctulata Handlirsch 1906
G. †Adynasia Handlirsch 1906
+
†Adynasia lyelli Heer 1864
G. †Allognosis Handlirsch 1906
+
†Allognosis nitens Geinitz 1894
†Allognosis striatus Ponomarenko 1985
G. †Allopliosilpha Gersdorf 1971
+
†Allopliosilpha inclavata Gersdorf 1971
Subor. †Alphacoleoptera Engel et al. 2022
+
Superfm. †Coleopsoidea Kirejtshuk and Nel 2016
+
Fm. †Coleopsidae Kirejtshuk and Nel 2016
+
G. †Coleopsis Kirejtshuk et al. 2014
+
†Coleopsis archaica Kirejtshuk et al. 2014
Superfm. †Permocupedoidea Martynov 1933
+
Fm. †Labradorocoleidae Ponomarenko 1969
+
G. †Labradorocoleus Ponomarenko 1969
+
†Labradorocoleus carpenteri Ponomarenko 1969
Fm. †Oborocoleidae Kukalová 1968
+
G. †Liberocoleus Kukalová 1968
+
†Liberocoleus intactus Kukalová 1968
G. †Oborocoleus Kukalová 1968
+
†Oborocoleus rohdendorfi Kukalová 1968
Invalid names: Tshekardocoleoidea Rodendorf 1944 [synonym]
Superfm. †Permosynoidea Tillyard 1924
+
Fm. †Permosynidae Tillyard 1924
+
G. †Alveolacupes Lin 1983
+
†Alveolacupes primus Lin 1983
G. †Alveolicupes Lin 1983
G. †Artematopodites Ponomarenko 1990
+
+
Invalid names: Ovivagina immediata Zhang 1997 [synonym]
†Artematopodites latissimus Ponomarenko 2014
†Artematopodites latus Ponomarenko 1990
†Artematopodites leskoviensis Yan and Zhang 2010
+
Invalid names: Artematopodites longus Ponomarenko 1990 [replaced]
†Artematopodites lozowskii Ponomarenko 2015
†Artematopodites major Ponomarenko 1990
G. †Bistrisyne Lin 1986
+
†Bistrisyne tenua Lin 1986
G. †Delpuentesyne Martins-Neto and Gallego 2007
+
†Delpuentesyne menendezi Martins-Neto and Gallego 2007
G. †Diarcuipenna Lin 1986
+
†Diarcuipenna bennettitophila Ponomarenko 2011
G. †Dinoharpalus Handlirsch 1906
+
†Dinoharpalus coptoclavoides Ponomarenko 2015
†Dinoharpalus latus Ponomarenko 2015
†Dinoharpalus liasinus Giebel 1856
†Dinoharpalus martynovi Ponomarenko 2018
†Dinoharpalus rugosus Ponomarenko 1985
G. †Dzeregia Ponomarenko 1985
+
†Dzeregia ampla Ponomarenko 1990
†Dzeregia byrrhoides Ponomarenko 1990
†Dzeregia crassa Ponomarenko 1990
†Dzeregia juxta Lin 1986
†Dzeregia lata Ponomarenko 1985
†Dzeregia longa Ponomarenko 1985
†Dzeregia pilula Ponomarenko 1990
†Dzeregia platis Ponomarenko 2014
†Dzeregia striata Ponomarenko 1985
G. †Hydrobiites Heer 1864
+
†Hydrobiites anglicus Handlirsch 1906
†Hydrobiites bellus Geinitz 1894
†Hydrobiites convexus Ponomarenko 1985
†Hydrobiites crassus Ponomarenko 1985
†Hydrobiites dobbertinensis Handlirsch 1939
†Hydrobiites giebeli Handlirsch 1906
+
Invalid names: Berosus liasinus Giebel 1856 [replaced]
†Hydrobiites handlirschi Ponomarenko 2011
†Hydrobiites liasinus Giebel 1856
†Hydrobiites longus Ponomarenko 1990
†Hydrobiites minor Ponomarenko 2014
†Hydrobiites minutissimus Ponomarenko 1985
†Hydrobiites mongolicus Ponomarenko 2014
†Hydrobiites permianus Ponomarenko 2018
†Hydrobiites punctatostriatus Handlirsch 1939
†Hydrobiites purbeccensis Giebel 1856
†Hydrobiites sulcatus Ponomarenko 1986
†Hydrobiites tillyardi Ponomarenko 2011
†Hydrobiites veteranus Heer 1852
†Hydrobiites vladimiri Ponomarenko 2011
G. †Ischichucasyne Martins-Neto and Gallego 2005
+
†Ischichucasyne cladocosta Martins-Neto and Gallego 2009
†Ischichucasyne santajuanaensis Martins-Neto and Gallego 2005
G. †Palademosyne Rohdendorf 1961
+
†Palademosyne martynovae Rohdendorf 1961
†Palademosyne natalensis Ponomarenko 2005
G. †Permocrossos Martynov 1932
+
†Permocrossos elongatus Martynov 1932
G. †Permosyne Tillyard 1924
+
†Permosyne affinis Tillyard 1924
†Permosyne belmontensis Tillyard 1924
†Permosyne dentata Ponomarenko 2003
†Permosyne elongata Ponomarenko 2005
†Permosyne mitchelli Tillyard 1924
†Permosyne rasnitsyni Ponomarenko 2011
G. †Platycrossos Dunstan 1923
+
†Platycrossos caroli Ponomarenko 2011
†Platycrossos elongatus Ponomarenko 1985
†Platycrossos latus Ponomarenko 2014
†Platycrossos ligulatus Dunstan 1923
†Platycrossos longus Ponomarenko 2014
†Platycrossos loxonicus Ponomarenko 2014
†Platycrossos mongolicus Ponomarenko 2014
†Platycrossos ovum Ponomarenko 2014
†Platycrossos petalus Ponomarenko 1985
†Platycrossos punctatus Ponomarenko 1985
†Platycrossos subtumidus Dunstan 1923
†Platycrossos tumidus Tillyard 1916
G. †Pseudorhynchophora Handlirsch 1906
+
†Pseudorhynchophora convexa Ponomarenko 2008
G. †Sakmaracoleus Ponomarenko 2013
+
†Sakmaracoleus orenburgensis Ponomarenko 2013
G. †Tychtocoleus Rohdendorf 1961
+
†Tychtocoleus europeus Ponomarenko 2003
†Tychtocoleus neuburgae Rohdendorf 1961
†Tychtocoleus popovi Ponomarenko 2018
G. †Umkomaasia Zeuner 1960
+
†Umkomaasia depressa Zeuner 1960
G. †Anancaeon Handlirsch 1939
+
†Anancaeon microcephalum Handlirsch 1939
G. †Anapiptus Handlirsch 1906
+
†Anapiptus brodiei Handlirsch 1906
G. †Anepismus Handlirsch 1906
+
†Anepismus vanus Giebel 1856
G. †Angelinella Handlirsch 1906
+
†Angelinella angelini Heer 1878
G. †Anhuistoma Lin 1985
+
†Anhuistoma hyla Lin 1985
G. †Anhydrophilus Handlirsch 1906
+
†Anhydrophilus brodiei Handlirsch 1906
G. †Anomerus Handlirsch 1939
+
†Anomerus punctifer Handlirsch 1939
G. †Anypostatus Handlirsch 1939
+
†Anypostatus taurus Handlirsch 1939
G. †Apheloodes Dunstan 1923
+
†Apheloodes obliquum Dunstan 1923
†Apheloodes rugosum Dunstan 1923
G. †Apicasia Bode 1953
+
†Apicasia inolata Bode 1953
G. †Apioderes Handlirsch 1939
+
†Apioderes punctatus Handlirsch 1939
G. †Apiopyrenides Bode 1953
+
G. †Apistotes Handlirsch 1906
+
†Apistotes purbeccensis Giebel 1856
G. †Apsychus Handlirsch 1939
+
†Apsychus alutaceus Handlirsch 1939
G. †Aptilotitus Carpenter 1986
+
Invalid names: Aptilotus Bode 1953 [replaced]
G. †Archosyne Ponomarenko et al. 2014
+
†Archosyne permiana Ponomarenko et al. 2014
G. †Auchenophorites Carpenter 1986
+
Invalid names: Auchenophorus Bode 1953 [replaced]
G. †Babuskaya Martins-Neto and Gallego 2009
+
†Babuskaya elaterata Martins-Neto and Gallego 2009
G. †Bareus Handlirsch 1939
+
†Bareus strigipennis Handlirsch 1939
G. †Bathygerus Handlirsch 1906
+
†Bathygerus bellus Geinitz 1894
†Bathygerus divergens Geinitz 1894
G. †Bellingera Heer 1852
+
†Bellingera ovalis Heer 1852
G. †Bellingeropsis Handlirsch 1906
+
G. †Biadelater Handlirsch 1906
+
†Biadelater werneri Giebel 1856
G. †Blaptoides Frič 1901
+
†Blaptoides dubius Frič 1901
G. †Bothroptera Handlirsch 1906
+
†Bothroptera westwoodi Giebel 1856
G. †Brachylaimon Bode 1953
+
†Brachylaimon striatus Bode 1953
G. †Brachytrachelites Bode 1953
+
G. †Brevispinnatus Gomez Pallerola 1979
+
†Brevispinnatus paludis Gomez Pallerola 1979
G. †Brodiola Handlirsch 1906
+
†Brodiola nana Handlirsch 1906
G. †Bucklandula Handlirsch 1906
+
†Bucklandula striata Handlirsch 1906
G. †Buprestium Westwood 1854
+
†Buprestium gorgus Westwood 1854
G. †Campopsis Braun 1860
+
G. †Carabicina Germar 1842
+
†Carabicina decipiens Germar 1842
G. †Carabocera Brauer et al. 1889
+
†Carabocera prisca Brauer et al. 1889
G. †Cardioides Bode 1953
G. †Cardiosyne Martins-Neto and Gallego 2006
+
†Cardiosyne elegans Martins-Neto and Gallego 2006
†Cardiosyne obesa Martins-Neto and Gallego 2006
G. †Cerylonopsis Handlirsch 1906
+
†Cerylonopsis striata Brodie 1845
G. †Chimaerocoleites Ponomarenko 1985
+
†Chimaerocoleites gyrinoides Ponomarenko 1985
