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Ferae
Taxonomy
Ferae was named by Linnaeus (1758). It is extant.
It was reranked as the superorder Ferae by Lopatin (2006); it was reranked as the unranked clade Ferae by McKenna (1975), Waddell et al. (2001), Kielan-Jaworowska et al. (2004), Spaulding et al. (2009) and Asher and Helgen (2010).
It was assigned to Unguiculata by Bonaparte (1850); to Monodelphia by Hay (1902); to Therictoidea by Gregory (1910); to Tokotheria by McKenna (1975); to Fereuungulata by Waddell et al. (2001); to Epitheria by Kielan-Jaworowska et al. (2004); to Mammalia by Linnaeus (1758) and Lopatin (2006); to Laurasiatheria by Spaulding et al. (2009); and to Scrotifera by Asher and Helgen (2010).
It was reranked as the superorder Ferae by Lopatin (2006); it was reranked as the unranked clade Ferae by McKenna (1975), Waddell et al. (2001), Kielan-Jaworowska et al. (2004), Spaulding et al. (2009) and Asher and Helgen (2010).
It was assigned to Unguiculata by Bonaparte (1850); to Monodelphia by Hay (1902); to Therictoidea by Gregory (1910); to Tokotheria by McKenna (1975); to Fereuungulata by Waddell et al. (2001); to Epitheria by Kielan-Jaworowska et al. (2004); to Mammalia by Linnaeus (1758) and Lopatin (2006); to Laurasiatheria by Spaulding et al. (2009); and to Scrotifera by Asher and Helgen (2010).
Subtaxa
Synonyms
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Synonymy list
Year | Name and author |
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1758 | Ferae Linnaeus p. 37 |
1850 | Ferae Bonaparte p. 1 |
1902 | Ferae Hay p. 743 |
1910 | Ferae Gregory p. 465 |
1975 | Ferae McKenna p. 42 |
2001 | Ferae Waddell et al. p. 148 |
2004 | Ferae Kielan-Jaworowska et al. pp. 494, 506-507 |
2006 | Ferae Lopatin |
2009 | Ferae Spaulding et al. p. 5 figs. Figure 2 |
2010 | Ferae Asher and Helgen p. 3 figs. Figure 10 |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Unr. Ferae Linnaeus 1758
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Unr. Carnivoramorpha Wyss and Flynn 1993
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Unr. †Carnivoraformes Flynn et al.
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Or. Carnivora Bowdich 1821 [carnivoran]
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G. †Adracon Filhol 1884
Fm. Canidae Fischer de Waldheim 1817 [canid]
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Subfm. †Borophaginae Simpson 1945 [bone-crushing dog]
Subfm. Caninae Gill 1872 [canine]
G. †Chailicyon Chow 1975
G. †Cubacyon Arredondo and Varona 1974
G. Cuon Hodgson 1838 [dhole]
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Invalid names: Anurocyon Heude 1888 [synonym], Chrysaeus Smith 1839 [synonym], Crassicuon Kretzoi 1941 [synonym], Cyon Agassiz 1846 [objective synonym], Primaevus Gray 1843 [synonym], Semicuon Kretzoi 1941 [synonym]
G. †Cynotherium Studiati 1857
G. Dysodus Cope 1879
Subfm. †Hesperocyoninae Tedford 1978
G. †Hydrocyon Lartet 1851
G. †Icticyon Lund 1839
G. Lupulella Hilzheimer 1906
G. †Lupus
G. Lycaon Brookes 1827
G. †Lycorus Bourguignat 1875
G. †Palaeocyon Lund 1843
G. †Plesiocyon Schlosser 1887
G. Schaeffia Hilzheimer 1906
G. Simenia Gray
G. †Synagodus Cope 1879
G. Thous Hamilton Smith 1839
G. Vulpes Frisch 1775 [fox]
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Invalid names: Alopex Kaup 1829 [synonym], Cynalopex Smith 1839 [synonym], Fennecus Desmarest 1804 [synonym], Leucocyon Woldrich 1868 [synonym], Mamvulpesus Herrera 1899 [synonym], Megalotis Smith 1839 [synonym], Vulpis Gray 1821 [synonym], Xenalopex Kretzoi 1954 [synonym]
