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Coelurosauria (coelurosaur)

Reptilia

Taxonomy
Coelurosauria was named by Huene (1914). It is extant.

It was reranked as the infraorder Coelurosauria by Roxo (1937), Lapparent (1947), Delair (1960), Simmons (1965), Romer (1966), Charig (1967), Reig (1970), Haubold (1971), Ellenberger (1972), Delair (1973), Haubold (1974), Anderson and Cruickshank (1978), Baird and Horner (1979), Benton (1979), McGinnis (1982), Battail (1986), Buscalioni and Sanz (1987); it was reranked as the superfamily Coelurosauria by Kuhn (1946), Parsch (1963); it was synonymized subjectively with Coeluroidea by Tatarinov (1964), Maleev (1968); it was synonymized subjectively with Coeluria by Kuhn (1966); it was reranked as the infraorder Coelurosaurida by Kuhn (1971); it was reranked as the unranked clade Coelurosauria by Gauthier and Padian (1985), Gauthier (1986), Gauthier et al. (1989), Osmolska (1990), Novas (1992), Chatterjee (1993), Sereno et al. (1994), Holtz (1994), Pérez-Moreno et al. (1994), Sereno (1995), Holtz and Padian (1995), Holtz (1995), Currie (1995), Sereno et al. (1996), Holtz (1996), Currie (1997), Hutchinson and Padian (1997), Sereno (1997), Sereno (1998), Holtz (1998), Sereno (1999), Holtz (2000), Rauhut (2003), Holtz et al. (2004), Yates (2005), Coria and Currie (2006), Sánchez-Hernández et al. (2007), Zhang et al. (2008), Bittencourt and Langer (2011), Zanno et al. (2011), Carrano et al. (2012), Xu et al. (2012), Choiniere et al. (2013), Hendrickx and Mateus (2014), Lü and Brusatte (2015), Pei et al. (2017), Sereno (2017), Xu et al. (2017), Yao et al. (2019), Aranciaga Rolando et al. (2022), Porfiri et al. (2024).

It was assigned to Saurischia by Huene (1914), Huene (1914), Huene (1914), Huene (1914), Huene (1914), Matthew (1915), Huene (1920), Nopcsa (1923), Lull (1924), Huene (1926), Huene (1926), Huene (1927), Parks (1928), Huene (1929), Huene (1929), Kuhn (1939), Huene (1948), Bock (1952), Huene (1954), Huene (1958), Krebs (1974), Bonaparte (1978), Zhao (1983); to Theropoda by Roxo (1937), Kuhn (1946), Lapparent (1947), Delair (1960), Parsch (1963), Simmons (1965), Romer (1966), Charig (1967), Reig (1970), Kuhn (1971), Haubold (1971), Ellenberger (1972), Delair (1973), Haubold (1974), Anderson and Cruickshank (1978), Baird and Horner (1979), Benton (1979), McGinnis (1982), Battail (1986), Buscalioni and Sanz (1987); to Neotetanurae by Sereno et al. (1994), Sereno et al. (1996), Sereno (1997), Sereno (1998), Sereno (1999); to Theropoda by Lull (1953), Romer (1956), Estes (1964), Walker (1964), Colbert (1964), Colbert and Russell (1969), Swinton (1970), Steel (1970), Colbert (1970), Osmólska et al. (1972), Barsbold (1976), Barsbold (1977), Ostrom (1980), Santafé et al. (1982), Dong et al. (1983), Barsbold (1983), Russell (1984), Zinke (1998), Gao and Liu (2005), Xu et al. (2010), Xu et al. (2011), Xu et al. (2012), Agnolin et al. (2012); to Avetheropoda by Holtz (1994), Holtz (1995), Holtz (1996), Holtz (2000), Holtz et al. (2004), Yates (2005), Carrano et al. (2012), Hendrickx and Mateus (2014); to Theropoda by Gauthier and Padian (1985), Gauthier et al. (1989), Currie (1995), Zhang et al. (2008), Zanno et al. (2011), Xu et al. (2011), Loewen et al. (2013), Lü and Brusatte (2015), Pei et al. (2017), Sereno (2017), Xu et al. (2017), Yao et al. (2019); and to Tetanurae by Gauthier (1986), Benton (1990), Osmolska (1990), Novas (1992), Chatterjee (1993), Pérez-Moreno et al. (1994), Sereno (1995), Holtz and Padian (1995), Currie (1997), Hutchinson and Padian (1997), Holtz (1998), Rauhut (2003), Coria and Currie (2006), Sánchez-Hernández et al. (2007), Novas et al. (2008), Bittencourt and Langer (2011), Choiniere et al. (2013), Aranciaga Rolando et al. (2022), Porfiri et al. (2024).

Synonyms
Synonymy list
YearName and author
1866Calamospondylus oweni Anonymous p. 383
1866Calamospondylus Fox p. 740
1866Calamospondylus oweni Fox p. 740
1883Megalosaurus lonzeensis Dollo pp. 212-214 fig. 4
1909Megalosaurus lonzeensis Dollo p. 102
1914Coelurosauria Huene p. 11
1915Coelurosauria Matthew p. 32
1920Coelurosauria Huene p. 161
1923Coelurosauria Nopcsa p. 125
1924Coelurosauria Lull p. 231
1926Ornithomimidorum lonzeensis Huene pp. 87-88
1926Coelurosauria Huene pp. 103-104
1927Coelurosauria Huene p. 256
1928Coelurosauria Parks p. 3
1929Coelurosauria Huene p. 155
1937Coelurosauria Roxo p. 44
1939Coelurosauria Kuhn p. 33
1939Struthiomimus lonzeensis Kuhn p. 54
1946Coelurosauria Kuhn p. 65
1947Coelurosauria Lapparent pp. 16-17
1947Megalosaurus lonzeensis Lapparent p. 20
1948Coelurosauria Huene p. 88
1952Coelurosauria Bock p. 401
1953Coelurosauria Lull p. 123
1954Coelurosauria Huene p. 45
1956Coelurosauria Romer p. 610
1958Coelurosauria Huene p. 205
1960Coelurosauria Delair p. 76
1963Coelurosauria Parsch p. 4
1964Coelurosauria Colbert p. 21 fig. 6
1964Coelurosauria Estes p. 142
1964Coelurosauria Walker p. 105
1965Coelurosauria Simmons p. 53
1966Coelurosauria Romer p. 369
1967Coelurosauria Charig p. 713
1969Coelurosauria Colbert and Russell p. 45
1970Coelurosauria Colbert p. 27
1970Coelurosauria Reig p. 277
1970Coelurosauria Steel p. 7
1970Megalosaurus lonzeensis Steel p. 35
1970Coelurosauria Swinton p. 117
1971Coelurosauria Haubold p. 65
1971Coelurosaurida Kuhn p. 15
1972Coelurosauria Ellenberger p. 152
1972Coelurosauria Osmólska et al. p. 105
1973Coelurosauria Delair p. 1
1974Coelurosauria Haubold p. 41
1974Coelurosauria Krebs p. 22
1976Coelurosauria Barsbold p. 74
1977Coelurosauria Barsbold p. 50
1978Coelurosauria Anderson and Cruickshank p. 31
1978Coelurosauria Bonaparte p. 316
1979Coelurosauria Baird and Horner p. 9
1979Coelurosauria Benton p. 142
1980Coelurosauria Ostrom p. 21
1982Coelurosauria McGinnis p. 28
1982Coelurosauria Santafé et al. p. 81
1983Coelurosauria Barsbold p. 86
1983Coelurosauria Dong et al.