†Chimaerocoleites parvulus Ponomarenko 1985
†Chimaerocoleites punctatus Ponomarenko 1985
†Chimaerocoleites seriatus Ponomarenko 1985
G. †Chlaeniopsis Handlirsch 1906
+
†Chlaeniopsis solitarius Deichmüller 1886
G. †Chrysomelidium Handlirsch 1907
+
†Chrysomelidium simplex Frič 1901
G. †Chrysomelophana Handlirsch 1906
+
†Chrysomelophana rara Weyenbergh 1869
G. †Cistelites Heer 1864
+
†Cistelites insignis Heer 1864
†Cistelites longipes Hong 1985
†Cistelites minor Heer 1874
G. †Clathropenna Fujiyama 1973
+
†Clathropenna rugosa Fujiyama 1973
G. †Clinomerus Carpenter 1986
+
†Clinomerus laticollis Handlirsch 1939
Invalid names: Catomerus Handlirsch 1939 [replaced]
G. †Coccinellophana Handlirsch 1906
+
†Coccinellophana murchisoni Handlirsch 1906
G. †Coleopterites Braun 1860
+
G. †Coleopteron Handlirsch 1906
+
†Coleopteron antiquus Weyenbergh 1869
†Coleopteron aquaticus Brodie 1845
†Coleopteron beyrichi Giebel 1856
†Coleopteron bractoides Blake 1876
†Coleopteron lentissimus Weyenbergh 1874
†Coleopteron semicrematus Jarzembowski 2003
†Coleopteron sibiricus Heer 1876
†Coleopteron striatus Oppenheim 1888
†Coleopteron studeri Giebel 1856
†Coleopteron vetustus Giebel 1856
†Coleopteron wittsi Brodie 1845
†Coleopteron zekelii Giebel 1856
Invalid names: Coleopteron harpalinus Heer 1864 [nomen nudum], Coleopteron spectabilis Heer 1864 [nomen nudum], Coleopteron troglodytes Heer 1864 [nomen nudum]
G. †Colymbetopsis Handlirsch 1906
+
G. †Cretaceites Wang 1998
+
G. †Cretofalselaterium Alexeev 2008
+
G. †Cretogermen Alexeev 2008
+
†Cretogermen turonicum Alexeev 2008
G. †Cretohelophorus Ponomarenko 1987
+
†Cretohelophorus cupedoides Ponomarenko 1987
G. †Critoderma Carpenter 1986
+
†Critoderma deplanatum Heer 1864
Invalid names: Cycloderma Heer 1864 [replaced]
G. †Cryptauchenia Bode 1953
G. †Curculidium Handlirsch 1907
G. †Curculium Westwood 1854
+
†Curculium syrichthus Westwood 1854
G. †Cyphospheron Bode 1953
G. †Darbiopelta Ponomarenko 1986
+
†Darbiopelta crassa Ponomarenko 1986
†Darbiopelta dubia Ponomarenko 1986
G. †Diachoristes Bode 1953
+
†Diachoristes collinus Bode 1953
G. †Diaperidium Westwood 1854
+
†Diaperidium mithrax Westwood 1854
G. †Diatarastus Handlirsch 1906
+
†Diatarastus westwoodi Giebel 1856
G. †Diatrypamene Bode 1953
+
†Diatrypamene excavata Bode 1953
G. †Dicyphelus Carpenter 1986
+
Invalid names: Dicyphus Bode 1953 [replaced]
G. †Diphymation Bode 1953
+
†Diphymation corrosum Bode 1953
G. †Diplocelides Bode 1953
+
†Diplocelides minutus Bode 1953
G. †Diplothece Bode 1953
+
†Diplothece scissa Bode 1953
G. †Doggeria Handlirsch 1906
+
†Doggeria bucklandi Mantell 1844
†Doggeria murchisoni Handlirsch 1906
†Doggeria sibirica Handlirsch 1906
G. †Dysarestus Handlirsch 1906
+
†Dysarestus vetustus Handlirsch 1906
G. †Ecthlimma Handlirsch 1939
+
†Ecthlimma forficuloides Handlirsch 1939
G. †Elateridopsis Zalessky 1929
+
†Elateridopsis permiensis Zalessky 1929
G. †Elaterina Gardiner 1961
+
†Elaterina liassica Gardiner 1961
G. †Elytrulum Handlirsch 1907
+
G. †Enamma Handlirsch 1906
+
†Enamma striatum Handlirsch 1906
G. †Entomocantharus Bode 1953
+
G. †Eocoleopteron Handlirsch 1906
+
†Eocoleopteron roemeri Handlirsch 1906
G. †Episcepes Bode 1953
+
†Episcepes rotundatus Bode 1953
G. †Epomenus Handlirsch 1906
+
†Epomenus rugosus Handlirsch 1906
G. †Erotylites Cockerell 1920
+
†Erotylites wallacei Cockerell 1920
G. †Etheridgea Handlirsch 1906
+
†Etheridgea australis Handlirsch 1906
†Etheridgea petrica Tillyard 1916
G. †Eurynotellus Carpenter 1986
+
Invalid names: Eurynotus Bode 1953 [replaced]
G. †Eurysphinctus Bode 1953
+
G. †Eusarcantarus Bode 1953
+
G. †Explansimarginis Lin 1997
+
G. †Fengningia Hong 1984
+
†Fengningia punctata Hong 1984
G. †Flichea Handlirsch 1906
+
†Flichea elongata Ponomarenko 1985
†Flichea lotharingiaca Fliche 1901
†Flichea sibirica Ponomarenko 1985
G. †Fuscicupes Hong and Wang 1990
+
†Fuscicupes parvus Hong and Wang 1990
G. †Gastrodelus Bode 1953
+
G. †Gastroratus Bode 1953
+
G. †Gemelina Martins-Neto and Gallego 2006
+
†Gemelina triangularis Martins-Neto and Gallego 2006
G. †Glaphoptera Handlirsch 1906
+
†Glaphoptera anglica Handlirsch 1906
G. †Glaphyropterites Handlirsch 1906
+
G. †Glaphyropterula Handlirsch 1906
+
†Glaphyroptula anglica Cockerell 1915
†Glaphyroptula liasina Giebel 1856
G. †Grahamelytron Zeuner 1959
+
†Grahamelytron crofti Zeuner 1959
G. †Grasselites Bode 1953
+
†Grasselites pusillus Bode 1953
G. †Gurvanocoleus Ponomarenko 1986
+
†Gurvanocoleus hirsutus Ponomarenko 1986
G. †Gyrinites Heer 1852
+
†Gyrinites troglodytes Heer 1852
G. †Gyrinulopsis Handlirsch 1906
+
†Gyrinulopsis nanus Handlirsch 1906
G. †Halticophana Handlirsch 1906
+
†Halticophana westwoodi Handlirsch 1906
G. †Harpalidium Westwood 1854
+
†Harpalidium anactus Westwood 1854
G. †Heeriaopsis Carpenter 1986
+
Invalid names: Heeriella Handlirsch 1906 [replaced]
G. †Helophoropsis Handlirsch 1906
+
†Helophoropsis brodiei Giebel 1856
G. †Helopidium Westwood 1854
+
†Helopidium brodiei Giebel 1856
†Helopidium dubium Handlirsch 1906
†Helopidium dunkeri Giebel 1856
†Helopidium neoridas Westwood 1854
†Helopidium rugosum Handlirsch 1906
†Helopidium westwoodi Giebel 1856
G. †Helopium Westwood 1854
+
†Helopium agabus Westwood 1854
G. †Hubeicoleus Hong 1985
+
†Hubeicoleus tenuis Hong 1985
G. †Hydroicetes Bode 1953
+
†Hydroicetes affictus Bode 1953
G. †Hydrophilites Heer 1864
+
†Hydrophilites stygius Heer 1864
G. †Hydroporopsis Handlirsch 1906
+
†Hydroporopsis neptuni Giebel 1856
G. †Hyperomima Handlirsch 1906
+
†Hyperomima antiqua Giebel 1856
G. †Kamaroma Handlirsch 1906
+
†Kamaroma breve Handlirsch 1906
G. †Katapiptus Handlirsch 1906
+
†Katapiptus striolatus Handlirsch 1906
G. †Keleusticus Handlirsch 1906
+
†Keleusticus acutus Ponomarenko 1985
†Keleusticus duplicatus Ponomarenko 1985
†Keleusticus zirkeli Geinitz 1894
G. †Kibdelia Handlirsch 1906
+
†Kibdelia oolitica Brodie 1845
G. Kryptophagus Herbst 1792
G. †Laimocenos Bode 1953
+
G. †Lamiites Frič 1889
+
†Lamiites simillimus Frič 1889
G. †Lamiophanes Handlirsch 1906
+
†Lamiophanes schroeteri Giebel 1856
G. †Laodiscis Lin 1977
+
†Laodiscis melanis Lin 1977
G. †Latridiites Heer 1864
+
†Latridiites schaumi Heer 1864
G. †Leioodes Dunstan 1923
+
†Leioodes planum Dunstan 1923
†Leioodes pygmaeum Dunstan 1923
G. †Leptilostis Ponting 2024
+
†Leptilostis parvula Hong 1982
Invalid names: Tetillopsis Hong 1982 [replaced]
G. †Leptomites Bode 1953
+
†Leptomites procerus Bode 1953
G. †Leptosolenophorus Bode 1953
+
G. †Longimaxilla Hong 1982
+
†Longimaxilla sinica Hong 1982
G. †Loxocamarotus Bode 1953
+
G. †Macrotrachelites Bode 1953
+
G. †Masselytron Handlirsch 1939
+
†Masselytron quinquestriatum Handlirsch 1939
G. †Megacentrus Heer 1852
+
†Megacentrus tristis Heer 1852
G. †Megachorites Bode 1953
+
G. †Melanophilites Handlirsch 1906
+
G. †Melanophilopsis Handlirsch 1906
+
G. †Menephiloides Fujiyama 1973
+
†Menephiloides minensis Fujiyama 1973
Unr. Mesocoleoptera Boudinot et al. 2022
+
Unr. Metacoleoptera Boudinot et al. 2022
+
Fm. †Ademosynidae Ponomarenko 1968
+
G. †Ademosyne Handlirsch 1906
+
†Ademosyne adunca Dunstan 1923
†Ademosyne arcucciae Martins-Neto and Gallego 2006
†Ademosyne australiensis Tillyard 1916
