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Testudinata (turtle)

Reptilia

Taxonomy
Testudinata was named by Oppel (1811). It is extant.

It was reranked as the subclass Testudinata by Bonaparte (1850), Estes (1964), Estes and Berberian (1970), Haas (1978), Carroll (1988), Cvancara and Hoganson (1993), Joyce et al. (2004) and Martinelli and Forasiepi (2004); it was reranked as the suborder Testudinata by Lydekker (1889); it was reranked as the superorder Testudinata by Nopcsa (1928) and Roxo (1937); it was synonymized subjectively with Testudines by Kuhn (1966) and Dundee (1989); it was reranked as the unranked clade Testudinata by Fitzinger (1826), Cope (1891), Gardiner (1982) and Sterli (2008).

It was assigned to Monopnoa by Fitzinger (1826); to Chelonia by Lydekker (1889); to Synapsida by Osborn (1903) and Osborn (1903); to Anomodontia by Osborn (1904); to Paratheria by Jaekel (1911); to Astatoglyptica by Hay (1930); to Anapsida by Nopcsa (1928) and Roxo (1937); to Reptiliomorphoidea by Huene (1954); to Sauropsida by Haas (1978); to Euamniota by Gardiner (1982); to Pantestudines by Joyce et al. (2004); and to Reptilia by Bonaparte (1850), Cope (1871), Cope (1875), Zittel (1890), Cope (1891), Kuhn (1946), Estes (1964), Estes and Berberian (1970), Carroll (1988), Cvancara and Hoganson (1993), Martinelli and Forasiepi (2004), Sterli (2008) and Li et al. (2008).

Subtaxa
Amblypus, Ancyropus, Aplax oberndorferi, Athecae, Australochelyidae, Australochelys, Chelonipedidae, Chelycarapookus, Chelycarapookus arcuatus, Chelys blakii, Chengyuchelyidae, Chengyuchelys, Chinlechelys, Emydhipus, Emys neumayri, Emys paranensis, Heckerochelyidae, Helcura, Helochelydridae (syn. Solemydidae), Kallokibotion, Megasternon, Mesochelydia, Mongolochelyidae, Odontochelyidae, Palaeochersis, Paracryptodira, Perichelydia, Platemys sehuensis, Platycheloides, Proganochelys (syn. Psammochelys), Proterochersidae, Protochelys, Saltosauropus, Siamochelys, Testudines (syn. Chelonii, Chelonia), Testudinites, Testudo elata, Testudo stricklandi, Testudofiexoolithidae, Testudoolithidae, Thalassochelydia, Thecophora, Tretosternon punctatum, Trionyx mammillaris
Sister taxa
Testudines (syn. Chelonii, Chelonia)

Synonyms
Synonymy list
YearName and author
1811Testudinata Oppel
1826Testudinata Fitzinger p. 4
1842Megasternon Gray
1842Tretosternon punctatum Owen p. 165
1843Aplax oberndorferi Meyer
1843Aplax oberndorferi von Meyer
1850Testudinata Bonaparte p. 1
1863Chelys blakii Mackie p. 41 fig. 1
1863Protochelys blakii Mackie p. 41 fig. 1
1870Megasternon Gray
1871Testudinata Cope p. 235
1871Testudo stricklandi Phillips p. 182
1873Aplax oberndorferi Rütimeyer p. 120
1875Testudinata Cope p. 16
1877Trionyx mammillaris Cope
1877Testudo elata Gervais
1881Emys neumayri Seeley p. 698
1885Emys paranensis Burmeister
1889Platemys sehuensis Ameghino
1889Testudinata Lydekker p. 3
1889Tretosternum punctatum Lydekker pp. 141-143
1889Protochelys Lydekker p. 220
1889Protochelys stricklandi Lydekker pp. 220-222
1890Testudinata Zittel p. 449
1891Testudinata Cope p. 5
1899Platemys sehuensis Ameghino
1902Trionyx mammilaris Hay p. 454
1903Testudinata Osborn p. 276
1904Testudinata Osborn p. 105
1911Testudinata Jaekel p. 183
1928Testudinata Nopcsa p. 164
1930Testudinata Hay p. 64
1937Testudinata Roxo p. 43
1946Testudinata Kuhn p. 58
1954Testudinata Huene p. 23
1958Tretosternum punctatum Delair p. 49
1964Testudinata Estes p. 96
1969Chelycarapookus Warren p. 26
1969Chelycarapookus arcuatus Warren p. 26 figs. plate 2, figs 1, 2A
1970Testudinata Estes and Berberian p. 5
1974Chelonoidis elata Auffenberg
1978Testudinata Haas
1981Testudo elata Campos and de Broin
1982Testudinata Gardiner p. 228
1988Chelycarapookus Carroll
1988Protochelys Carroll
1988Testudinata Carroll
1993Testudinata Cvancara and Hoganson p. 15
1998Chelycarapookus Gaffney et al. p. 6
2004Testudinata Joyce et al.
2004Testudinata Martinelli and Forasiepi p. 260
2008Protochelys blakii Anquetin and Claude
2008Testudinata Li et al. p. 497
2008Testudinata Sterli p. 286