1983Coelurosauria Zhao p. 296
1984Coelurosauria Russell p. 28
1985Coelurosauria Gauthier and Padian pp. 189-190 fig. 2
1986Coelurosauria Battail p. 47
1986Coelurosauria Gauthier p. 54 fig. 7
1987Coelurosauria Buscalioni and Sanz p. 66
1989Coelurosauria Gauthier et al. p. 347
1990Coelurosauria Benton p. 19
1990Coelurosauria Osmolska p. 149
1992Coelurosauria Novas p. 144
1993Coelurosauria Chatterjee p. 283
1994Coelurosauria Holtz
1994Coelurosauria Pérez-Moreno et al. p. 184 fig. 1
1994Coelurosauria Sereno et al. p. 268 fig. 3
1995Coelurosauria Currie p. 25A
1995Coelurosauria Holtz, Jr. p. 35A
1995Coelurosauria Holtz, Jr. and Padian p. 35A
1995Coelurosauria Sereno p. 53A
1996Coelurosauria Holtz, Jr. p. 42A
1996Calamospondylus oweni Hutt et al. p. 636
1996Coelurosauria Sereno et al. p. 989 fig. 4
1997Coelurosauria Currie p. 732
1997Coelurosauria Hutchinson and Padian p. 129
1997Calamospondylus oweni Norman p. 784
1997Coelurosauria Sereno p. 444
1998Coelurosauria Holtz, Jr. p. 39
1998Coelurosauria Sereno p. 64
1998Coelurosauria Zinke p. 181
1999Calamospondylus oweni Naish pp. 367, 371
1999Coelurosauria Sereno p. 2139 fig. 2
2000Coelurosauria Holtz, Jr. p. 23 fig. 5
2003Coelurosauria Rauhut p. 143
2004Coelurosauria Holtz, Jr. et al. p. 76
2005Coelurosauria Gao and Liu p. 313
2005Coelurosauria Yates p. 118 fig. 9
2006Coelurosauria Coria and Currie p. 108 fig. 35
2007Coelurosauria Sánchez-Hernández et al. p. 192
2008Coelurosauria Novas et al. p. 469
2008Coelurosauria Zhang et al. p. 1105
2010Coelurosauria Xu et al. p. 2
2011Coelurosauria Bittencourt and Langer p. 35
2011Coelurosauria Xu et al. p. 2338
2011Coelurosauria Xu et al. p. 2
2011Coelurosauria Zanno et al. p. 3
2012Coelurosauria Agnolin et al. p. 35
2012Coelurosauria Carrano et al. p. 248 fig. 7
2012Coelurosauria Xu et al. p. 141
2013Coelurosauria Choiniere et al.
2013Coelurosauria Loewen et al. p. 2
2014Coelurosauria Hendrickx and Mateus
2015Coelurosauria Lü and Brusatte
2017Coelurosauria Pei et al.
2017Coelurosauria Sereno
2017Coelurosauria Xu et al.
2019Coelurosauria Yao et al.
2022Coelurosauria Aranciaga Rolando et al.
2024Coelurosauria Porfiri et al. p. 5

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
—Osteichthyes()
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
—Tetrapoda
—Reptiliomorpha
—Anthracosauria
subclassAmphibiosauriaKuhn 1967
—Cotylosauria()
—Amniota
—Sauropsida
classReptilia
RankNameAuthor
subclassEureptilia()
—Romeriida
—Diapsida()
—Archosauromorpha(Huene 1946)
—Crocopoda
—ArchosauriformesGauthier 1986
—Eucrocopoda
—Archosauria()
informalAvemetatarsalia
—Ornithodira
—Dinosauromorpha
—Dinosauriformes
—Dinosauria()
—Theropoda()
—Neotheropoda
—AverostraPaul 2002
—Tetanurae
—Coelurosauria()
—Coelurosauria()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Unr. Coelurosauria Huene 1914 [coelurosaur]
show all | hide all
Infraor. †Alvarezsauria Bonaparte 1991
+
Unr. †Alvarezsauroidea Choiniere et al. 2010
+
Fm. †Alvarezsauridae Bonaparte 1991
+
G. †Achillesaurus Martinelli and Vera 2007
+
†Achillesaurus manazzonei Martinelli and Vera 2007
G. †Albertonykus Longrich and Currie 2009 [Alberta claw]
+
†Albertonykus borealis Longrich and Currie 2009
G. †Alnashetri Makovicky et al. 2012
+
†Alnashetri cerropoliciensis Makovicky et al. 2012
G. †Alvarezsaurus Bonaparte 1991
+
†Alvarezsaurus calvoi Bonaparte 1991
G. †Elopteryx Andrews 1913
+
†Elopteryx nopcsai Andrews 1913
Subfm. †Parvicursorinae Karhu and Rautian 1996
+
G. †Albinykus Nesbitt et al. 2011
G. †Ceratonykus Alifanov and Barsbold 2009
G. †Dzharaonyx Averianov and Sues 2022
G. †Jaculinykus Kubo et al. 2023
G. †Khulsanurus Averianov and Lopatin 2021
G. †Linhenykus Xu et al. 2011
Unr. †Mononykini Chiappe et al. 1998
G. †Mononykus Perle et al. 1993
+
Invalid names: Mononychus Perle et al. 1993 [replaced]
G. †Ondogurvel Averianov and Lopatin 2022
G. †Parvicursor Karhu and Rautian 1996
G. †Shuvuuia Chiappe et al. 1998
G. †Xixianykus Xu et al. 2010
Invalid names: Ceratonykini Agnolin et al. 2012 [empty]
G. †Trierarchuncus Fowler et al. 2020
+
†Trierarchuncus prairiensis Fowler et al. 2020
G. †Bannykus Xu et al. 2018
+
†Bannykus wulatensis Xu et al. 2018
G. †Bonapartenykus Agnolin et al. 2012
+
†Bonapartenykus ultimus Agnolin et al. 2012
G. †Haplocheirus Choiniere et al. 2010
+
†Haplocheirus sollers Choiniere et al. 2010
G. †Nemegtonykus Lee et al. 2019
+
†Nemegtonykus citus Lee et al. 2019
G. †Patagonykus Novas 1994
+
†Patagonykus puertai Novas 1994
G. †Qiupanykus Lü et al. 2018
+
†Qiupanykus zhangi Lü et al. 2018
G. †Tugulusaurus Dong 1973
+
†Tugulusaurus faciles Dong 1973
G. †Xiyunykus Xu et al. 2018
+
†Xiyunykus pengi Xu et al. 2018
Invalid names: Patagonykinae Agnolin et al. 2012 [empty]
G. †Aorun Choiniere et al. 2013
+
†Aorun zhaoi Choiniere et al. 2013
Fm. †Arriagadoolithidae Agnolin et al. 2012
+
G. †Arriagadoolithus Agnolin et al. 2012
+
†Arriagadoolithus patagoniensis Agnolin et al. 2012
G. †Shishugounykus Qin et al. 2019
+
†Shishugounykus inexpectus Qin et al. 2019
G. †Aratasaurus Sayão et al. 2020
+
†Aratasaurus museunacionali Sayão et al. 2020
G. †Bicentenaria Novas et al. 2012
+
†Bicentenaria argentina Novas et al. 2012
G. †Brasileosaurus Huene 1931
+
G. †Bressanichnus Calvo 1991
+
G. †Britoichnium Bertini and Carvalho 2001
+
†Britoichnium coelurosauroides Bertini and Carvalho 2001
G. †Coelurus Marsh 1879
+
†Coelurus fragilis Marsh 1879
+
Invalid names: Coelurus agilis Marsh 1884 [objective synonym]
Invalid names: Coelurus gracilis Marsh 1888 [nomen dubium]
Fm. †Compsognathidae Cope 1871 [compsognathid]
+
G. †Aniksosaurus Martínez and Novas 2006
+
†Aniksosaurus darwini Martínez and Novas 2006
G. †Aristosuchus Seeley 1887
+
†Aristosuchus pusillus Owen 1876
Subfm. †Compsognathinae Nopcsa 1923
+
G. †Saurornithoides Nopcsa 1928
G. †Compsognathus Wagner 1861 [elegant jaw]
+
†Compsognathus longipes Wagner 1861
+
Invalid names: Compsognathus corallestris Bidar et al. 1972 [synonym]
G. †Huaxiagnathus Hwang et al. 2004
+
†Huaxiagnathus orientalis Hwang et al. 2004
G. †Mirischia Naish et al. 2004
+
†Mirischia asymmetrica Naish et al. 2004