†Ademosyne bacca Ponomarenko 1969
†Ademosyne brevis Dunstan 1923
†Ademosyne cameroni Tillyard 1916
†Ademosyne congener Tillyard 1916
†Ademosyne curvata Dunstan 1923
†Ademosyne elliptica Ponomarenko 1969
†Ademosyne elongatus Martins-Neto and Gallego 2006
†Ademosyne hexacostata Martins-Neto and Gallego 2006
†Ademosyne intermedia Dunstan 1923
†Ademosyne kirghizica Ponomarenko 1969
†Ademosyne lata Dunstan 1923
†Ademosyne llantenesensis Brauckmann et al. 2010
†Ademosyne major Handlirsch 1906
†Ademosyne olliffi Handlirsch 1906
†Ademosyne parva Dunstan 1923
†Ademosyne polyzetete Ponomarenko 2011
†Ademosyne prisca Riek 1974
†Ademosyne punctata Tillyard 1916
†Ademosyne punctuada Martins-Neto and Gallego 2006
†Ademosyne ramocostata Dunstan 1923
†Ademosyne reducta Riek 1976
†Ademosyne rosenfeldi Brauckmann et al. 2010
†Ademosyne rugulosa Dunstan 1923
†Ademosyne sibirica Rohdendorf 1961
†Ademosyne speciosa Riek 1974
†Ademosyne umutu Martins-Neto and Gallego 2009
†Ademosyne vittamargina Dunstan 1923
†Ademosyne wianamattensis Tillyard 1918
G. †Atalosyne Ren 1995
+
†Atalosyne sinuolata Ren 1995
G. †Cephalosyne Ponomarenko 1969
+
†Cephalosyne capitata Ponomarenko 1969
G. †Chaocoleus Ponomarenko et al. 2014
+
†Chaocoleus limnebius Ponomarenko et al. 2014
G. †Dolichosyne Ponomarenko 1969
+
†Dolichosyne confragosa Ponomarenko 1969
†Dolichosyne rostrata Ponomarenko 1969
†Dolichosyne sulcata Ponomarenko 1969
G. †Eremisyne Wang 1998
+
G. †Gnathosyne Ponomarenko 1969
+
†Gnathosyne akkolkensis Ponomarenko 1969
G. †Grammositum Dunstan 1923
+
†Grammositum bilineatus Dunstan 1923
G. †Petrosyne Ponomarenko 1969
+
†Petrosyne liassica Ponomarenko 1969
G. †Sphaerosyne Ponomarenko 1969
+
†Sphaerosyne globosa Ponomarenko 1969
Subor. Adephaga Clairville 1806
+
G. †Amarodes Handlirsch 1906
+
†Amarodes pseudozabrus Deichmüller 1886
G. †Amphoxyne Bode 1953
+
†Amphoxyne darbiensis Ponomarenko 1986
†Amphoxyne lineata Bode 1953
+
Invalid names: Cryptauchenia simplex Bode 1953 [synonym]
†Amphoxyne minuta Bode 1953
+
Invalid names: Coilotrachelus lineatus Bode 1953 [synonym], Cricotrachelites rotundatus Bode 1953 [synonym], Loxoncus procerus Bode 1953 [synonym]
Invalid names: Coilotrachelus Bode 1953 [synonym], Cricotrachelites Bode 1953 [synonym], Loxostelidotus Bode 1953 [synonym]
Fm. Carabidae Latreille 1802 [ground beetle]
+
G. †Antemnacrassa Gomez Pallerola 1979
G. Apotomopterus Hope 1838
G. †Ataktosites Haupt 1956
G. Bembidium Latreille 1802
Subfm. Brachininae Bonelli 1810
G. †Brachinites Fritsch 1882
Subfm. Broscinae Hope 1838
G. †Carabidium Westwood 1854
Subfm. Carabinae Latreille 1802
G. †Carabites Ponomarenko 1977
G. †Carabites Heer 1852
G. †Carabopteron Martynov 1926
G. †Cavicarabus Hong 1985
Unr. Celeripedes Latreille 1802
G. †Cicindelites Meunier 1898
Subfm. †Conjunctiinae Ponomarenko 1977
Subfm. Elaphrinae Latreille 1802
G. †Elaphrotites Haupt 1956
G. †Fangshania Hong 1981
G. †Feronites Frič 1884
G. †Fonsecacarabus Martins-Neto and Mendes 2002
Subfm. Gehringiinae Darlington 1933
G. †Glenopterus Heer 1847
G. †Hadrocephalus Handlirsch 1906
Subfm. Harpalinae Bonelli 1810
+
Invalid names: Licininae Bonelli 1810 [synonym]
G. †Harpalomimes Handlirsch 1906
G. †Hebeicarabus Hong 1984
G. †Lithocantharis Lin 1997
Subfm. Loricerinae Bonelli 1810
G. †Meileyingia Hong 1987
G. †Metagrilium Handlirsch 1906
Subfm. Migadopinae Chaudoir 1861
Subfm. Nebriinae Laporte 1834
G. †Nebrorabus Ponomarenko 1989
G. †Neothanes Scudder 1890
G. †Notokistus Handlirsch 1906
Subfm. Omophroninae Bonelli 1810
Subfm. Paussinae Latreille 1806 [ant nest beetle]
+
Invalid names: Ozaenidae Hope 1838 [synonym]
Subfm. Platyninae Bonelli 1810
G. †Procarabites Handlirsch 1906
Subfm. Protopaussinae Gestro 1892
Subfm. †Protorabinae Ponomarenko 1977
G. †Rhopalochlaenius Zhang et al. 1994
Subfm. Scaritinae Bonelli 1810
+
Invalid names: Promecognathinae Le Conte 1853 [synonym]
G. †Sinocarabus Hong 1982
G. Stereocerus Kirby 1837
G. †Sunocarabus Hong 1987
Subfm. Trechinae Bonelli 1810
+
Invalid names: Patrobinae Kirby 1837 [synonym]
G. †Trechoides Motschulsky 1856
G. †Tremembecarabus Martins-Neto 1998
G. †Wongyekokia Lin 1997
G. †Wuchangicarabus Hong 1985
G. †Yunnanocarabus Lin 1977
Invalid names: Cratacanthus florissantensis Wickham 1917 [nomen dubium], Fossores Latreille 1802 [empty], Loricera exita Scudder 1900 [nomen dubium], Rhysodinae Laporte 1840 [empty], Siagoninae Bonelli 1813 [empty], Trechus fractus Wickham 1912 [nomen dubium]
Infraor. Carabiformia Lameere 1900
+
Superfm. Caraboidea Latreille 1802
G. †Chimaerocoleus Ponomarenko 1985
+
†Chimaerocoleus aenigma Ponomarenko 1985
Fm. Cicindelidae Latreille 1802 [tiger beetle]
+
Tr. Cicindelini Latreille 1802
G. †Cretotetracha Zhao et al. 2019
G. †Goriresina Matalin et al. 2021
G. †Hujia Song et al. 2022
Tr. Manticorini Laporte 1834
+
Invalid names: Amblycheilini Csiki 1903 [synonym]
Tr. Megacephalini Laporte 1834
G. †Opsis Handlirsch 1906
G. †Oxycheilopsis Cassola and Werner 2004
G. †Palaeopronyssiformia Wiesner et al. 2017
Invalid names: Oxycheilini Thomson 1857 [empty]
Fm. †Colymbotethidae Ponomarenko 1993
+
G. †Colymbotethis Ponomarenko 1993
Fm. †Coptoclavidae Ponomarenko 1961
+
G. †Agrascapha Lin 1992
G. †Amblycephalonius Bode 1953
+
Invalid names: Barocephalus Carpenter 1986 [synonym], Barycephalus Bode 1953 [synonym], Tricyrtus Bode 1953 [synonym]
Subfm. †Charonoscaphinae Ponomarenko 1977
G. †Coptoclavia Ping 1928
Subfm. †Coptoclavinae Ponomarenko 1961
Subfm. †Coptoclaviscinae Soriano et al. 2007
G. †Euroscapha Lin 1992
Subfm. †Hispanoclavinae Soriano et al. 2007
Subfm. †Timarchopsinae Wang et al. 2010
+
Invalid names: Necronectinae Ponomarenko 1977 [replaced]
G. †Curculionites Oppenheim 1888
+
†Curculionites striatus Oppenheim 1888
Fm. Dytiscidae Leach 1815 [predaceous diving beetle]
+
Subfm. Agabinae Thomson 1867
G. †Angaragabus Ponomarenko 1963
Subfm. Colymbetinae Erichson 1837
Subfm. Copelatinae Van den Branden 1885
Subfm. Coptotominae Van den Branden 1885
G. †Cretodytes Ponomarenko 1977
Subfm. Cybisterinae Sharp 1880
Subfm. Dytiscinae Leach 1815
Subfm. Hydroporinae Aubé 1836
Subfm. Laccophilinae Gistel 1848
+
Invalid names: Palaeogyrininae von Schlechtendal 1894 [synonym]
Subfm. †Liadytiscinae Prokin and Ren 2010
G. †Palaeodytes Ponomarenko 1987
G. †Sinoporus Prokin and Ren 2010
Invalid names: Aubehydrinae Guignot 1942 [empty], Necticus palustris Aymard 1856 [nomen nudum]
Fm. Gyrinidae Latreille 1810 [whirligig beetle]
+
G. †Anagyrinus Handlirsch 1906
G. †Coptogyrinus Handlirsch 1906
Subfm. Gyrininae Latreille 1810
G. †Gyrinopsis Handlirsch 1906
G. †Protogyrininus Hatch 1927
G. †Tunguskagyrus Yan et al. 2018
Invalid names: Spanglerogyrinae Folkerts 1979 [empty]
Subfm. Haliplidae Aubé 1836 [crawling water beetle]
+
G. †Cretihaliplus Ren et al. 1995
G. Haliplus Latreille 1802
G. Peltodytes Regimbart 1878
Fm. Hygrobiidae Regimbart 1878
+
G. Hygrobia Latreille 1804
+
Invalid names: Paelobius Schoenherr 1808 [synonym]
Invalid names: Paelobiidae Erichson 1837 [synonym]
Fm. †Liadytidae Ponomarenko 1977
+
G. †Liadytes Ponomarenko 1963
G. †Ovidytes Ren et al. 1995
G. †Longurius Hong 1982
+