Invalid names: Amphicyon angustidens Marsh 1871 [nomen vanum], Otocyoninae Trouessart 1885 [empty]
Subor. Caniformia Kretzoi 1943
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Fm. †Amphicyonidae Haeckel 1886 [bear-dog]
Infraor. Canoidea Simpson 1931
G. †Lycophocyon Tomiya 2011
Invalid names: Cynoidea Flower 1869 [empty]
Subor. Feliformia Kretzoi 1945
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Infraor. Aeluroida Flynn and Galiano 1982
Unr. Aeluroidea Flower 1869
G. †Alagtsavbaatar Egi et al. 2016
G. †Asiavorator Spassov and Lange-Badré 1995
Superfm. Feloidea Hay 1930
Fm. Nandiniidae Pocock 1929
Fm. †Nimravidae Cope 1880 [false sabre-tooth]
Fm. †Palaeogalidae Martin and Lim 2001
Invalid names: Didymictida Flynn and Galiano 1982 [empty]
Subor. Fissipedia Blumenbach 1791
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Invalid names: Arctictidae Cope 1889 [empty], Bassarididae Cope 1889 [empty], Cynictidae Cope 1882 [empty]
G. †Monachiopsis Kretzoi 1941
Fm. †Percrocutidae Werdelin and Solounias 1991
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G. †Dinocrocuta Schmidt-Kittler 1976
G. †Percrocuta Kretzoi 1938
G. †Pseudailurus Gervais 1848
Invalid names: Eosictis Scott 1945 [nomen dubium], Felidi Jaekel 1911 [empty], Phocidi Jaekel 1911 [empty], Protelidi Jaekel 1911 [empty], Protoferi Jaekel 1911 [empty], Ursina Gray 1825 [empty], Ursocanidi Jaekel 1911 [empty], Viverra memphitis Linnaeus 1758 [nomen vanum]
G. †Gracilocyon Smith and Smith 2010
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†Gracilocyon igniculus Beard and Dawson 2009
†Gracilocyon rosei Heinrich et al. 2008
†Gracilocyon rundlei Hooker 2010
†Gracilocyon solei Smith and Smith 2010
†Gracilocyon winkleri Gingerich 1983
G. †Miacis Cope 1872
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†Miacis bathygnathus Scott 1888
†Miacis boqinghensis Huang et al. 1999
†Miacis deutschi Gingerich 1983
†Miacis edax Leidy 1872
†Miacis exiguus Matthew and Granger 1915
†Miacis gracilis Clark 1939
†Miacis hookwayi Stock 1934
†Miacis invictus Matthew and Granger 1925
†Miacis latidens Matthew and Granger 1915
†Miacis lushiensis Chow 1975
†Miacis parvivorus Cope 1872
†Miacis petilus Gingerich 1983
†Miacis rundlei Hooker 2010
†Miacis vorax Leidy 1872
†Miacis vulpinus Scott and Osborn 1887
Invalid names: Harpalodon Marsh 1872 [synonym], Prodaphaenus Wortman and Matthew 1899 [synonym]
G. †Neovulpavus Wortman 1901
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†Neovulpavus mccarrolli Tomiya et al. 2021
†Neovulpavus washakius Wortman 1901
G. †Procynodictis Wortman and Matthew 1899
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†Procynodictis progressus Stock 1935
†Procynodictis vulpiceps Wortman and Matthew 1899
Invalid names: Plesiomiacis Simpson 1945 [synonym]
G. †Quercygale Kretzoi 1945
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†Quercygale angustidens Filhol 1872
†Quercygale hastingsae Davies 1884
†Quercygale helvetica Rütimeyer 1862
†Quercygale smithi Solé 2014
Invalid names: Humbertia De Beaumont 1965 [objective synonym]
G. †Uintacyon Leidy 1873
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†Uintacyon acutus Thorpe 1923
†Uintacyon asodes Gazin 1952
†Uintacyon gingerichi Heinrich et al. 2008
†Uintacyon hookeri Solé 2014
†Uintacyon jugulans Matthew 1909
†Uintacyon massetericus Cope 1882
†Uintacyon rudis Rose 1981
G. †Vassacyon Matthew 1909