G. †Ornitholestes Osborn 1903
+
†Ornitholestes hermanni Osborn 1903
G. †Scipionyx Dal Sasso and Signore 1996
+
†Scipionyx samniticus Dal Sasso and Signore 1996
G. †Sinocalliopteryx Ji et al. 2007
+
†Sinocalliopteryx gigas Ji et al. 2007
G. †Xunmenglong Xing et al. 2020
+
†Xunmenglong yingliangis Xing et al. 2020
Invalid names: Beipiaognathus Hu et al. 2016 [nomen dubium]
G. †Cridotrisauropus Ellenberger 1965
+
†Cridotrisauropus cruentus Ellenberger 1970
†Cridotrisauropus ginsburgi Ellenberger 1965
†Cridotrisauropus minor Ellenberger 1965
†Cridotrisauropus unguiferus Ellenberger 1965
G. †Deferrariischnium Calvo 1991
+
G. †Juravenator Göhlich and Chiappe 2006
+
†Juravenator starki Göhlich and Chiappe 2006
G. †Kainomoyenisauropus Ellenberger 1970
+
†Kainomoyenisauropus ranivorus Ellenberger 1970
G. †Kol Turner et al. 2009
+
†Kol ghuva Turner et al. 2009
G. †Lapparentichnus Haubold 1971
+
G. †Lourinhanosaurus Mateus 1998
+
Unr. Maniraptora Gauthier 1986 [maniraptoran]
+
Fm. †Anchiornithidae Foth and Rauhut 2017
+
G. †Ostromia Foth and Rauhut 2017
+
†Ostromia crassipes Meyer 1857
Unr. Avialae Gauthier 1986 [avialan]
+
G. †Alcmonavis Rauhut et al. 2019
+
†Alcmonavis poeschli Rauhut et al. 2019
G. †Archaeopteryx Meyer 1861
+
†Archaeopteryx albersdoerferi Kundrát et al. 2019 [Ancient bird]
+
Invalid names: Archaeopteryx bavarica Wellnhofer 1993 [synonym], Archaeopteryx macrura Owen 1862 [synonym], Archaeopteryx oweni Petronievics 1921 [synonym], Archaeornis siemensii Dames 1897 [synonym], Griphosaurus problematicus Woodward 1862 [synonym], Pterodactylus crassipes Meyer 1857 [synonym], Wellnhoferia grandis Elzanowski 2001 [synonym]
Invalid names: Archaeornis Petronievics 1917 [synonym], Griphosaurus Wagner 1861 [synonym], Jurapteryx Howgate 1985 [synonym]
G. †Aurornis Godefroit et al. 2013
+
†Aurornis xui Godefroit et al. 2013
G. †Baminornis Chen et al. 2025
+
†Baminornis zhenghensis Chen et al. 2025
G. †Dalianraptor Gao and Liu 2005
+
†Dalianraptor cuhe Gao and Liu 2005
G. †Fujianvenator Xu et al. 2023
+
†Fujianvenator prodigiosus Xu et al. 2023
G. †Fukuipteryx Imai et al. 2019
+
†Fukuipteryx prima Imai et al. 2019
Or. †Jeholornithiformes Zhou and Zhang 2006
+
G. †Kompsornis Wang et al. 2020
+
†Kompsornis longicaudus Wang et al. 2020
G. †Neimengornis Wang et al. 2021
+
†Neimengornis rectusmim Wang et al. 2021
Invalid names: Jeholornithidae Zhou and Zhang 2006 [empty]
Unr. Ornithothoraces Chiappe and Calvo 1994
+
G. †Alamitornis Agnolin and Martinelli 2009
+
†Alamitornis minutus Agnolin and Martinelli 2009
Subcl. †Enantiornithes Walker 1981
+
Or. †Alexornithiformes Brodkorb 1976
G. †Avimaia Bailleul et al. 2019
Fm. †Avisauridae Brett-Surman and Paul 1985
Fm. †Bohaiornithidae Wang et al. 2014
Or. †Boluochiformes Zhou and Zhang 2006
G. †Brevirostruavis Li et al. 2021
G. †Castignovolucris Buffetaut et al. 2023
G. †Cathayornis Zhou et al. 1992
Or. †Cathayornithiformes Zhou et al. 1992
G. †Concornis Sanz and Buscalioni 1992
G. †Cruralispennia Wang et al. 2017
G. †Dunhuangia Wang et al. 2015
G. †Elektorornis Xing et al. 2019
G. †Elsornis Chiappe et al. 2007 [sand bird]
G. †Eoalulavis Sanz et al. 1996
G. †Eocathayornis Zhou 2002
G. †Eoenantiornis Hou et al. 1999
Or. †Eoenantiornithiformes Hou et al. 1999
Superor. †Euenantiornithes Chiappe 2002
Or. †Euornithiformes Kurochkin 1996
G. †Evgenavis O'Connor et al. 2014
G. †Falcatakely O'Connor et al. 2020
G. †Feitianius O'Connor et al. 2016
G. †Flexomornis Tykoski and Fiorillo 2010
G. †Fortipesavis Clark and O’Connor 2021
G. †Fortunguavis Wang et al. 2014
G. †Gobipipus Kurochkin et al. 2013
G. †Gobipteryx Elzanowski 1974
G. †Gretcheniao Chiappe et al. 2019
G. †Holbotia Zelenkov and Averianov 2015
G. †Houornis Wang and Liu 2015
G. †Huoshanornis Wang et al. 2010
G. †Iberomesornis Sanz et al. 1990
G. †Junornis Liu et al. 2017
G. †Lectavis Chiappe 1993
G. †Linyiornis Wang et al. 2016
Fm. †Longipterygidae Zhang et al. 2000
Or. †Longirostravisiformes Zhou and Zhang 2006
G. †Microenantiornis Wei and Li 2017
G. †Mirusavis Wang et al. 2020
G. †Monoenantiornis Hu and O'Connor 2017
G. †Musivavis Wang et al. 2022
G. †Navaornis Chiappe et al. 2024
G. †Noguerornis Lacasa Ruiz 1989
G. †Orienantius Liu et al. 2019
G. †Otogornis Hou 1994
G. †Paraprotopteryx Zheng et al. 2007
G. †Parvavis Wang et al. 2014
Fm. †Pengornithidae Wang et al. 2014
G. †Piscivorenantiornis Wang and Zhou 2017
Or. †Protopterygiformes Zhou and Zhang 2006
G. †Protopteryx Zhang and Zhou 2000
G. †Pterygornis Wang et al. 2014
G. †Qiliania Ji et al. 2011
G. †Shangyang Wang and Zhou 2019
G. †Sinornis Sereno and Rao 1992
G. †Vescornis Zhang et al. 2004
G. †Xiangornis Hu et al. 2012
G. †Yatenavis Álvarez Herrera et al. 2023
G. †Yuanjiawaornis Hu et al. 2015
G. †Yungavolucris Chiappe 1993
G. †Yuornis Xu et al. 2021
Invalid names: Aberratiodontuidae Gong et al. 2004 [empty], Cathayornis aberransis Hou 2002 [nomen dubium], Cathayornis chabuensis Li et al. 2008 [nomen dubium], Enantiornithiformes [empty], Longipterygiformes Zhang et al. 2000 [empty]
Unr. Euornithes Stejneger 1885
+
Or. Cecomorphae Huxley 1867
+
Invalid names: Pygopodes [synonym]
Invalid names: Chenomorphae Huxley 1867 [empty], Grallae Linnaeus 1758 [empty], Herodii [empty], Picariae Blanchard 1859 [empty]
Unr. †Ornithuromorpha Chiappe 1999
+
G. †Ambiortus Kurochkin 1982
G. †Antarcticavis Cordes-Person et al. 2020
G. †Apsaravis Norell and Clarke 2001
G. †Archaeorhynchus Zhou and Zhang 2006
G. †Bellulia Wang et al. 2016
G. †Changmaornis Wang et al. 2013
G. †Chaoyangia Hou and Zhang 1993
G. †Dingavis O'Connor et al. 2016
G. †Eogranivora Zheng et al. 2018
G. †Gansus Hou and Liu 1984
G. †Hollanda Bell et al. 2010
Fm. †Hongshanornithidae O'Connor et al. 2010
G. †Iteravis Zhou et al. 2014
G. †Jianchangornis Zhou et al. 2009
G. †Jiuquanornis Wang et al. 2013
G. †Juehuaornis Wang et al. 2015
G. †Kaririavis Carvalho et al. 2021
G. †Khinganornis Wang et al. 2020
G. †Liaoningornis Hou 1997
Subcl. Ornithurae Haeckel 1866
G. †Patagopteryx Alvarenga and Bonaparte 1992
G. †Piscivoravis Zhou et al. 2013
G. †Schizooura Zhou et al. 2012