†Longurius xigeduensis Hong 1982
G. †Memptus Handlirsch 1906
+
†Memptus ganglbaueri Ponomarenko 1985
†Memptus guttula Ponomarenko 1985
†Memptus handlirschi Ponomarenko 1985
†Memptus redtenbacheri Handlirsch 1906
+
Invalid names: Memptus braueri Handlirsch 1906 [synonym]
Fm. Noteridae Thomson 1860 [burrowing water beetle]
+
Subfm. Noterinae Thomson 1860
Subfm. Phreatodytinae Ueno 1957
G. †Oryctites Oppenheim 1888
+
†Oryctites fossilis Oppenheim 1888
Fm. †Parahygrobiidae Ponomarenko 1977
+
G. †Parahygrobia Ponomarenko 1977
Fm. Trachypachidae Thomson 1857 [false ground beetle]
+
G. †Apermunda Kirejtshuk and Ansorge 2022
G. †Denudirabus Ren 1995
G. †Dundorabus Ponomarenko 1989
Subfm. †Eodromeinae Ponomarenko 1977
G. †Eodromeites Ponomarenko 1985
G. †Necronectulus Ponomarenko 1977
Subfm. Trachypachinae Thomson 1857
Invalid names: Karadromeus xiangfanggouensis Hong 1995 [nomen dubium]
G. †Triadogyrus Ponomarenko 1977
+
†Triadogyrus sternalis Ponomarenko 1977
Fm. †Triaplidae Ponomarenko 1977
+
G. †Avocatinus Ponomarenko 1969
G. †Catinius Ponomarenko 1968
G. †Catinoides Ponomarenko 1969
G. †Clypeus Hong 1983
G. †Macrocatinius Ponomarenko 1969
G. †Tomiaplus Ponomarenko 2021
G. †Triaplus Ponomarenko 1977
G. †Triassocatinius Ponomarenko 1969
Invalid names: Catiniidae Ponomarenko 1968 [replaced]
Invalid names: Amphizoidae Leconte 1853 [empty], Dytiscoidea Leach 1815 [invalid subgroup], Pseudomorphinae Hope 1838 [empty], Tritarsusidae Hong 2002 [invalid subgroup]
Subor. Archostemata Kolbe 1908
+
G. †Anthocoleus Hong 1983
+
G. †Argentinocupes Martins-Neto and Gallego 2006
+
†Argentinocupes abdalai Martins-Neto and Gallego 2006
†Argentinocupes pulcher Martins-Neto and Gallego 2006
†Argentinocupes sara Martins-Neto and Gallego 2009
Superfm. †Asiocoleoidea Rohdendorf 1961
+
Fm. †Tricoleidae Ponomarenko 1969
G. †Bothynophora Heer 1864
+
†Bothynophora elegans Heer 1864
G. †Brachilatus Strelnikova and Yan 2023
+
†Brachilatus caducus Ponomarenko 1969
†Brachilatus dundulaensis Ponomarenko 1994
†Brachilatus foersteri Ponomarenko 1971
+
Invalid names: Notocupes reticulatus Ponomarenko 1968 [replaced]
†Brachilatus longicoxa Soriano and Delclos 2006
†Brachilatus nigrimonticola Ponomarenko 1968
†Brachilatus oculatus Soriano and Delclos 2006
†Brachilatus premeris Lee et al. 2022
†Brachilatus spinosus Li and Cai 2023
†Brachilatus viridis Soriano and Delclos 2006
G. †Celocoleus Hong 1983
+
†Celocoleus densus Hong 1983
G. †Clathrocupes Hong 1980
+
G. †Conexicoxa Lin 1986
+
†Conexicoxa brachycephala Ponomarenko 1994
†Conexicoxa crassa Ponomarenko 1969
†Conexicoxa denticollis Jiang et al. 2020
†Conexicoxa epicharis Tan et al. 2005
†Conexicoxa eumeura Tan et al. 2006
†Conexicoxa exigua Ponomarenko 1994
†Conexicoxa homora Lin 1986
†Conexicoxa jurassica Hong 1983
†Conexicoxa khetanensis Ponomarenko 1993
†Conexicoxa kirghizica Ponomarenko 1969
†Conexicoxa longicollis Ponomarenko 1994
†Conexicoxa martinclosas Soriano and Delclos 2006
†Conexicoxa porrecta Tan et al. 2006
†Conexicoxa psilata Tan et al. 2005
†Conexicoxa siniestri Soriano and Delclos 2006
†Conexicoxa stabilis Tan et al. 2005
†Conexicoxa tripartita Oppenheim 1888
G. †Cupedites Ponomarenko 1985
+
†Cupedites minutissimus Ponomarenko 1985
Superfm. Cupedoidea Laporte 1836
+
Fm. Cupedidae Laporte 1836 [reticulated beetle]
G. †Magnocoleus Hong 1998
Fm. Micromalthidae Barber 1913 [telephone pole beetle]
Fm. Ommatidae Sharp and Muir 1912
Invalid names: Crowsoniellidae Iablokoff-Khnzorian 1983 [empty], Magnocoleidae Hong 1998 [invalid subgroup]
G. †Dictyocoleus Hong 1982
+
G. †Doggeriopsis Handlirsch 1906
+
†Doggeriopsis stonesfieldiana Handlirsch 1906
G. †Eumecoleus Haupt 1950
+
†Eumecoleus tenuis Haupt 1950
G. †Euteticoleus Hong 1983
+
†Euteticoleus radiatus Hong 1983
G. †Gansucupes Hong 1982
+
†Gansucupes attenuatus Hong 1982
G. †Hebeicoleus Hong 1992
+
G. †Hebeicupes Zhang 1986
+
G. †Helopides Roemer 1876
+
†Helopides hildesiensis Roemer 1876
G. †Ironicus Handlirsch 1906
+
†Ironicus nothrus Westwood 1854
G. †Kargalarva Prokin et al. 2019
+
†Kargalarva permosialis Prokin et al. 2019
G. †Katapontisus Handlirsch 1906
+
†Katapontisus brodiei Giebel 1856
G. †Kelidus Handlirsch 1906
+
†Kelidus bolbus Westwood 1854
G. †Lecanopteryx Haupt 1950
+
†Lecanopteryx miranda Haupt 1950
G. †Longaevicupes Ren 1995
+
G. †Metacupes Gardiner 1961
+
†Metacupes harrisi Gardiner 1961
G. †Nannocurculionites Handlirsch 1906
+
G. †Notocupes Ponomarenko 1964
+
†Notocupes alienus Tan and Ren 2006
†Notocupes caudatus Ponomarenko 1966
†Notocupes cyclodontus Tan et al. 2006
†Notocupes daohugouensis Li and Cai 2023
†Notocupes diazromerali Soriano and Delclos 2006
†Notocupes elegans Ponomarenko 1994
†Notocupes excellens Ponomarenko 1966
†Notocupes kezuoensis Hong 1987
†Notocupes khasurtyiensis Strelnikova 2019
†Notocupes laetus Lin 1976
†Notocupes laiyangensis Hong and Wang 1990
†Notocupes lapidarius Ponomarenko 1968
†Notocupes latus Ponomarenko 1969
†Notocupes lini Ponomarenko et al. 2012
†Notocupes ludongensis Wang and Liu 1996
†Notocupes mongolicus Ponomarenko 1994
†Notocupes neli Tihelka et al. 2019
†Notocupes ohmkuhnlei Jarzembowski et al. 2019
†Notocupes picturatus Ponomarenko 1964
†Notocupes pingi Ponomarenko and Ren 2010
†Notocupes pulcher Ponomarenko 1968
†Notocupes reticulatus Oppenheim 1888
†Notocupes robustus Li and Cai 2023
†Notocupes rudis Tan et al. 2005
†Notocupes shiluoensis Hong 1984
†Notocupes tuanwangensis Hong and Wang 1990
†Notocupes validus Lin 1976
Invalid names: Amblomma Tan et al. 2005 [synonym], Chengdecupes Hong 1983 [synonym], Forticupes Hong and Wang 1990 [synonym], Lupicupes Ren 1995 [synonym], Ovatocupes Tan and Ren 2006 [synonym], Picticupes Hong and Wang 1990 [synonym], Sinocupes Lin 1976 [synonym]
G. †Parabuprestites Handlirsch 1906
+
G. †Paracurculionites Handlirsch 1906
+
G. †Paradoggeria Handlirsch 1906
+
†Paradoggeria acuminata Handlirsch 1906
G. †Pseudosilphites Zeuner 1930
+
G. †Rhabdocupes Ponomarenko 1966
+
†Rhabdocupes baculatus Ponomarenko 1969
†Rhabdocupes cellulosus Ponomarenko 1969
†Rhabdocupes dischides Zhang 1986
†Rhabdocupes issykkulensis Ponomarenko 1969
†Rhabdocupes laticella Ponomarenko 1969
†Rhabdocupes lentus Ren 1995
†Rhabdocupes longus Ponomarenko 1966
†Rhabdocupes minisculus Tan et al. 2006
†Rhabdocupes minor Ponomarenko 1966
†Rhabdocupes oxypygus Ponomarenko 1969
†Rhabdocupes protensus Tan et al. 2006
†Rhabdocupes rostratus Ponomarenko 1969
†Rhabdocupes sogutensis Ponomarenko 1969
†Rhabdocupes tenuis Ponomarenko 1969
†Rhabdocupes vitimensis Ponomarenko 1966
Superfm. †Rhombocoleoidea Rohdendorf 1961
+
Fm. †Rhombocoleidae Rohdendorf 1961
Superfm. †Schizocoleoidea Rohdendorf 1961
+
Fm. †Phoroschizidae Bouchard and Bousquet 2020
+
Invalid names: Schizophoridae Ponomarenko 1968 [replaced]
Fm. †Schizocoleidae Rohdendorf 1961
Invalid names: Schizophoroidea Ponomarenko 1968 [synonym]
G. †Stigmenamma Handlirsch 1906
+
†Stigmenamma heeri Giebel 1856
G. †Synodus Hong 1982
+
†Synodus changmaensis Hong 1982
G. †Tentyridium Westwood 1854
+
†Tentyridium dilatatum Handlirsch 1906
†Tentyridium peleus Westwood 1854
G. †Tetrocupes Hong 1983
+
†Tetrocupes cavernasus Hong 1983