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†Vassacyon bowni Heinrich et al. 2008
†Vassacyon prieuri Solé et al. 2016
†Vassacyon promicrodon Wortman and Matthew 1899
†Vassacyon taxidiotis Solé et al. 2013
G. †Vulpavus Marsh 1871
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†Vulpavus completus Matthew 1909
†Vulpavus farsonensis Gunnell 1998
†Vulpavus hargeri Wortman 1901
†Vulpavus ovatus Matthew 1909
†Vulpavus palustris Marsh 1871
†Vulpavus profectus Matthew 1909
G. †Oodectes Wortman 1901
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†Oodectes herpestoides Wortman 1901
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Invalid names: Oodectes proximus Matthew 1909 [synonym], Paeneprolimnocyon amissadomus Guthrie 1967 [synonym], Triacodon nanus Marsh 1872 [synonym]
†Oodectes pugnax Wortman and Matthew 1899
Invalid names: Paeneprolimnocyon Guthrie 1967 [synonym]
Fm. †Viverravidae Wortman and Matthew 1899
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G. †Didymictis Cope 1875
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†Didymictis altidens Cope 1880
†Didymictis dellensis Dorr 1952
†Didymictis leptomylus Cope 1880
†Didymictis proteus Polly 1997
†Didymictis vancleveae Robinson 1966
G. †Protictis Matthew
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Subg. †Protictis (Bryanictis) MacIntyre 1966
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†Bryanictis microlestes Simpson 1935
†Bryanictis paulus Meehan and Wilson 2002
Subg. †Protictis (Protictis) Matthew 1937
Subg. †Protictis (Protictoides) Flynn and Galiano 1982
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†Protictoides aprophatos Flynn and Galiano 1982
†Protictis agastor Gingerich and Winkler 1985
†Protictis haydenianus Cope 1882
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Invalid names: Didymictis primus Cope 1884 [synonym], Prolimnocyon macfaddeni Rigby, Jr. 1980 [synonym]
†Protictis minor Meehan and Wilson 2002
†Protictis paralus Holtzman 1978
†Protictis simpsoni Meehan and Wilson 2002
G. †Simpsonictis MacIntyre 1962
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†Simpsonictis jaynanneae Rigby, Jr. 1980
†Simpsonictis pegus Gingerich and Winkler 1985
†Simpsonictis tenuis Simpson 1935
G. †Viverravus Marsh 1872
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†Viverravus acutus Matthew and Granger 1915
†Viverravus gracilis Marsh 1872
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Invalid names: Didymictis dawkinsianus Cope 1881 [synonym], Harpalodon vulpinus Marsh 1872 [synonym], Triacodon fallax Marsh 1871 [synonym], Ziphacodon rugatus Marsh 1872 [synonym]
†Viverravus lawsoni Hooker 2010
†Viverravus laytoni Gingerich and Winkler 1985
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Invalid names: Viverravus bowni Gingerich 1987 [synonym]
†Viverravus lutosus Gazin 1952
†Viverravus minutus Wortman 1901
†Viverravus nitidus Marsh 1872
†Viverravus politus Matthew and Granger 1915
†Viverravus rosei Polly 1997
†Viverravus schaffi Gingerich and Winkler 1985
†Viverravus sicarius Matthew 1909
Invalid names: Ziphacodon Marsh 1872 [synonym]
Invalid names: Deltatherium durini Van Valen 1978 [nomen dubium], Didymictidae Flynn and Galiano 1982 [synonym]
Or. †Creodonta Cope 1875 [creodont]
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Or. †Hyaenodonta Van Valen 1967
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Superfm. †Hyaenodontoidea Leidy 1869
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Fm. †Hyaenodontidae Leidy 1869