Fm. †Schizoouridae Wang et al. 2020
G. †Vorona Forster et al. 1996
G. †Yumenornis Wang et al. 2013
G. †Zhongjianornis Zhou et al. 2010
Invalid names: Songlingornithidae Hou 1997 [empty]
G. †Xinghaiornis Wang et al. 2013
+
†Xinghaiornis lini Wang et al. 2013
G. †Wellnhoferia Elzanowski 2001
Invalid names: Archaeopterygidae Huxley 1871 [invalid subgroup], Llerdopteryx Lacasa 1985 [nomen dubium], Llerdopteryx viai Lacasa 1985 [nomen dubium], Wyleyia Harrison and Walker 1973 [nomen dubium], Wyleyia valdensis Harrison and Walker 1973 [nomen dubium]
G. †Migmanychion Wang et al. 2023
+
†Migmanychion laiyang Wang et al. 2023
Fm. †Montanoolithidae Zelenitsky and Therrien 2008
+
G. †Montanoolithus Zelenitsky and Therrien 2008
+
†Montanoolithus labadousensis Vila et al. 2017
†Montanoolithus strongorum Zelenitsky and Therrien 2008
Infraor. †Oviraptorosauria Barsbold 1976
+
G. †Avimimus Kurzanov 1981
+
†Avimimus nemegtensis Funston et al. 2018
†Avimimus portentosus Kurzanov 1981
Fm. †Caenagnathidae Sternberg 1940
+
G. †Anomalipes Yu et al. 2018
+
†Anomalipes zhaoi Yu et al. 2018
G. †Beibeilong Pu et al. 2017
+
†Beibeilong sinensis Pu et al. 2017
Subfm. †Caenagnathinae Paul 1988
+
G. †Anzu Lamanna et al. 2014 [chicken from hell]
G. †Apatoraptor Funston and Currie 2016
G. †Caenagnathasia Currie et al. 1994
G. †Caenagnathus Sternberg 1940
G. †Citipes Funston 2020
G. †Elmisaurus Osmolska 1981
+
Invalid names: Nomingia Barsbold et al. 2000 [synonym]
G. †Epichirostenotes Sullivan et al. 2011
G. †Hagryphus Zanno and Sampson 2005
G. †Chirostenotes Gilmore 1924
+
†Chirostenotes pergracilis Gilmore 1924
+
Invalid names: Caenagnathus sternbergi Cracraft 1971 [synonym], Macrophalangia canadensis Sternberg 1932 [synonym]
Invalid names: Macrophalangia Sternberg 1932 [synonym]
G. †Eoneophron Atkins-Weltman et al. 2024
+
†Eoneophron infernalis Atkins-Weltman et al. 2024
G. †Gigantoraptor Xu et al. 2007
+
†Gigantoraptor erlianensis Xu et al. 2007
G. †Leptorhynchos Longrich et al. 2013
+
†Leptorhynchos elegans Parks 1933
†Leptorhynchos gaddisi Longrich et al. 2013
G. †Microvenator Ostrom 1970
+
†Microvenator celer Ostrom 1970
†Microvenator chagyabi Zhao 1986
Invalid names: Elmisauridae Osmolska 1981 [synonym], Elmisaurinae Currie 2000 [empty], Ojoraptorsaurus Sullivan et al. 2011 [nomen dubium], Ojoraptorsaurus boerei Sullivan et al. 2011 [nomen dubium]
Fm. †Caudipterygidae Osmólska et al. 2004
+
G. †Similicaudipteryx He et al. 2008
+
Invalid names: Caudipteridae Zhou and Wang 2000 [replaced]
G. †Caudipteryx Ji et al. 1998
+
†Caudipteryx dongi Zhou and Wang 2000
†Caudipteryx zoui Ji et al. 1998
G. †Incisivosaurus Xu et al. 2002
+
†Incisivosaurus gauthieri Xu et al. 2002
G. †Ningyuansaurus Ji et al. 2012
+
†Ningyuansaurus wangi Ji et al. 2012
Fm. †Oviraptoridae Barsbold 1976
+
Subfm. †Citipatinae Funston 2020
+
G. †Citipati Clark et al. 2001
G. †Corythoraptor Lü et al. 2017
G. †Huanansaurus Lü et al. 2015
G. †Rinchenia Barsbold 1997
G. †Tongtianlong Lü et al. 2016
G. †Wulatelong Xu et al. 2013
G. †Gobiraptor Lee et al. 2019
+
†Gobiraptor minutus Lee et al. 2019
Subfm. †Heyuanninae Yun 2019
+
G. †Banji Xu and Han 2010
G. †Conchoraptor Barsbold 1986
G. †Heyuannia Lü 2003
+
Invalid names: Ajancingenia Easter 2013 [synonym]
G. †Jiangxisaurus Wei et al. 2013
G. †Khaan Clark et al. 2001
G. †Machairasaurus Longrich et al. 2010
G. †Nemegtomaia Lü et al. 2005
+
Invalid names: Nemegtia Lü et al. 2004 [replaced]
G. †Oksoko Funston et al. 2020
G. †Shixinggia Lü and Zhang 2005
Invalid names: Ingeniinae Barsbold 1981 [replaced]
G. †Luoyanggia Lü et al. 2009
+
†Luoyanggia liudianensis Lü et al. 2009
G. †Nankangia Lü et al. 2013
+
†Nankangia jiangxiensis Lü et al. 2013
G. †Oviraptor Osborn 1924
+
†Oviraptor philoceratops Osborn 1924
G. †Yulong Lü et al. 2013
+
†Yulong mini Lü et al. 2013
Invalid names: Oviraptorinae Barsbold 1981 [empty]
G. †Protarchaeopteryx Ji and Ji 1997
+
†Protarchaeopteryx robusta Ji and Ji 1997
G. †Saurexallopus Harris 1997
+
†Saurexallopus cordata McCrea et al. 2014
†Saurexallopus lovei Harris et al. 1996
†Saurexallopus zerbsti Lockley et al. 2003
Invalid names: Exallopus Harris et al. 1996 [replaced]
G. †Yuanyanglong Hao et al. 2025
+
†Yuanyanglong bainian Hao et al. 2025
Invalid names: Avimimidae Kurzanov 1981 [empty], Caenagnathoidea Sternberg 1940 [empty], Ingeniidae Barsbold 1981 [empty], Oviraptoroidea Barsbold 1976 [empty]
Unr. Paraves Sereno 1997
+
Cl. Aves Linnaeus 1758 [bird]
+
Or. Accipitriformes Vieillot 1816
+
Fm. Accipitridae Viellot 1816 [hawk]
Fm. Pandionidae Sclater and Salvin 1873 [osprey]
G. †Alaripeda Sarjeant and Reynolds 2001
+
†Alaripeda logfreni Sarjeant and Reynolds 2001
G. †Archaeornithipus Fuentes Vidarte 1996
+
†Archaeornithipus meijidei Fuentes Vidarte 1996
G. †Aucornis Ameghino 1898
+
Invalid names: Aucornis solidus Ameghino 1899 [nomen dubium]
Fm. †Avipedidae Sarjeant and Langston 1994
+
G. †Aquatilavipes Currie 1981
G. †Avipeda Vialov 1965
G. †Fuscinapeda Sarjeant and Langston 1994
G. †Barrosopus Coria et al. 2002
+
†Barrosopus slobodai Coria et al. 2002
Or. Bucerotiformes Fürbringer 1888 [hornbill]
+
Fm. Bucerotidae Rafinesque 1815 [hornbill]
Fm. Upupidae Bonaparte 1831
Or. Charadriiformes Huxley 1867 [shorebird]
+
Superfm. Alcoidae Stejneger 1885
Fm. Burhinidae Mathews 1912 [stone-curlew]
Subor. Charadrii Huxley 1867
G. †Charadriisimilis Mayr and Kitchener 2023
Fm. Chionidae Bonaparte 1832 [sheathbill]
Fm. †Cimolopterygidae Brodkorb 1963
Fm. †Dakotornithidae Erickson 1975
G. Elseyornis Mathews 1914
G. Erythrogonys Gould 1838
Fm. †Graculavidae Fürbringer 1888
G. †Graculavus Marsh 1872
Fm. Haematopodidae Bonaparte 1838 [oystercatchers]
G. Hydrophasianus Wagler 1832
Fm. Jacanidae Stejneger 1885
Subor. Lari Sharpe 1891
Subor. Limicolae
Unr. Panalcidae Smith 2011
Subor. Parrae
Fm. Pedionomidae Gadow 1893
G. Peltohyas Sharpe 1896
G. Phegornis Gray 1845
G. Pluvianellus Gray 1846
Fm. Pluvianidae MacGillivray 1852 [Egyptian plover]
Fm. Recurvirostridae Bonaparte 1854
G. Rhinoptilus Strickland 1852
G. †Sarjeantopodus Lockley et al. 2003
Subor. Scolopaci Strauch 1978