G. †Tillyardiopsis Dunstan 1923
+
†Tillyardiopsis granulata Dunstan 1923
G. †Zygadenia Handlirsch 1906
+
†Zygadenia alexrasnitsyni Strelnikova and Yan 2021
†Zygadenia angliae Giebel 1856
†Zygadenia cornigera Lee et al. 2023
†Zygadenia floodpagei Jarzembowski et al. 2015
†Zygadenia giebeli Ponomarenko 2014
†Zygadenia handlirschi Ponomarenko 2014
†Zygadenia liui Jarzembowski et al. 2015
†Zygadenia martinae Jarzembowski et al. 2015
†Zygadenia semen Ponomarenko 2000
†Zygadenia sibirica Ponomarenko 2000
†Zygadenia simpsoni Jarzembowski et al. 2015
†Zygadenia sinitzae Ponomarenko 2000
†Zygadenia tuberculata Giebel 1856
Invalid names: Camtodontus Dejean 1826 [synonym], Kakoselia Handlirsch 1906 [synonym]
Invalid names: Cupediformia [invalid subgroup], Moravocoleus perditus Kukalová 1968 [nomen dubium], Notocupedini Ponomarenko 1966 [invalid subgroup], Priacma corrupta Ponomarenko 1986 [nomen dubium], Priacma longicapitis Ponomarenko 1997 [nomen dubium], Priacma oculata Ponomarenko 1997 [nomen dubium], Priacma striata Ponomarenko 2000 [nomen dubium]
Fm. †Asiocoleidae Rohdendorf 1961
+
G. †Asiocoleopsis Ponomarenko et al. 2014
+
†Asiocoleopsis hongi Ponomarenko et al. 2014
G. †Asiocoleus Rohdendorf 1961
+
†Asiocoleus novojilovi Rohdendorf 1961
G. †Bicoleus Ponomarenko 2013
+
†Bicoleus kuplensis Ponomarenko 2013
†Bicoleus laticella Ponomarenko 2013
G. †Gondvanocoleus Ponomarenko 2020
+
†Gondvanocoleus chikatunovi Ponomarenko et al. 2020
G. †Loculitricoleus Tan and Ren 2009
+
†Loculitricoleus flatus Tan and Ren 2009
†Loculitricoleus mongolicus Ponomarenko 2014
†Loculitricoleus tenuatus Tan and Ren 2009
G. †Schizotaldycupes Rohdendorf 1961
+
†Schizotaldycupes ananjevi Rohdendorf 1961
†Schizotaldycupes pubescens Ponomarenko 2013
Invalid names: Carinicupes Rohdendorf 1961 [synonym]
G. †Tetracoleus Ponomarenko 2008
+
†Tetracoleus aristovi Ponomarenko 2018
†Tetracoleus golubevi Ponomarenko 2015
†Tetracoleus meyeni Ponomarenko 2013
†Tetracoleus minimus Ponomarenko 2008
†Tetracoleus orenburgensis Ponomarenko 2013
†Tetracoleus sennikovi Ponomarenko 2011
†Tetracoleus tricoleoides Ponomarenko 2013
†Tetracoleus tshalyshevi Ponomarenko 2008
†Tetracoleus vasilenkoi Ponomarenko 2018
Subor. Myxophaga Crowson 1955
+
Superfm. Lepiceroidea Hinton 1936
+
Fm. Lepiceridae Hinton 1936
+
Invalid names: Cyathoceridae Scharp 1882 [synonym], Haplochelidae Kirejtshuk and Poinar 2006 [synonym]
+
Fm. Hydroscaphidae LeConte 1874 [skiff beetle]
Fm. Sphaeriusidae Erichson 1845
+
Invalid names: Microsporidae Crotch 1873 [synonym]
Invalid names: Sphaeriidae Erichson 1845 [empty], Torridincolidae Steffan 1964 [empty]
Invalid names: Sphaeriiformia [empty]
Fm. †Peltosynidae Yan et al. 2017
+
G. †Gnathopeltos Yan et al. 2017
+
†Gnathopeltos dixis Yan et al. 2017
G. †Ofthalmopeltos Yan et al. 2017
+
†Ofthalmopeltos synkritos Yan et al. 2017
G. †Peltosyne Ponomarenko 1977
+
†Peltosyne triassica Ponomarenko 1977
†Peltosyne varyvrosa Yan et al. 2017
Subor. Polyphaga Emery 1886
+
G. †Angarolarva Ponomarenko 1985
+
†Angarolarva aquatica Ponomarenko 1985
†Angarolarva incerta Ponomarenko 1990
Infraor. Bostrichiformia Forbes 1926
+
Superfm. Bostrichoidea Latreille 1802
Infraor. Bostrychiformia Forbes 1926
+
Invalid names: Dermestoidea Latreille 1804 [empty]
Unr. Clambiformia Cai and Tihelka 2022
+
Superfm. Clamboidea Fischer von Waldheim 1821
+
Invalid names: Derodontoidea LeConte 1861 [synonym], Eucinetoidea Lacordaire 1857 [synonym]
G. †Cryptophagites Ponomarenko 1990
+
†Cryptophagites capitatus Ponomarenko 1990
†Cryptophagites elongatus Ponomarenko 1990
Infraor. Cucujiformia Lameere 1938
+
Superfm. Cleroidea Latreille 1802
Superfm. Coccinelloidea Latreille 1807
Superfm. Cucujoidea Latreille 1802
Superfm. Erotyloidea Latreille 1802
Superfm. Lymexyloidea Fleming 1821
Superfm. Nitiduloidea Latreille 1802
Fm. †Parandrexidae Kirejtshuk 1994
G. †Petratypus Kaddumi 2005
Superfm. Tenebrionoidea Latreille 1802
Fm. †Wabbelidae Alekseev 2017
Infraor. Elateriformia Crowson 1960
+
Superfm. Buprestoidea Leach 1815
Superfm. Byrrhoidea Latreille 1804
Superfm. Cantharoidea
Superfm. Dascilloidea Gue'rin-Me'neville 1843
Superfm. Dryopoidea
Superfm. Elateroidea Leach 1815
+
Invalid names: Artematopoidea Lacordaire 1857 [synonym]
Invalid names: Mesosperchus angulatus Ponomarenko 1985 [nomen dubium], Podabrocephalidae Pic 1930 [empty]
G. †Elmadulescens Peris et al. 2015
+
†Elmadulescens rugosus Peris et al. 2015
G. †Jingxidiscus Hong 1990
+
Invalid names: Discus Hong 1981 [replaced]
Fm. Jurodidae Ponomarenko 1985
+
G. †Jurodes Ponomarenko 1985
Invalid names: Sikhotealiniidae Lafer 1996 [synonym]
G. †Larvula Ponomarenko 1985
+
†Larvula cassa Ponomarenko 1985
†Larvula triassica Prokin and Ponomarenko 2013
Fm. †Lasiosynidae Kirejtshuk et al. 2010
+
G. †Anacapitis Yan 2009
+
Invalid names: Brachysyne Tan and Ren 2009 [synonym]
G. †Bupredactyla Kirejtshuk et al. 2010
G. †Crassisyne Yan et al. 2014
G. †Glaphyropterodes Handlirsch 1906
G. †Lasiosyne Tan et al. 2007
+
Invalid names: Pappisyne Tan and Ren 2009 [synonym]
G. †Microsyne Yan et al. 2014
Invalid names: Lasiosyne laxa Yan et al. 2014 [nomen dubium]
G. †Melanocantharis Bode 1953
+
G. †Mormolucoides Hitchcock 1858
+
†Mormolucoides articulatus Hitchcock 1858
Unr. Nosodendriformia Cai and Tihelka 2022
+
Superfm. Nosodendroidea Erichson 1846
G. †Plastelater Handlirsch 1906
+
†Plastelater glabratus Ponomarenko 1985
†Plastelater neptuni Giebel 1856
G. †Polysitum Dunstan 1923
+
†Polysitum elongatum Ponomarenko 1985
†Polysitum hosbayari Ponomarenko 1986
†Polysitum kuznetskiense Rohdendorf 1961
†Polysitum latum Ponomarenko 1985
†Polysitum majus Ponomarenko 1990
†Polysitum minutus Dunstan 1923
†Polysitum ovale Ponomarenko 1990
†Polysitum punctatus Dunstan 1923
†Polysitum rugosum Ponomarenko 1986
†Polysitum sharategense Ponomarenko 2014
G. †Potamophilites Haupt 1956
+
G. †Ranis Ponomarenko 1968
+
†Ranis collevus Ponomarenko 1969
†Ranis ovalis Ponomarenko 1968
Unr. Rhinorhipiformia Cai et al. 2022
+
Superfm. Rhinorhipoidea Lawrence 1988
G. †Rhysopsalis Bode 1953
+
†Rhysopsalis distorta Bode 1953
†Rhysopsalis granulum Ponomarenko 1986
Unr. Scirtiformia Lawrence et al. 2011
+
Superfm. Scirtoidea Fleming 1821
G. †Sphaerocantharis Bode 1953
+
†Sphaerocantharis sibirica Ponomarenko 1990
+
Invalid names: Mikrocarpides lineatus Bode 1953 [synonym]
Invalid names: Mikrocarpides Bode 1953 [synonym], Theornithion Bode 1953 [synonym]
Infraor. Staphyliniformia Lameere 1900
+
Superfm. Histeroidea Gyllenhal 1808
Superfm. Hydrophiloidea Latreille 1802 [water scavenger beetle]
Superfm. Scarabaeoidea Latreille 1802 [scarab beetle]
Superfm. Staphylinoidea Latreille 1802
Invalid names: Scarabaeiformia Crowson 1960 [synonym]
G. †Syntomopterus Bode 1953
+
†Syntomopterus latus Bode 1953
Invalid names: Agyrtes primoticus Scudder 1900 [nomen dubium], Derodontiformia Le Conte 1861 [invalid subgroup], Escheria crassipunctata Förster 1891 [nomen dubium], Escheria dimidiata Förster 1891 [nomen dubium], Escheria punctulata Förster 1891 [nomen dubium], Eucinetiformia [empty], Limnius perrini Theobald 1937 [nomen dubium], Malachius vertummni Heer 1847 [nomen dubium], Mesosperchus notatus Ponomarenko 1977 [nomen dubium], Tomocoleites Hong 2002 [nomen nudum], Troglops punctatissima Théobald 1935 [nomen dubium]