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G. †Acrohyaenodon Ameghino 1904
G. †Ahknatenavus Holroyd 1999
Subfm. †Arfiinae Solé et al. 2014
G. †Boritia Solé et al. 2014
G. †Cartierodon Solé and Mennecart 2019
G. †Eurotherium Polly and Lange-Badré 1993
Subfm. †Hyaenodontinae Trouessart 1885
G. †Hyaenodontipus
Subfm. †Indohyaenodontinae Solé et al. 2014
G. †Ischnognathus Stovall 1948
Subfm. †Koholiinae Crochet 1988
G. †Leonhardtina Matthes 1952
G. †Matthodon Lange-Badré and Haubold 1990
G. †Neohyaenodon Thorpe 1922
G. †Paracynohyaenodon Martin 1906
G. †Preregidens Solé et al. 2015
Unr. †Prionogalidae Morales et al. 2008
G. †Protopsalis Cope 1880
G. †Quercytherium Filhol 1880
G. †Schizophagus Lange-Badre 1975
Subfm. †Sinopinae Solé et al. 2014
G. †Thereutherium Filhol 1876
G. †Tinerhodon Gheerbrant 1995
Invalid names: Hyaenaelurinae Pilgrim 1932 [empty]
Subfm. †Proviverrinae Schlosser 1886
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G. †Alienetherium Lange-Badre 1981
G. †Consobrinus Lange-Badre 1979
G. †Lesmesodon Morlo and Habersetzer 1999
G. †Minimovellentodon Solé et al. 2014
G. †Morlodon Solé 2013
G. †Paenoxyaenoides Lange-Badre 1979
G. †Parvagula Lange-Badré 1987
G. †Praecodens Lange-Badre 1981
G. †Proviverra Rütimeyer 1862
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Invalid names: Geiselotherium Matthes 1952 [synonym], Mimocyon Peterson 1919 [synonym], Prorhyzaena Rutimeyer 1891 [synonym]
Superfm. †Hyainailouroidea Borths et al. 2016
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G. †Metasinopa Osborn 1909
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†Metasinopa fraasii Osborn 1909
†Metasinopa napaki Savage 1965
†Metasinopa osborni Holroyd 1994
G. †Paratritemnodon Ranga Rao 1973
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†Paratritemnodon indicus Ranga Rao 1973
†Paratritemnodon jandewalensis Thewissen et al. 2001
Unr. †Teratodontinae Savage 1965
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G. †Anasinopa Savage 1965
G. †Brychotherium Holroyd 1994
G. †Buhakia Morlo et al. 2007
G. †Dissopsalis Pilgrim 1910
G. †Ekweeconfractus Flink et al. 2021
G. †Furodon Solé et al. 2014
G. †Glibzegdouia Crochet et al. 2001
G. †Masrasector Simons and Gingerich 1974
G. †Pakakali Borths and Stevens 2017
G. †Teratodon Savage 1965
Subfm. †Limnocyoninae Wortman 1902
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G. †Limnocyon Marsh 1872
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†Limnocyon cuspidens Morlo and Gunnell 2005
†Limnocyon verus Marsh 1872
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Invalid names: Limnocyon riparius Marsh 1872 [synonym], Triacodon grandis Marsh 1872 [synonym]
Invalid names: Telmatocyon Marsh 1899 [synonym]
G. †Prolimnocyon Matthew and Granger 1915
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†Prolimnocyon antiquus Matthew and Granger 1915
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Invalid names: Prolimnocyon elisabethae Gazin 1952 [synonym]
†Prolimnocyon atavus Matthew and Granger 1915
†Prolimnocyon chowi Meng et al. 1998
†Prolimnocyon eerius Gingerich 1989
†Prolimnocyon haematus Gingerich and Deutsch 1989
G. †Thinocyon Marsh 1872
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†Thinocyon sichowensis Chow 1975
†Thinocyon velox Marsh 1872
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Invalid names: Thinocyon cledensis Matthew 1909 [synonym], Thinocyon minimus Matthew 1909 [synonym], Thinocyon mustelinus Matthew 1909 [synonym], Thinocyon wyomingensis Morlo et al. 2000 [synonym]