Fm. Scolopacidae Rafinesque 1815 [snipe]
+
Invalid names: Phalaropodidae Bonaparte 1831 [synonym]
Fm. Stercorariidae Brisson 1760 [skua]
G. †Telmatornis Marsh 1870
Fm. Thinocoridae Gray 1845
G. Thinornis Vieillot 1789
Fm. Turnicidae Gray 1840 [buttonquail]
+
Invalid names: Turnipacidae Mayr 2000 [synonym]
G. †Volgavis Nessov and Yarkov 1989
Invalid names: Alcae [empty], Pedionomae [empty], Telmatornithidae Cracraft 1972 [empty]
G. †Dolichopterus Milne-Edwards 1867
+
†Dolichopterus viator Milne-Edwards 1867
Fm. †Dromiceiidae Wetmore
+
G. Dromaeus Vieillot 1816
G. †Dromiceius Vieillot
Unr. †Eoaves Sibley 1988
+
Invalid names: Lithornithiformes Houde 1988 [empty]
G. †Eociconia Hou 1989
+
G. †Eopteryx Meyer 1887
+
Unr. †Eucavitaves Kimball 2013
+
Unr. Bucerotes Fürbringer 1888
Superor. Eurhipidurae Gill 1874
+
Or. †Hesperornithomorphi Hay 1930
+
Invalid names: Odontolcae Marsh 1875 [synonym]
G. †Euryonotus Mercerat 1897
+
†Euryonotus argentinus Mercerat 1897
†Euryonotus brachypterus Mercerat 1897
Unr. Falconiformes Sharpe 1874 [raptor]
+
Subor. Accipitres Linnaeus 1758
Subor. Cathartae
+
Invalid names: Sarcorhamphi [synonym]
Fm. Falconidae Vigors 1824
Superfm. Falconoidae
Fm. †Masillaraptoridae Mayr 2009
Superfm. Vulturoidae Hay 1930
Invalid names: Cathartoidea Lafresnaye 1839 [empty], Falcones Lambrecht 1933 [empty], Horusornithidae [empty], Teratornithi [empty]
G. †Fontinalis Lesquereux 1883
+
Fontinalis pristina Lesuquereux 1883
Unr. †Gansuiformes Hou and Liu 1984
+
Invalid names: Gansuidae Hou and Liu 1984 [empty]
Fm. †Gobioolithidae Mikhailov 1996
+
G. †Gobioolithus Mikhailov 1996
G. †Gruipeda Panin and Avram 1962
+
†Gruipeda dominguensis de Valais and Melchor 2008
†Gruipeda maxima Panin and Avram 1962
†Gruipeda vegrandiunus Fiorillo et al. 2011
Invalid names: Antarctichnus Covacevich and Lamperein 1970 [synonym], Charadriipeda Panin and Avram 1962 [synonym]
G. †Gyeongsangornipes Kim et al. 2013
+
†Gyeongsangornipes lockleyi Kim et al. 2013
G. †Ibidopodia Milne-Edwards 1868
+
†Ibidopodia palustris Milne-Edwards 1868
Fm. †Ignotornidae Lockley et al. 1992
+
G. †Goseongornipes Lockley et al. 2006
G. †Hwangsanipes Yang et al. 1995
G. †Ignotornis Mehl 1931
G. †Jeholornis Zhou and Zhang 2002
+
†Jeholornis curvipes Lefevre et al. 2014
†Jeholornis palmapenis O'Connor et al. 2012
†Jeholornis prima Zhou and Zhang 2002
Fm. †Jindongornipodidae Lockley et al. 2006
+
G. †Jindongornipes Lockley et al. 1992
G. †Jixiangornis Ji et al. 2002
+
†Jixiangornis orientalis Ji et al. 2002
Fm. †Koreanaornipodidae Lockley et al. 2006
+
G. †Koreanaornis Kim 1969
G. †Koreanornis Kim 1969
G. †Pullornipes Lockley et al. 2006
Fm. †Limiavipedidae McCrea et al. 2014
+
G. †Limiavipes McCrea et al. 2014
G. †Wupus Xing et al. 2007
G. †Loncornis Ameghino 1899
G. †Manu Marples 1946
+
†Manu antiquus Marples 1946
+
Subor. †Micropodii
Invalid names: Trochili [empty]
G. †Moguiornipes Xing et al. 2011
+
†Moguiornipes robusta Xing et al. 2011
Or. †Mystiornithiformes Kurochkin et al. 2011
+
Fm. †Mystiornithidae Kurochkin et al. 2011
Subcl. Neornithes Gadow 1893
+
Unr. Aequornithes Mayr 2011
Or. Alectromorphae Gadow
Or. Cariamiformes Fürbringer 1888
Or. Cathartiformes Coues 1884
Or. Colymbomorphae Gadow
Or. Eurypygiformes Hackett et al. 2008
Or. Galbuliformes Vigors 1825
Or. Leptosomiformes Sharpe 1891
Or. Mesitornithiformes Wetmore 1960
Unr. Neognathae Pycraft 1900
Or. Otidiformes Wagler 1830
Superor. Palaeognathae Pycraft 1900
Or. Phoenicopteriformes Howard 1955 [flamingo]
Unr. Picocoraciae Mayr 2011
Or. Pterocliformes Huxley 1868
Invalid names: Coraciomorphae Gadow [empty], Lonchodytes pterygius Brodkorb 1963 [nomen dubium], Odontognathae Wetmore 1930 [empty], Pelargomorphae Gadow [empty]
Or. †Odontopterygiformes Howard 1957
+
Fm. †Pelagornithidae Fürbringer 1888
+
Invalid names: Dasornithidae Harrison and Walker 1976 [synonym], Odontopterygidae Lydekker 1891 [synonym], Pseudodontornithidae Lambrecht 1933 [synonym]
Or. †Omnivoropterygiformes Czerkas and Ji 2002
+
Fm. †Omnivoropterygidae Czerkas and Ji 2002
G. †Onychopteryx Cracraft 1971
+
†Onychopteryx simpsoni Cracraft 1971
Fm. †Pachycorioolithidae Lawver et al. 2016
+
G. †Pachycorioolithus Lawver et al. 2016
Unr. †Palaeoaves Livezey and Zusi 2007
+
Superor. †Archaeornithes Gadow 1893
G. †Palaeolestes DeVis 1911
G. †Palaeopsittacus Harisson 1982
+
†Palaeopsittacus georgei Harrison 1982
Unr. †Pangalliformes Clarke 2004
+
G. †Austinornis Clarke 2004
G. †Parvoolithus Mikhailov 1996
+
†Parvoolithus tortuosus Mikhailov 1996
Fm. †Paxavipedidae McCrea et al. 2015
+
G. †Paxavipes McCrea et al. 2015
Or. Phaethontiformes Sharpe 1891
+
G. †Clymenoptilon Mayr et al. 2023
Fm. Phaethontidae Brandt 1831 [tropicbird]
G. †Phaethusavis Bourdon et al. 2008
Fm. †Prophaethontidae Harrison and Walker 1976
G. †Platanavis Nessov 1992
+
†Platanavis nana Nessov 1992
G. Plectophora Gray 1834
Subcl. †Praeornithes Rautian 1978
+
Or. †Praeornithiformes Rautian 1978
G. †Proornis Lim 1993
+
†Proornis coreae Lim 1993
Or. †Protoaviformes Chatterjee 1991
+
Invalid names: Protoavidae Chatterjee 1991 [empty]
G. †Protornis Jaekel 1929
Unr. †Psittacopasseres Suh 2011
+
G. †Namapsitta Mourer-Chauvire et al. 2015
Unr. †Parapasseres Mayr 2015
Fm. †Psittacopedidae Mayr 2015
+
G. †Eofringillirostrum Ksepka et al. 2019
G. †Parapsittacopes Mayr 2020
G. †Psittacomimus Mayr and Kitchener 2022
G. †Psittacopes Mayr and Daniels 1998
G. †Pumiliornis Mayr 1999
Unr. Pygostylia Chiappe 2002
+
Or. †Confuciusornithiformes Hou et al. 1995
Fm. †Jinguofortisidae Wang et al. 2018
G. †Sapeornis Zhou and Zhang 2002
+
Invalid names: Shenshiornis Hu et al. 2010 [synonym]
Or. †Sapeornithiformes Zhou and Zhang 2006
Fm. †Qianshanornithidae Mayr et al. 2013
+
G. †Qianshanornis Mayr et al. 2013
G. †Querandiornis Rusconi 1958
+
†Querandiornis romani Rusconi 1958
G. †Riacama Ameghino 1899
+
†Riacama caliginea Ameghino 1899
Fm. †Shandongornipodidae Lockley et al. 2007
+
G. †Shandongornipes Li et al. 2005 [Shandong bird track]
G. †Shenzhouraptor Ji et al. 2002
+
†Shenzhouraptor sinensis Ji et al. 2002