Fm. †Ponomarenkiumidae Lawrence and Bouchard 2022
+
G. †Ponomarenkium Yan et al. 2018
+
†Ponomarenkium belmonthense Yan et al. 2017
Invalid names: Ponomarenkia Yan et al. 2017 [replaced]
Invalid names: Ponomarenkiidae Yan et al. 2017 [invalid subgroup]
Fm. †Taldycupedidae Rohdendorf 1961
+
G. †Afrotaldycupes Kirejtshuk 2020
+
†Afrotaldycupes africanus Ponomarenko 2005
†Afrotaldycupes lidgettoniensis Ponomarenko 2005
G. †Mesothoris Tillyard 1916
+
†Mesothoris clathrata Tillyard 1916
†Mesothoris grandis Dunstan 1923
†Mesothoris punctomarginum Dunstan 1923
G. †Proterocupes Ponomarenko 2015
+
†Proterocupes major Ponomarenko 2015
†Proterocupes nedubrovensis Ponomarenko 2015
†Proterocupes permiana Ponomarenko 2013
Invalid names: Lobanovia Kirejtshuk 2020 [objective synonym]
G. †Simmondsia Dunstan 1923
+
†Simmondsia cylindrica Dunstan 1923
†Simmondsia subpiriformis Dunstan 1923
G. †Taldycupes Rohdendorf 1961
+
†Taldycupes bergi Rohdendorf 1961
+
Invalid names: Cryptocupes rjabinini Rohdendorf 1961 [synonym]
†Taldycupes cellulosus Ponomarenko 2013
†Taldycupes khalfini Rohdendorf 1961
†Taldycupes moltshanovi Rohdendorf 1961
†Taldycupes pingi Ponomarenko et al. 2014
†Taldycupes reticulatus Ponomarenko 1969
†Taldycupes rosanovi Ponomarenko 2006
†Taldycupes vasjuchitshevi Rohdendorf 1961
Invalid names: Cryptocupes Rohdendorf 1961 [synonym], Taldycupidium Rohdendorf 1961 [synonym]
G. †Tecticupes Rohdendorf 1961
+
†Tecticupes heckeri Rohdendorf 1961
+
Invalid names: Stegocupes efremovi Rohdendorf 1961 [synonym], Stegocupes fedotovi Rohdendorf 1961 [synonym]
†Tecticupes indistinctus Rohdendorf 1961
+
Invalid names: Tychticupes tchihatcheffi Rohdendorf 1961 [synonym]
†Tecticupes martynovi Ponomarenko 2013
Invalid names: Stegocupes Rohdendorf 1961 [synonym]
G. †Tychticupes Rohdendorf 1961
+
†Tychticupes beljanini Rohdendorf 1961
†Tychticupes radtshenkoi Rohdendorf 1961
†Tychticupes ragozini Rohdendorf 1961
G. †Tychticupoides Rohdendorf 1961
+
†Tychticupoides grjazevi Rohdendorf 1961
G. †Wuchangia Hong 1985
+
†Wuchangia latilimbata Hong 1985
G. †Yiyangicupes Hong 1988
+
G. †Yuxianocoleus Hong 1985
+
Invalid names: Tychticupes ussovi Rohdendorf 1961 [nomen dubium], Uskaticupes Rohdendorf 1961 [nomen dubium], Uskaticupes javorskyi Rohdendorf 1961 [nomen dubium]
Fm. †Triadocupedidae Ponomarenko 1966
+
G. †Cupesia Ponomarenko 1966
+
†Cupesia monilicornia Ponomarenko 1966
†Cupesia sepulta Ponomarenko 1969
†Cupesia serricornia Ponomarenko 1969
G. †Moltenocupes Zeuner 1960
+
†Moltenocupes townrowi Zeuner 1960
G. †Platycupes Ponomarenko 1966
+
†Platycupes dolichocerus Ponomarenko 1966
†Platycupes major Ponomarenko 1966
†Platycupes pusillus Ponomarenko 1966
†Platycupes reticulatus Ponomarenko 1966
G. †Procupes Ponomarenko 1966
+
†Procupes mandibularis Ponomarenko 1966
G. †Triadocupes Ponomarenko 1966
+
†Triadocupes ellipticus Ponomarenko 1966
†Triadocupes ferghanensis Ponomarenko 1966
†Triadocupes latus Ponomarenko 1966
Fm. †Permocupedidae Martynov 1932
+
Subfm. †Permocupedinae Martynov 1932
+
G. †Afrocupes Geertsema and van den Heever 1996
+
†Afrocupes firmae Geertsema and van den Heever 1996
G. †Archicupes Rohdendorf 1961
+
†Archicupes jacobsoni Rohdendorf 1961
+
Invalid names: Palaeocupes kaltanicus Rohdendorf 1961 [synonym]
Invalid names: Palaeocupes Rohdendorf 1961 [synonym]
G. †Cytocupes Rohdendorf 1961
+
†Cytocupes angustus Rohdendorf 1961
G. †Cytocupoides Ponomarenko 1969
+
†Cytocupoides elongatus Ponomarenko 1969
G. †Eocupes Rohdendorf 1961
+
†Eocupes cellulosus Ponomarenko 1969
†Eocupes lukjanovitshi Rohdendorf 1961
G. †Frankencupes Ponomarenko and Bashkuev 2018
+
†Frankencupes ultimus Ponomarenko and Bashkuev 2018
G. †Ichthyocupes Rohdendorf 1961
+
†Ichthyocupes kuznetskiensis Ponomarenko 1969
†Ichthyocupes skoki Rohdendorf 1961
†Ichthyocupes tyzhnovi Rohdendorf 1961
G. †Kaltanicupes Rohdendorf 1961
+
†Kaltanicupes acutus Ponomarenko 1963
†Kaltanicupes kargalensis Ponomarenko 1963
†Kaltanicupes kitjakensis Ponomarenko 1963
†Kaltanicupes richteri Rohdendorf 1961
G. †Kaltanocoleus Rohdendorf 1961
+
†Kaltanocoleus pospelovi Rohdendorf 1961
G. †Linicupes Ponomarenko et al. 2014
+
†Linicupes yinpinensis Ponomarenko et al. 2014
G. †Maricoleus Shcherbakov 2009
+
†Maricoleus valentinae Shcherbakov 2009
G. †Permocupes Martynov 1932
+
†Permocupes distinctus Martynov 1932
†Permocupes grandis Ponomarenko 1963
†Permocupes latus Ponomarenko 2013
†Permocupes semenovi Martynov 1932
†Permocupes sojanensis Ponomarenko 1963
Invalid names: Permocupoides Martynov 1932 [synonym]
G. †Pintolla Kirejtshuk 2020
+
†Pintolla ponomarenkoi Pinto 1987
G. †Protocupoides Rohdendorf 1961
+
†Protocupoides acer Rohdendorf 1961
†Protocupoides carinatus Rohdendorf 1961
†Protocupoides elongatus Ponomarenko 2013
†Protocupoides esini Ponomarenko 2013
†Protocupoides plavilstshikovi Rohdendorf 1961
†Protocupoides sharovi Rohdendorf 1961
Invalid names: Protocupoides rohdendorfi Pinto 1987 [nomen dubium], Tomiocupes Rohdendorf 1961 [synonym], Tricupes Rohdendorf 1961 [synonym]
G. †Tatarocupes Ponomarenko 2004
+
†Tatarocupes granulatus Ponomarenko 2004
G. †Uralocupes Ponomarenko 1969
+
†Uralocupes major Ponomarenko 1963
Invalid names: Protocupes Rohdendorf 1961 [nomen dubium], Taldycupes chachlovi Rohdendorf 1961 [nomen dubium]
Invalid names: Archicupes reichardti Rohdendorf 1961 [nomen dubium], Kaltanocoleidae Rohdendorf 1961 [synonym], Protocupes martynovi Rohdendorf 1961 [nomen dubium]
G. †Mesoncus Carpenter 1986
+
†Mesoncus striatulus Bode 1953
Invalid names: Loxoncus Bode 1953 [replaced]
G. †Mesorhynchophora Tillyard 1916
G. †Mesotricupes Hong 1982
+
†Mesotricupes lineatus Hong 1982
G. †Mesotylites Bode 1953
+
G. †Metanastes Bode 1953
+
†Metanastes denudatus Bode 1953
G. †Micrelaterium Handlirsch 1906
+
†Micrelaterium triopas Westwood 1854
G. †Microcoleopteron Handlirsch 1906
+
†Microcoleopteron heydeni Weyenbergh 1869
†Microcoleopteron jurassicum Weyenbergh 1869
†Microcoleopteron minimum Oppenheim 1888
G. †Microentomus Poinar 2015
+
†Microentomus epibatus Poinar 2015
G. †Mimelater Handlirsch 1906
+
†Mimelater angulatus Giebel 1856
G. †Nannoodes Handlirsch 1906
+
†Nannoodes pseudocistela Handlirsch 1906
G. †Nebrioides Handlirsch 1906
+
†Nebrioides dobbertinensis Geinitz 1894
Unr. Nubili Latreille 1804
+
Invalid names: Herbicolae Latreille 1804 [empty], Sabulosi Latreille 1804 [empty]
G. †Omogongylus Bode 1953
+
†Omogongylus ovatus Bode 1953
G. †Onkedodimus Handlirsch 1939
+
†Onkedodimus discicollis Handlirsch 1939
G. †Ooidellus Carpenter 1986
+
†Ooidellus denudatus Bode 1953
Invalid names: Ooides Bode 1953 [replaced]
G. †Ooperiglyptus Bode 1953
+
G. †Ooperioristus Bode 1953
+
Tr. †Oropseini Legalov and Kirejtshuk 2016
+
G. †Libanorhinus Kuschel and Poinar 1993
+
†Libanorhinus succinus Kuschel and Poinar 1993
G. †Oropsis Legalov and Kirejtshuk 2016
+
†Oropsis marinae Legalov and Kirejtshuk 2016
G. †Oxycephalites Bode 1953
+
G. †Oxytoroptera Handlirsch 1939
+
†Oxytoroptera mediocris Handlirsch 1939