Invalid names: Limnocyonini Wortman 1902 [empty], Machaeroidini Matthew 1909 [empty]
Unr. †Oxyaenodonta Van Valen 1971
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Fm. †Oxyaenidae Cope 1877
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G. †Dipsalidictis Matthew and Granger 1915
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†Dipsalidictis aequidens Matthew and Granger 1915
†Dipsalidictis krausei Gunnell and Gingerich 1991
†Dipsalidictis platypus Matthew and Granger 1915
†Dipsalidictis transiens Matthew and Granger 1915
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Invalid names: Dipsalidictis amplus Jepsen 1930 [synonym]
Invalid names: Dipsalidictides Denison 1938 [synonym]
G. †Dormaalodon Lange-Badré 1987
G. †Galethylax Gervais 1850
G. †Machaeroides Matthew 1909
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†Machaeroides eothen Matthew 1909
†Machaeroides simpsoni Dawson et al. 1986
Subfm. †Oxyaeninae Trouessart 1885
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G. †Malfelis Stucky and Hardy 2007
G. †Patriofelis Leidy 1870
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Invalid names: Aelurotherium Adams 1896 [synonym], Limnofelis Marsh 1872 [synonym], Oreocyon Marsh 1872 [synonym]
G. †Sarkastodon Granger 1938
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†Sarkastodon henanensis Tong and Lei 1986
†Sarkastodon mongoliensis Granger 1938
Invalid names: Ambloctonidae Cope 1877 [empty], Amblyctonidae Cope 1877 [empty], Arctocyonoidae Trouessart 1885 [empty], Hyaenodontida Leidy 1869 [empty], Limnocyonidae [empty], Oxyaenoidae Cope 1877 [empty], Uintacyonidae Hay 1902 [empty]
Unr. †Pholidotamorpha Gaudin et al. 2009
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Subor. †Palaeanodonta Matthew 1918
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Fm. †Epoicotheriidae Simpson 1927
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G. †Alocodontulum Rose et al. 1978
G. †Epoicotherium Simpson 1927
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†Epoicotherium unicum Douglass 1905
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Invalid names: Pseudochrysochloris yoderensis Turnbull and Reed 1967 [synonym]
Invalid names: Pseudochrysochloris Turnbull and Reed 1967 [synonym], Xenotherium Douglass 1905 [replaced]
G. †Tetrapassalus Simpson 1959
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†Tetrapassalus mckennai Simpson 1959
†Tetrapassalus proius West 1973
G. †Tubulodon Jepsen 1932
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†Tubulodon atopum Rose et al. 1978
†Tubulodon pearcei Gazin 1952
†Tubulodon taylori Jepsen 1932
†Tubulodon woodi Guthrie 1967
Invalid names: Alocodon Rose et al. 1977 [synonym], Pentapassalus Gazin 1952 [synonym]
Fm. †Escavadodontidae Rose and Lucas 2000
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Fm. †Metacheiromyidae Wortman 1903
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G. †Metacheiromys Wortman 1903
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†Metacheiromys marshi Wortman 1903
†Metacheiromys tatusia Osborn 1904
G. †Palaeanodon Matthew 1918
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†Palaeanodon ignavus Matthew 1918
†Palaeanodon nievelti Gingerich 1989
†Palaeanodon parvulus Matthew 1918
Or. Pholidota Weber 1904 [pangolin]
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Unr. †Eupholidota Gaudin et al. 2009
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Unr. †Manoidea Gaudin et al.