G. †Smiliornis Ameghino 1899
+
†Smiliornis penetrans Ameghino 1899
G. †Tantalatos Reichenbach 1852
G. †Tatarornipes Lockley et al. 2012
+
†Tatarornipes chabuensis Lockley et al. 2012
Unr. †Telluraves Yuri 2013
+
Unr. †Afroaves Ericson 2012
Unr. †Australaves Ericson 2012
G. †Calcardea Gingerich 1987
+
Subor. †Stereornithes Moreno and Mercerat 1891
Invalid names: Larolimicolae Lambrecht 1933 [empty]
G. †Teracus Milne-Edwards 1871
+
†Teracus littoralis Milne-Edwards 1871
G. †Tridentigerpes Xing et al. 2019
+
†Tridentigerpes huashibanleei Xing et al. 2019
†Tridentigerpes pinuelai Xing et al. 2019
G. †Tshulia Nessov 1988
+
†Tshulia litorea Nessov 1988
G. †Uhangrichnus Yang et al. 1995
+
†Uhangrichnus chuni Yang et al. 1995
+
Invalid names: Dongyangornipes sinensis Azuma et al. 2013 [synonym]
Invalid names: Dongyangornipes Azuma et al. 2013 [synonym]
G. †Yacoraitichnus Alonso and Marquillas 1986
+
†Yacoraitichnus avis Alonso and Marquillas 1986
G. †Yalavis Shufeldt 1913
+
†Yalavis tenuipes Shufeldt 1913
Invalid names: Archornithes Carus [empty], Avimimiformes Chatterjee 1991 [empty], Chaoyangornithidae [empty], Climacarthrus Ameghino 1895 [nomen dubium], Eupterornis remensis Lemoine 1878 [nomen dubium], Kuszholiidae Nessov 1992 [empty], Liaoxiornithidae [empty], Longipennes [empty], Metornithes Perle et al. 1993 [empty], Minerva Shufeldt 1915 [nomen dubium], Mirandornithes Sangster 2005 [empty], Musophagiformes Seebohm 1890 [empty], Odontoglossae [empty], Odontornithes Marsh 1873 [empty], Paludicolae [empty], Pancharadriiformes Hood et al. 2019 [empty], Panmirandornithes Hood et al. 2019 [empty], Pansphenisciformes Clarke et al. 2003 [empty], Patagonichnornis Casamiquela 1987 [nomen vanum], Proherodius Lydekker 1891 [nomen dubium], Proherodius oweni Lydekker 1891 [nomen dubium], Protopelargus Reichenbach 1852 [nomen nudum], Sauriurae Haeckel 1866 [invalid subgroup], Tantaleus Reichenbach 1852 [nomen nudum], Tricorynopus brinkmani McCrea and Sarjeant 2001 [nomen dubium]
G. †Caihong Hu et al. 2018
+
†Caihong juji Hu et al. 2018
Infraor. †Deinonychosauria Colbert and Russell 1969
+
Fm. †Dromaeopodidae Li et al. 2007
+
G. †Dromaeopodus Li et al. 2007 [runner foot]
G. †Dromaeosauripus Kim et al. 2008
G. †Menglongipus Xing et al. 2009
Fm. †Dromaeosauridae Colbert and Russell 1969
+
G. †Balaur Csiki et al. 2010
G. †Bambiraptor Burnham et al. 2000
G. †Daurlong Wang et al. 2022
G. †Dromaeosauroides Christiansen and Bonde 2003
Unr. †Eudromaeosauria Longrich and Currie 2009
Subfm. †Halszkaraptorinae Cau et al. 2017
G. †Imperobator Ely and Case 2019
G. †Kuru Napoli et al. 2021
G. †Luanchuanraptor Lü 2007
Subfm. †Microraptorinae Senter et al. 2004
G. †Nuthetes Owen 1854
G. †Pyroraptor Allain and Taquet 2000
G. †Richardoestesia Currie et al. 1990
+
Invalid names: Asiamericana Nessov 1995 [synonym]
G. †Shri Turner et al. 2021
G. †Tianyuraptor Zheng et al. 2009
Subfm. †Unenlagiinae Bonaparte 1999
G. †Variraptor Le Loeuff and Buffetaut 1998
G. †Velociraptorichnus Zhen et al. 1994
G. †Wulong Poust et al. 2020
G. †Zhenyuanlong Lü and Brusatte 2015
G. †Pamparaptor Porfiri et al. 2011
+
†Pamparaptor micros Porfiri et al. 2011
G. †Paravipus Mudroch et al. 2011
+
†Paravipus didactyloides Mudroch et al. 2011
Fm. †Troodontidae Gilmore 1924
+
G. †Albertavenator Evans et al. 2017
G. †Almas Pei et al. 2017
G. †Anchiornis Xu et al. 2009
G. †Archaeornithoides Elzanowski and Wellnhofer 1992
G. †Daliansaurus Shen et al. 2017
G. †Geminiraptor Senter et al. 2010
G. †Gobivenator Tsuihiji et al. 2014
G. †Harenadraco Lee et al. 2024
G. †Hesperornithoides Hartman et al. 2019
Subfm. †Jinfengopteryginae Turner et al. 2012
G. †Papiliovenator Pei et al. 2022
G. †Pectinodon Carpenter 1982
Subfm. †Sinovenatorinae Shen et al. 2017
G. †Sinusonasus Xu and Wang 2004
G. †Tochisaurus Kurzanov and Osmólska 1991
Subfm. †Troodontinae van der Reest and Currie 2017
G. †Xixiasaurus Lü et al. 2010
Invalid names: Bradycneme Harrison and Walker 1975 [nomen dubium], Heptasteornis Harrison and Walker 1975 [nomen dubium], Koparion Chure 1994 [nomen dubium], Laelaps cristatus Cope 1876 [nomen dubium], Laelaps laevifrons Cope 1876 [nomen dubium], Polyodontosaurus grandis Gilmore 1932 [nomen dubium], Saurornithoididae Barsbold 1974 [synonym], Saurornitholestes robustus Sullivan 2006 [nomen dubium]
G. †Eosinopteryx Godefroit et al. 2013
+
†Eosinopteryx brevipenna Godefroit et al. 2013
G. †Epidendrosaurus Zhang et al. 2002
+
†Epidendrosaurus ningchengensis Zhang et al. 2002
G. †Epidexipteryx Zhang et al. 2008
+
†Epidexipteryx hui Zhang et al. 2008
Unr. Eumaniraptora Padian et al. 1999
+
Invalid names: Dromaeosauria Chatterjee and Templin 2007 [empty]
G. †Overoraptor Motta et al. 2020
+
†Overoraptor chimentoi Motta et al. 2020
G. †Paronychodon Cope 1876
+
†Paronychodon asiaticus Nessov 1995
Invalid names: Euronychodon Antunes and Sigogneau-Russell 1991 [synonym], Paronychodon lacustris Cope 1876 [nomen dubium], Tripriodon Marsh 1889 [synonym]
G. †Pedopenna Xu and Zhang 2005
+
†Pedopenna daohugouensis Xu and Zhang 2005
G. †Pneumatoraptor Osi et al. 2010
+
†Pneumatoraptor fodori Osi et al. 2010
G. †Serikornis Lefèvre et al. 2017
+
†Serikornis sungei Lefèvre et al. 2017
G. †Zhongornis Gao et al. 2008
+
†Zhongornis haoae Gao et al. 2008
G. †Pseudogeckoolithus Vianey-Liaud and López-Martínez 1997
+
†Pseudogeckoolithus nodosus Vianey-Liaud and López-Martínez 1997
†Pseudogeckoolithus tirboulensis Vianey-Liaud and Garcia 2003
Fm. †Scansoriopterygidae Czerkas and Yuan 2002
+
G. †Ambopteryx Wang et al. 2019
+
†Ambopteryx longibrachium Wang et al. 2019
G. †Scansoriopteryx Czerkas and Yuan 2002
+
†Scansoriopteryx heilmanni Czerkas and Yuan 2002
G. †Yi Xu et al. 2015
+
†Yi qi Xu et al. 2015
G. †Yixianosaurus Xu and Wang 2003
+
†Yixianosaurus longimanus Xu and Wang 2003
Invalid names: Palaeopteryx Jensen 1981 [nomen dubium]
Unr. Maniraptoriformes Holtz, Jr. and Padian 1995
+
Unr. †Arctometatarsalia Holtz 1994
+
Infraor. †Ornithomimosauria Barsbold 1976 [ornithomimosaur]
+
G. †Afromimus Sereno 2017
+
†Afromimus tenerensis Sereno 2017
G. †Arkansaurus Hunt and Quinn 2018