†Oxytoroptera punctata Ponomarenko 1985
G. †Pachycoleon Handlirsch 1906
+
†Pachycoleon woodlei Westwood 1854
G. †Pachypleurites Haupt 1952
+
†Pachypleurites nodosus Haupt 1952
G. †Paklapwania Lin 1997
+
G. †Palaeotrachys Bode 1953
+
Subfm. †Palaeotylinae Poinar et al. 2018
+
G. †Palaeotylus Poinar et al. 2018
+
†Palaeotylus femoralis Poinar et al. 2018
G. †Pallax Handlirsch 1906
+
†Pallax prevosti Handlirsch 1906
G. †Pantodapus Handlirsch 1906
+
†Pantodapus ewaldi Giebel 1856
†Pantodapus knorri Giebel 1856
†Pantodapus westwoodi Handlirsch 1906
G. †Parabuprestium Handlirsch 1906
+
†Parabuprestium pseudocarabus Handlirsch 1906
†Parabuprestium teleas Westwood 1854
G. †Paracurculium Handlirsch 1906
+
†Paracurculium punctatum Geinitz 1894
G. †Paragrilium Handlirsch 1906
+
†Paragrilium barypus Westwood 1854
G. †Paragyrinus Handlirsch 1906
+
†Paragyrinus dubius Giebel 1856
G. †Parasilphites Handlirsch 1906
+
†Parasilphites angusticollis Oppenheim 1888
G. †Parnidium Geinitz
+
†Parnidium frechi Handlirsch 1906
†Parnidium geinitzi Handlirsch 1906
G. †Parnosoma Carpenter 1986
+
†Parnosoma detectum Bode 1953
G. †Pedinosoma Bode 1953
G. †Peltocoleops Ponomarenko 1990
+
†Peltocoleops onokhojensis Ponomarenko 1990
G. †Periboloptera Handlirsch 1939
+
†Periboloptera rotunda Handlirsch 1939
G. †Peridosoma Carpenter 1986
+
†Peridosoma praecisum Bode 1953
Invalid names: Perosoma Bode 1953 [replaced]
G. †Petalocupes Hong 1982
+
†Petalocupes arcus Hong 1982
G. †Petrorophus Heer 1852
+
†Petrorophus truncatus Heer 1852
G. †Phaulogyrinus Handlirsch 1906
+
†Phaulogyrinus minimus Heer 1864
G. †Pholipheron Bode 1953
+
†Pholipheron armatus Bode 1953
G. †Phytoplesion Bode 1953
+
†Phytoplesion ovatus Bode 1953
G. †Pimeliodes Frič 1901
+
†Pimeliodes parvus Frič 1901
G. †Plastobuprestites Handlirsch 1906
+
†Plastobuprestites elegans Geinitz 1894
G. †Plastonebria Handlirsch 1906
+
†Plastonebria scudderi Geinitz 1894
G. †Pleuralocista Bode 1953
+
G. †Polypamon Handlirsch 1906
+
†Polypamon byrrhoides Geinitz 1894
G. †Praedodromeus Strada et al. 2014
+
†Praedodromeus sangiorgiensis Strada et al. 2014
G. †Prionophana Handlirsch 1906
+
†Prionophana antiqua Giebel 1856
G. †Prochrysomela Handlirsch 1906
+
†Prochrysomela jurassica Oppenheim 1888
G. †Proheuristes Bode 1953
+
†Proheuristes striatus Bode 1953
G. †Prophasis Handlirsch 1906
+
†Prophasis dubia Giebel 1856
†Prophasis ignota Giebel 1856
G. †Prosthecogmus Ponomarenko 1985
+
†Prosthecogmus additus Ponomarenko 1985
G. †Prosthenostictus Handlirsch 1906
+
†Prosthenostictus ungeri Giebel 1856
G. †Protocuneus Cockerell 1915
+
†Protocuneus punctatus Cockerell 1915
G. †Prototoma Heer 1852
+
†Prototoma striata Heer 1852
G. †Pseudobuprestites Handlirsch 1906
+
G. †Pseudocarabites Handlirsch 1906
+
G. †Pseudocurculionites Handlirsch 1906
+
G. †Pseudocymindis Handlirsch 1906
+
†Pseudocymindis antiqua Giebel 1856
G. †Pseudocyphon Handlirsch 1906
+
†Pseudocyphon geinitzi Handlirsch 1906
G. †Pseudoelateropsis Handlirsch 1906
+
Invalid names: Elateropsis Roemer 1876 [replaced]
G. †Pseudohydrophilites Handlirsch 1906
+
G. †Pseudoprionites Handlirsch 1906
+
†Pseudoprionites liasinus Geinitz 1894
G. †Pseudotelephorus Handlirsch 1906
+
†Pseudotelephorus grandis Cockerell 1915
†Pseudotelephorus haueri Giebel 1856
†Pseudotelephorus punctulatus Cockerell 1915
G. †Pseudotenebrio Handlirsch 1906
+
†Pseudotenebrio innominatus Weyenbergh 1869
†Pseudotenebrio relictus Weyenbergh 1869
G. †Pseudus Handlirsch 1906
+
†Pseudus purbeccensis Handlirsch 1906
G. †Rhomaleus Bode 1953
+
†Rhomaleus ornatus Bode 1953
G. †Scalopoides Bode 1953
+
†Scalopoides inscissus Bode 1953
G. †Semiglobus Handlirsch 1906
+
†Semiglobus chrysomelinus Handlirsch 1906
†Semiglobus jurassicus Handlirsch 1906
†Semiglobus neptuni Giebel 1856
G. †Sideriosemion Bode 1953
+
G. †Silphidium Handlirsch 1907
+
†Silphidium priscum Fritsch 1882
Invalid names: Silphites Fritsch 1882 [replaced]
G. †Sitonites Heer 1864
+
†Sitonites melanarius Heer 1864
G. †Smodicoptera Handlirsch 1906
+
†Smodicoptera liasina Heer 1852
Fm. †Sojanocoleidae Martynov 1932
+
G. †Sojanocoleus Martynov 1932
+
†Sojanocoleus reticulatus Martynov 1932
G. †Sphaericites Bode 1953
+
G. †Stenelytron Handlirsch 1906
+
G. †Stictulus Handlirsch 1906
+
†Stictulus brodiei Handlirsch 1906
G. †Strongylites Heer 1864
+
†Strongulites latus Ponomarenko 1985
†Strongylites morio Heer 1864
†Strongylites stygius Heer 1864
G. †Strullendorfia Kuhn 1938
+
†Strullendorfia parva Kuhn 1938
G. †Telephorium Westwood 1854
+
†Telephorium abgarus Westwood 1854
Fm. Tenebrionites Latreille 1802 [darkling beetle]
+
Invalid names: Pimeliariae Latreille 1802 [empty], Tenebrionites Latreille 1802 [empty]
G. †Tetragonides Bode 1953
+
†Tetragonides magnus Bode 1953
G. †Thurmannia Heer 1852
+
†Thurmannia punctulata Heer 1852
G. †Toksunius Ponomarenko et al. 2012
+
G. †Tolype Bode 1953
+
†Tolype rotundata Bode 1953
G. †Triadocoleopteron Zeuner 1930
+
G. †Trialarva Prokin and Bashkuev 2021
+
†Trialarva coburgensis Prokin and Bashkuev 2021
G. †Trichelepturgetes Bode 1953
+
G. †Tripsalis Bode 1953
+
†Tripsalis praecisa Bode 1953
G. †Trixagites Heer 1864
+
†Trixagites floralis Heer 1864
G. †Trochiscites Bode 1953
+
G. †Trochmalus Bode 1953
+
†Trochmalus compressus Bode 1953
Fm. Trogossitarii Latreille 1802 [bark gnawing beetle]
+
G. Parandre Latreille 1802
G. †Trypocoleus Hong 1982
+
†Trypocoleus ramulosus Hong 1982
Fm. †Tshekardocoleidae Rohdendorf 1944
+
G. †Avocoleus Ponomarenko 1969
+
†Avocoleus fractus Kukalová 1968
†Avocoleus neglegens Kukalová 1968
G. †Boscoleus Kukalová 1968
+
†Boscoleus blandus Kukalová 1968
G. †Eocoleus Kukalová 1968
+
†Eocoleus scaber Kukalová 1968
G. †Moravocoleus Kukalová 1968
+
†Moravocoleus permianus Kukalová 1968
G. †Permocoleus Lubkin and Engel 2005
+
†Permocoleus wellingtonensis Lubkin and Engel 2005
G. †Prosperocoleus Kukalová 1968
+
†Prosperocoleus prosperus Kukalová 1968
G. †Retelytron Kukalová 1965
+
†Retelytron conopeum Kukalová 1965
G. †Sylvacoleodes Ponomarenko 1969
+
†Sylvacoleodes admirandus Ponomarenko 1969
G. †Sylvacoleus Ponomarenko 1963
+
†Sylvacoleus richteri Ponomarenko 1963
†Sylvacoleus sharovi Ponomarenko 1963
G. †Tshekardocoleus Rohdendorf 1944
+
†Tshekardocoleus magnus Rohdendorf 1944
†Tshekardocoleus minor Ponomarenko 1963
G. †Umoricoleus Kukalová 1968
+
†Umoricoleus perplex Kukalová 1968
G. †Uralocoleus Zalessky 1947
+
†Uralocoleus splendens Zalessky 1947
G. †Votocoleus Kukalová 1968
+
†Votocoleus submissus Kukalová 1968
Invalid names: Moravocoleidae Kukalová-Peck and Beutel 2012 [synonym], Uralocoleidae Zalessky 1947 [synonym], Uralocoleus ultimus Ponomarenko 2000 [nomen dubium]
G. †Tychon Handlirsch 1908
+
†Tychon antiquum Giebel 1856
G. †Ulomites Tillyard 1916
+
†Ulomites willcoxi Tillyard 1916
G. †Velenovskya Frič 1889
+
†Velenovskya inornata Frič 1889
G. †Wollastonia Heer 1852
+
†Wollastonia ovalis Heer 1852
G. †Xenogyrinus Handlirsch 1906
+
†Xenogyrinus natans Brodie 1845
G. †Xinbinia Hong 1983
+
†Xinbinia foveolata Hong 1983
G. †Xylotupia Handlirsch 1906
+
†Xylotupia brodiei Handlirsch 1906
G. †Yanqingia Hong and Xiao 1997
+