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G. †Euromanis Storch and Martin 1994
Fm. Manidae Gray 1821
Fm. †Patriomanidae Szalay and Schrenk 1998
Invalid names: Eomanidae Storch 2003 [empty], Squamata Huxley 1872 [synonym]
Invalid names: Ostentoria [synonym]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
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Source: s = species, subc = subclass, c = class, subp = subphylum | |||||
References: Martin 1990, Carroll 1988, Hendy et al. 2009, Lillegraven 1979, Ji et al. 2002 |
Age range
Maximum range based only on fossils: base of the Late/Upper Maastrichtian to the top of the Holocene or 70.60000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 66.0 Ma
Minimum age of oldest fossil (stem group age): 66.0 Ma
Collections (6206 total)
Oldest occurrences | |||
Time interval | Ma | Country or state | Original ID and collection number |
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Lancian | USA (Montana) | Cimolestes magnus (14516 14532) | |
Lancian | Canada (Alberta) | Cimolestes magnus (14630 14633 14635 14637) | |
Lancian | Canada (Saskatchewan) | Cimolestes magnus (14558) | |
Lancian | USA (Wyoming) | Cimolestes magnus (14519 14598) | |
Late/Upper Maastrichtian | USA (North Dakota) | Cimolestes magnus (45144) | |
Puercan | Canada (Saskatchewan) | Carnivora indet., Ravenictis krausei (14768) | |
Early/Lower Paleocene | China (Anhui) | Pappictidops orientalis (132943 146556) | |
Early/Lower Paleocene | China (Guangdong) | Pappictidops obtusus, Pappictidops acies (140273 176607) | |
Danian | USA (North Dakota) | Protictis paralus, Miacidae indet., Viverravinae indet. (14962) | |
Cenozoic | Mongolia (South Gobi Aimag) | Carnivora indet. (137861) | |
To2 | Canada (Alberta) | Protictis sp., Simpsonictis jaynanneae (14894) | |
Torrejonian | Canada (Alberta) | Bryanictis microlestes (143200) Protictis sp. (14822) Simpsonictis jaynanneae, Creodonta indet. (14828) | |
Torrejonian | USA (Texas) | Bryanictis sp. (14829) Protictis sp. (14887) | |
Torrejonian | USA (Wyoming) | Bryanictis microlestes, Protictis haydenianus, Simpsonictis tenuis, Protictis sp. (14912) Protictis haydenianus, Prolimnocyon macfaddeni, Simpsonictis jaynanneae, Bryanictis vanvaleni (14886) Protictis haydenianus, Protictis microlestes (14896) Protictis microlestes (14899 14900 14901 14904 14905) Protictis sp., Protictis microlestes (14897 14898) | |
Torrejonian | USA (Montana) | Didymictis haydenianus (14919 14920 14923) Didymictis microlestes (14917) Didymictis microlestes, Didymictis haydenianus (14913) Protictis haydenianus, Ictidopappus mustelinus, Protictis microlestes, Protictis tenuis, Bryanictis vanvaleni (14906) | |
Torrejonian | USA (New Mexico) | Bryanictis paulus, Didymictis sp. a, Didymictis sp. b (14816) Didymictis haydenianus (14819 14873) Protictis haydenianus (14812 14815 14824 14832 14862) Protictis haydenianus, Escavadodon zygus (14835) Protictis haydenianus, Protictis sp. (14865) Protictis haydenianus, Protictis vanvaleni (14833 14893) Protictis sp. (14868) Protictis vanvaleni (14825) | |
Torrejonian | USA (Utah) | Protictis haydenianus (14830) | |
Torrejonian - Tiffanian | USA (Wyoming) | Intyrictis vanvaleni (14853) | |
Middle Paleocene | China (Guangdong) | Ernanodon antelios (132935 176606) | |
Ti1 | Canada (Alberta) | Simpsonictis tenuis, Simpsonictis pegus, Pristinictis connata, Creodonta indet. (14939) |