+
†Arkansaurus fridayi Hunt and Quinn 2018
G. †Aviatyrannis Rauhut 2003
+
†Aviatyrannis jurassica Rauhut 2003
G. †Beishanlong Makovicky et al. 2010
+
†Beishanlong grandis Makovicky et al. 2010
Fm. †Deinocheiridae Kielan-Jaworowska 1969
+
G. †Deinocheirus Kielan-Jaworowska 1969
G. †Garudimimus Barsbold 1981 [Garudi mimic]
G. †Harpymimus Barsbold and Perle 1984
G. †Paraxenisaurus Serrano-Brañas et al. 2020
G. †Tyrannomimus Hattori et al. 2023
G. †Hexing Jin et al. 2012
+
†Hexing qingyi Jin et al. 2012
G. †Kinnareemimus Buffetaut et al. 2009
+
†Kinnareemimus khonkaenensis Buffetaut et al. 2009
Fm. †Nqwebasauridae Alifanov and Saveliev 2015
+
G. †Lepidocheirosaurus Alifanov and Saveliev 2015
G. †Nqwebasaurus de Klerk et al. 2000
+
†Nqwebasaurus thwazi de Klerk et al. 2000
Fm. †Ornithomimidae Marsh 1890 [ornithomimid]
+
G. †Aepyornithomimus Chinzorig et al. 2017
G. †Anserimimus Barsbold 1988
G. †Archaeornithomimus Russell 1972
G. †Dromiceiomimus Russell 1972
G. †Gallimimus Osmólska et al. 1972
G. †Ornithomimidorum Huene 1926
G. †Ornithomimus Marsh 1890
G. †Qiupalong Xu et al. 2011
G. †Rativates McFeeters et al. 2016
G. †Sinornithomimus Kobayashi and Lü 2003
G. †Struthiomimus Osborn 1916
G. †Tototlmimus Serrano-Brañas et al. 2016
Invalid names: Deinocheirinae Paul 1988 [empty], Garudimiminae Paul 1988 [empty], Ornithomiminae Nopcsa 1923 [empty]
G. †Pelecanimimus Pérez-Moreno et al. 1994
+
†Pelecanimimus polyodon Pérez-Moreno et al. 1994
G. †Shenzhousaurus Ji et al. 2003
+
†Shenzhousaurus orientalis Ji et al. 2003
Invalid names: Coelosaurus Leidy 1865 [nomen dubium], Garudimimidae Barsbold 1981 [empty], Harpymimidae Barsbold and Perle 1984 [empty], Ornithomimoidea Sereno 1999 [empty], Pelecanimimidae Alifanov 2012 [empty], Thecocoelurus Huene 1923 [nomen dubium], Valdoraptor Olshevsky 1991 [nomen dubium]
Invalid names: Bullatosauria Holtz [empty]
Unr. †Therizinosauria Russell 1997
+
G. †Erlikosaurus Barsbold and Perle 1980
+
†Erlikosaurus andrewsi Barsbold and Perle 1980
G. †Falcarius Kirkland et al. 2005
+
†Falcarius utahensis Kirkland et al. 2005
G. †Fukuivenator Azuma et al. 2016
+
†Fukuivenator paradoxus Azuma et al. 2016
G. †Jianchangosaurus Pu et al. 2013
+
†Jianchangosaurus yixianensis Pu et al. 2013
G. †Lingyuanosaurus Yao et al. 2019
+
†Lingyuanosaurus sihedangensis Yao et al. 2019
Superfm. †Therizinosauroidea Russell and Dong 1994
+
G. †Alxasaurus Russell and Dong 1994
+
†Alxasaurus elesitaiensis Russell and Dong 1994
G. †Beipiaosaurus Xu et al. 1999
+
†Beipiaosaurus inexpectus Xu et al. 1999
Fm. †Dendroolithidae Zhao and Li 1988
+
G. †Placoolithus Zhao 1979
G. †Martharaptor Senter et al. 2012
+
†Martharaptor greenriverensis Senter et al. 2012
Fm. †Therizinosauridae Maleev 1954
+
G. †Enigmosaurus Barsbold and Perle 1983 [enigmatic lizard]
G. †Erliansaurus Xu et al. 2002
G. †Eshanosaurus Xu et al. 2001
G. †Nanshiungosaurus Dong 1979
G. †Neimongosaurus Zhang et al. 2001
G. †Nothronychus Kirkland and Wolfe 2001
G. †Paralitherizinosaurus Kobayashi et al. 2022
G. †Segnosaurus Perle 1979
G. †Suzhousaurus Li et al. 2007 [Suzhou lizard]
G. †Therizinosaurus Maleev 1954
G. †Tiantaisaurus Qian et al. 2012
Invalid names: Enigmosauridae Barsbold and Perle 1983 [synonym], Erlikosauridae Eberth et al. 2001 [synonym], Nanshiungosauridae Dong and Yu 1997 [synonym], Segnosauridae Perle 1979 [synonym]
Invalid names: Alxasauridae Russell and Dong 1994 [empty], Segnosauria Barsbold and Perle 1980 [synonym], Segnosaurischia Dong 1992 [synonym]
Invalid names: Tyrannoraptora Sereno 1998 [empty]
Unr. †Megaraptora Benson et al. 2010
+
G. †Aoniraptor Motta et al. 2016
+
†Aoniraptor libertatem Motta et al. 2016
Fm. †Megaraptoridae Novas et al. 2013
+
G. †Aerosteon Sereno et al. 2008
+
†Aerosteon riocoloradensis Sereno et al. 2008
G. †Australovenator Hocknull et al. 2009
+
†Australovenator wintonensis Hocknull et al. 2009
G. †Maip Aranciaga Rolando et al. 2022
+
†Maip macrothorax Aranciaga Rolando et al. 2022
G. †Megaraptor Novas 1998
+
G. †Murusraptor Coria and Currie 2016
+
†Murusraptor barrosaensis Coria and Currie 2016
G. †Orkoraptor Novas et al. 2008
+
†Orkoraptor burkei Novas et al. 2008
G. †Tratayenia Porfiri et al. 2018
+
†Tratayenia rosalesi Porfiri et al. 2018
G. †Phuwiangvenator Samathi et al. 2019
+
†Phuwiangvenator yaemniyomi Samathi et al. 2019
G. †Siats Zanno and Makovicky 2013
+
†Siats meekerorum Zanno and Makovicky 2013
G. †Vayuraptor Samathi et al. 2019
+
†Vayuraptor nongbualamphuensis Samathi et al. 2019
G. †Nedcolbertia Kirkland et al. 1998
+
†Nedcolbertia justinhoffmanni Kirkland et al. 1998
G. †Neotripodiscus Ellenberger 1970
+
†Neotripodiscus makoetlani Ellenberger 1970
G. †Ngexisaurus Zhao 1983
+
Fm. †Ornithomimipodidae Lockley et al. 2011
+
G. †Columbosauripus Sternberg 1932
+
†Columbosauripus ungulatus Sternberg 1932
G. †Irenichnites Sternberg 1932
G. †Magnoavipes Lee 1997
+
†Magnoavipes asiaticus Matsukawa et al. 2014
†Magnoavipes caneeri Lockley et al. 2001
†Magnoavipes denaliensis Fiorillo et al. 2011
†Magnoavipes lowei Lee 1997
G. †Ornithomimipus Sternberg 1926
+
†Ornithomimipus angustus Sternberg 1926
†Ornithomimipus gracilis Sternberg 1932
†Ornithomimipus jaillardi Jaillard et al. 1993
G. †Plesiothornipus Harkness 1850
+
†Plesiothornipos binneyi Harkness 1850
G. †Qomoqomosauropus Ellenberger 1970
+
†Qomoqomosauropus acutus Ellenberger 1970
G. †Satapliasaurus Gabuniya 1951
+
†Satapliasaurus dsocenidzei Gabuniya 1951
†Satapliasaurus kandelakii Gabuniya 1951
G. †Shanbeipus Wang et al. 2016
+
†Shanbeipus caudatus Wang et al. 2016
G. †Sinosauropteryx Ji and Ji 1996
+
†Sinosauropteryx prima Ji and Ji 1996
G. †Tanycolagreus Carpenter et al. 2005
+
†Tanycolagreus topwilsoni Carpenter et al. 2005
G. †Taupezia Delair 1963
+
†Taupezia landeri Delair 1963
G. †Trisauropodiscus Ellenberger 1970
+
†Trisauropodiscus aviforma Ellenberger 1970
†Trisauropodiscus galliforma Ellenberger 1970
†Trisauropodiscus levis Ellenberger 1970
†Trisauropodiscus phasianiforma Ellenberger 1970
†Trisauropodiscus pompompoi Ellenberger 1970
†Trisauropodiscus superaviforma Ellenberger 1970