†Yanqingia jurassica Hong and Xiao 1997
G. †Zetemenos Bode 1953
+
†Zetemenos sexlineatus Bode 1953
Invalid names: Acmaeodera perreaui Piton 1940 [nomen dubium], Ancylocheira liasina Giebel 1856 [nomen dubium], Brochocoleus indibili Soriano and Delclos 2006 [nomen dubium], Carabiques Latreille 1802 [empty], Cebrionates Latreille 1802 [empty], Chrysomelinae Latreille 1802 [empty], Cicindeletae Latreille 1802 [empty], Cistela Geoffroy 1762 [nomen oblitum], Clerii Latreille 1802 [empty], Coleopteron depressus Heer 1879 [nomen nudum], Coleopteron interpunctatus Heer 1864 [nomen nudum], Coleopteron laevigatus Heer 1864 [nomen nudum], Coleopteron scabriusculus Heer 1879 [nomen nudum], Coprophagi Latreille 1802 [empty], Criocerina pristina Germar 1813 [nomen dubium], Curculionites liasinus Heer 1852 [nomen dubium], Curculiopsidae Martynov 1937 [empty], Cydnopsis nigromembranacea Jordan 1967 [nomen dubium], Diaperialae Latreille 1802 [empty], Elateridopsidae Zalessky 1929 [invalid subgroup], Eleuterata Fabricius 1775 [synonym], Entomophagi Latreille 1804 [empty], Eodromeus viriosus Zhang 1997 [nomen dubium], Erotilenae Latreille 1802 [empty], Euchroma arvernica Piton 1940 [nomen dubium], Eucoleoptera Engel et al. 2022 [invalid subgroup], Helophorus exilis Heer 1862 [nomen dubium], Helophorus magnus Heer 1862 [nomen dubium], Helopii Latreille 1802 [empty], Helops atticus Redtenbacher 1867 [nomen dubium], Hydrobius godeti Heer 1862 [nomen dubium], Hydrobius obsoletus Heer 1856 [nomen dubium], Hydrocanthares Latreille 1802 [empty], Hydrophilus stenopterus Heer 1862 [nomen dubium], Hydrous ebeninus Statz 1939 [nomen dubium], Hylesinus lineatus Förster 1891 [nomen dubium], Ipidae Bedel 1888 [empty], Isomira aurora Wickham 1914 [nomen dubium], Leptopodocoleidae Hong 1982 [invalid subgroup], Macrogastri Latreille 1802 [empty], Malacodermi Latreille 1802 [empty], Malacodermidae [empty], Megopis lineolatus Piton 1940 [nomen dubium], Metaxycoleoptera Engel et al. 2022 [invalid subgroup], Necrophagi Latreille 1802 [empty], Necticus minutus Aymard 1856 [nomen nudum], Nitidulariae Latreille 1802 [empty], Onthophagus luteus Oustalet 1874 [nomen dubium], Palpatores Latreille 1802 [empty], Platyprosopi Latreille 1804 [empty], Protocoleoptera Tillyard 1924 [empty], Rhynchophori Latreille 1804 [empty], Saperda valdensis Heer 1864 [nomen dubium], Saprophagi Latreille 1804 [empty], Staphyliniae Latreille 1802 [empty], Sternoxi Latreille 1802 [empty], Tridigitati Latreille 1802 [empty], Versicolores Latreille 1804 [empty], Xylophagi Latreille 1802 [empty], Zacoleoptera Engel et al. 2022 [invalid subgroup]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Kiessling 2004, Bush and Bambach 2015

Age range

Maximum range based only on fossils: base of the Asselian to the top of the Holocene or 298.90000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 295.5 Ma

Collections (1991 total)

Oldest occurrences

Time interval Ma Country or state Original ID and collection number
Asselian298.9 - 295.5Germany (Rhineland-Palatinate) Coleopsis archaica (156248)
Asselian298.9 - 295.5Germany (Thüringer Wald) Pachypleurites nodosus (128485)
Asselian298.9 - 295.5Czech Republic Oborocoleus rohdendorfi, Liberocoleus intactus, Moravocoleus perditus, Tshekardocoleidae indet., Boscoleus blandus, Moravocoleus neglegens, Moravocoleus fractus, Votocoleus submissus, Moravocoleus permianus, Retelytron conopeum, Eocoleus scaber, Umoricoleus perplex, Prosperocoleus prosperus (114186)
Artinskian290.1 - 279.3USA (Oklahoma) Permocoleus wellingtonensis (104816 104826)
Artinskian290.1 - 279.3South Africa (Western Cape) Afrocupes firmae (156986)
Kungurian279.3 - 272.3Russian Federation (Suksun) Tshekardocoleidae indet., Sylvacoleus sharovi, Sylvacoleus richteri, Sylvacoleodes admirandus, Tshekardocoleus minor (108206) Tshekardocoleus magnus (116652)
Kungurian279.3 - 272.3Brazil (Rio Grande do Sul) Kaltanicupes ponomarenkoi, Protocupoides rohdendorfi (116121)
Kungurian279.3 - 272.3Russian Federation Maricoleus valentinae (115214) Sylvacoleus richteri, Tshekardocoleus magnus (107787) Uralocoleus splendens (155819)
Roadian272.3 - 268.8Russian Federation Elateridopsis permiensis (156163) Stegosyne rugosa (124611)
Roadian272.3 - 268.8Russian Federation (Kirov) Rossocoleus altus, Kaltanicupes kitjakensis, Kaltanicupes acutus, Kaltanicupes major (105058)
Roadian272.3 - 268.8Russian Federation (Arkangelsk) Permocrossos elongatus (104993) Permocupes semenovi (104716) Permocupes sojanensis, Permocupes grandis (116933) Permocupes sojanensis, Uralocoleus ultimus (104992) Sojanocoleus reticulatus (116874)
Roadian272.3 - 268.8Russian Federation (Kemerovo) Ichthyocupes kuznetskiensis (122572) Kaltanicupes richteri (122568) Protocupes martynovi, Kaltanocoleus pospelovi (122569) Rossocoleus avus, Protocupes martynovi, Eocupes lukjanovitshi, Archicupes jacobsoni, Cytocupoides elongatus (122567) Schizocoleus kuznetskiensis (124604) Uskatocoleus kaltanicus, Carinicupes beckermigdisovae, Schizotaldycupes ananjevi, Asiocoleus novojilovi, Stegocupes efremovi, Archicupes reichardti, Ichthyocupes tyzhnovi, Eocupes cellulosus, Palaeocupes kaltanicus, Cytocupes angustus, Tomiocupes sharovi, Protocupoides plavilstshikovi, Kaltanocoleus pospelovi (104704)
Roadian272.3 - 268.8Russian Federation (Kazan) Permocupoides distinctus, Permocupes semenovi (116877)
Wordian266.9 - 264.28Russian Federation (Kemerovo) Palademosyne martynovae (124612) Schizocupes obrutshevi (124600) Uskatocoleus avitus, Uskatocoleus iljinskiensis, Uskatocoleus surijokovicus, Karakanocoleus janischewskyi, Rossocoleus grandis, Rossocoleus latior, Rossocoleus parvus, Rossocoleus surijokovensis, Rossocoleus latus, Tecticupes heckeri, Stegocupes fedotovi, Tecticupes indistinctus, Tychticupes tchihatcheffi, Taldycupes chachlovi, Protocupes ogloblini (122566) Uskatocoleus convexus, Rossocoleus verrucosus (124603)
Wordian266.9 - 264.28Russian Federation (Udmurtia) Coleoptera hindwing indet., Schizophoridae indet., Rossocoleus novojilovi, Rossocoleus angustus, Protocupoides elongatus, Permocupes latus (103502) Rhombocoleites uralensis, Schizotaldycupes pubescens (104750)
Wordian266.9 - 264.28South Africa Permocupedidae indet. (226396)
Wordian - Capitanian266.9 - 259.51Russian Federation Uskatocoleus kushejakovicus (124610)
Capitanian264.28 - 259.51China (Anhui) Archosyne permiana, Chaocoleus limnebius, Dikerocoleus divisus, Asiocoleopsis hongi, Taldycupes pingi, Linicupes yinpinensis (103239)
Capitanian264.28 - 259.51Russian Federation (Orenburg) Curculiopsis ellipticus (104994) Kargalarva permosialis, Rhombocoleites adumbratus, Tecticupes martynovi, Kaltanicupes kargalensis (104057) Sakmaracoleus orenburgensis, Schizocoleus major, Aenigmocoleus uralensis, Rossocoleus sakmara (152235) Schizocoleus minimus, Schizocoleus robustus, Schizocoleus longus, Schizocoleus glabrus, Erunakicupes brevis, Karakanocoleus europeus, Palademosyne ovum, Rhombocoleus gomankovi, Tetracoleus orenburgensis, Bicoleus kuplensis, Tatarocupes granulatus (104717)
Wuchiapingian259.9 - 254.17Russian Federation (Orenburg) Uskatocoleus uralensis (152236)