†Trisauropodiscus superavipes Ellenberger 1970
Superfm. †Tyrannosauroidea Walker 1964
+
G. †Alectrosaurus Gilmore 1933
+
†Alectrosaurus olseni Gilmore 1933
G. †Appalachiosaurus Carr et al. 2005
+
G. †Calamosaurus Lydekker 1891
+
†Calamosaurus foxi Lydekker 1889
Fm. †Coeluridae Marsh 1881 [coelurid]
+
Invalid names: Coeluridae Nopcsa 1901 [empty], Coelurosauridae Cope 1882 [synonym]
G. †Dilong Xu et al. 2004
+
†Dilong paradoxus Xu et al. 2004
G. †Dryptosaurus Marsh 1877
+
Invalid names: Laelaps Cope 1866 [synonym]
G. †Eotyrannus Hutt et al. 2001
+
†Eotyrannus lengi Hutt et al. 2001
G. †Gualicho Apesteguia et al. 2016
+
†Gualicho shinyae Apesteguia et al. 2016
G. †Jinbeisaurus Wu et al. 2020
+
†Jinbeisaurus wangi Wu et al. 2020
G. †Juratyrant Brusatte and Benson 2013
+
†Juratyrant langhami Benson 2008
G. †Moros Zanno et al. 2019
+
†Moros intrepidus Zanno et al. 2019
Fm. †Proceratosauridae Rauhut et al. 2010
+
G. †Guanlong Xu et al. 2006
+
†Guanlong wucaii Xu et al. 2006
G. †Kileskus Averianov et al. 2010
+
†Kileskus aristotocus Averianov et al. 2010
G. †Proceratosaurus Huene 1926
+
†Proceratosaurus bradleyi Woodward 1910
G. †Sinotyrannus Ji et al. 2009
+
†Sinotyrannus kazuoensis Ji et al. 2009
G. †Yutyrannus Xu et al. 2012
+
†Yutyrannus huali Xu et al. 2012
G. †Raptorex Sereno et al. 2009
+
†Raptorex kriegsteini Sereno et al. 2009
G. †Santanaraptor Kellner 1999
+
†Santanaraptor placidus Kellner 1999
G. †Stokesosaurus Madsen 1974
+
†Stokesosaurus clevelandi Madsen 1974
G. †Suskityrannus Nesbitt et al. 2019
+
†Suskityrannus hazelae Nesbitt et al. 2019
G. †Teihivenator Yun 2017
+
†Teihivenator macropus Cope 1868
G. †Theroplantigrada Casanovas Cladellas et al. 1993
+
†Theroplantigrada encisensis Casanovas Cladellas et al. 1993
G. †Timimus Rich and Vickers-Rich 1994
+
†Timimus hermani Rich and Vickers-Rich 1994
G. †Timurlengia Brusatte et al. 2016
+
†Timurlengia euotica Brusatte et al. 2016
Fm. †Tyrannosauridae Osborn 1906 [tyrannosaurid]
+
Subfm. †Albertosaurinae Currie et al. 2003
+
G. †Albertosaurus Osborn 1905 [Alberta lizard]
+
+
Invalid names: Albertosaurus arctunguis Parks 1928 [synonym]
G. †Gorgosaurus Lambe 1914
+
†Gorgosaurus libratus Lambe 1914
+
Invalid names: Dryptosaurus kenabekides Hay 1899 [synonym], Gorgosaurus sternbergi Matthew and Brown 1922 [synonym], Laelaps falculus Cope 1876 [synonym], Laelaps hazenianus Hay 1902 [synonym], Laelaps incrassatus Cope 1876 [synonym]
G. †Bagaraatan Osmólska 1996
+
†Bagaraatan ostromi Osmólska 1996
Subfm. †Tyrannosaurinae Matthew and Brown 1922 [tyrannosaurine]
+
Tr. †Alioramini Lü et al. 2014
+
G. †Alioramus Kurzanov 1976
G. †Qianzhousaurus Lü et al. 2014
G. †Asiatyrannus Zheng et al. 2024
+
†Asiatyrannus xui Zheng et al. 2024
Unr. †Daspletosaurini Voris et al. 2020
+
G. †Daspletosaurus Russell 1970
G. †Thanatotheristes Voris et al. 2020
G. †Lythronax Loewen et al. 2013
+
†Lythronax argestes Loewen et al. 2013
G. †Nanuqsaurus Fiorillo and Tykoski 2014
+
†Nanuqsaurus hoglundi Fiorillo and Tykoski 2014
Unr. †Teratophoneini Scherer and Voiculescu-Holvad 2024
+
G. †Bistahieversor Carr and Williamson 2010
G. †Dynamoterror McDonald et al. 2018
G. †Labocania Molnar 1974
G. †Teratophoneus Carr et al. 2011
Tr. †Tyrannosaurini Olshevsky et al. 1995
+
G. †Tarbosaurus Maleev 1955
+
Invalid names: Jenghizkhan Olshevsky 1995 [synonym], Maleevosaurus Carpenter 1992 [synonym], Shanshanosaurus Dong 1977 [synonym]
G. †Tyrannosaurus Osborn 1905 [tyrant lizard]
+
Invalid names: Dinotyrannus Olshevsky 1995 [synonym], Dynamosaurus Osborn 1905 [synonym], Manospondylus Cope 1892 [synonym], Nanotyrannus Bakker et al. 1988 [synonym], Stygivenator Olshevsky 1995 [synonym]
G. †Zhuchengtyrannus Hone et al. 2011
Invalid names: Shanshanosaurinae Olshevsky 1995 [synonym], Tarbosaurini Olshevsky et al. 1995 [empty], Tyrannosaurus zhuchengensis Hu et al. 2001 [nomen dubium]
Unr. †Tyrannosauripodidae McCrea et al. 2014
+
G. †Bellatoripes McCrea et al. 2014
+
†Bellatoripes fredlundi McCrea et al. 2014
G. †Tyrannosauripus Lockley and Hunt 1994
+
†Tyrannosauripus pillmorei Lockley and Hunt 1994
G. †Yezosaurus Muramoto and Obata 1977
+
†Yezosaurus mikasaensis Caldwell et al. 2008
Invalid names: Aublysodon Leidy 1868 [nomen dubium], Deinodontidae Brown 1914 [synonym], Ornithomimus grandis Marsh 1890 [nomen dubium]
G. †Xiongguanlong Li et al. 2009
+
†Xiongguanlong baimoensis Li et al. 2009
Invalid names: Chingkankousaurus Young 1958 [nomen nudum], Deinodon Leidy 1856 [nomen dubium], Diplotomodon Leidy 1868 [nomen dubium], Tomodon horrificus Leidy 1865 [nomen dubium], Tyrannosaurus lanpingensis Yeh 1975 [nomen dubium]
G. †Xinjiangovenator Rauhut and Xu 2005
+
†Xinjiangovenator parvus Rauhut and Xu 2005
G. †Xiuningpus Yu 1998
+
Invalid names: Xiuningpus qukouensis Yu 1998 [nomen nudum]
G. †Zuolong Choiniere et al. 2010
+
†Zuolong salleei Choiniere et al. 2010
Invalid names: Calamospondylus Fox 1866 [nomen dubium], Megalosaurus lonzeensis Dollo 1883 [nomen dubium]
Diagnosis
ReferenceDiagnosis
R. S. Lull 1924Small to middle-sized, slenderly built, bipedal dinosaurs, with cavernous vertebrae and limb bones; hind limbs, pubis, and ischium very slender, as are three fingers of hand; metatarsus elongated, bones being closely appressed, but without fusion; teeth of carnivorous type, but entirely lacking in Cretaceous forms, Struthiomimus and also probably Ornithomimus.
F. v. Huene 1948Large orbit; pointed skull; 9 elongated cervicals, 14 dorsals, 3 or more sacrals; long tail, distal vertebrae sometimes elongated; manus not reduced on ulnar side, mtc. V long; ilium with strong anterior and posterior process, acetabular opening small; pubic in earlier forms broad, plate-like, later elongated, often rod-like; trochanter quartus of femur in a high position; tibia longer than femur; calcaneum in earlier forms with spur; mt. II-IV long, without coalescence, I and V short; bird-like foot; thin walls of vertebrae and of all bones.
J. F. Bonaparte 1978Triásico sup. Jurásico-Cretácico; de hábitos bipedales; de talla chica o mediana; carnívoros y frugívoros.