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
—Osteichthyes()
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
RankNameAuthor
—Tetrapoda
—Reptiliomorpha
—Anthracosauria
subclassAmphibiosauriaKuhn 1967
—Cotylosauria()
—Amniota
—Sauropsida
classReptilia
—Testudinata(Oppel 1811)
—Testudinata(Oppel 1811)

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Unr. Testudinata Oppel 1811 [turtle]
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G. †Amblypus Hitchcock 1858
+
†Amblypus dextratus Hitchcock 1858
G. †Ancyropus Hitchcock 1845
+
†Ancyropus heteroclitus Hitchcock 1841
+
Invalid names: Sauroidichnites jacksoni Hitchcock 1841 [synonym]
Fm. †Australochelyidae Gaffney and Kitching 1994
+
G. †Waluchelys Sterli et al. 2020
+
†Waluchelys cavitesta Sterli et al. 2020
G. †Australochelys Gaffney and Kitching 1994
+
†Australochelys africanus Gaffney and Kitching 1994
G. †Chengyuchelys Young and Chow 1953
+
†Chengyuchelys baenoides Young and Chow 1953
†Chengyuchelys latimarginalis Young and Chow 1953
G. †Chinlechelys Joyce et al. 2009
G. †Emydhipus Fuentes Vidarte et al. 2003
G. †Helcura Hitchcock 1848
+
†Helcura anguinea Hitchcock 1858
†Helcura caudata Hitchcock 1858
†Helcura littoralis Hitchcock 1848
†Helcura surgens Hitchcock 1858
Unr. †Helochelydridae Nopcsa 1928
+
G. †Aragochersis Perez-Garcia et al. 2019
+
†Aragochersis lignitesta Perez-Garcia et al. 2019
G. †Helochelydra Nopcsa 1928
+
†Helochelydra anglica Lydekker 1889
†Helochelydra bakewelli Mantell 1833
†Helochelydra nopcsai Lapparent de Broin and Murelaga 1999
G. †Helochelys von Meyer 1854
+
†Helochelys danubina von Meyer 1854
G. †Naomichelys Hay 1908
+
G. †Plastremys Owen 1881
+
†Plastremys lata Owen 1881
†Plastremys rutteri Andrews 1920
Subfm. †Solemydinae Tong et al. 2023
+
G. †Solemys Lapparent de Broin and Murelaga 1996
+
†Solemys gaudryi Matheron 1869
†Solemys vermiculata Lapparent de Broin and Murelaga 1996
G. †Trachydermochelys Seeley 1869
+
G. †Tretosternon Owen 1842
Invalid names: Solemydidae Lapparent de Broin and Murelaga 1996 [objective synonym]
G. †Kallokibotion Nopcsa 1923
+
†Kallokibotion bajazidi Nopcsa 1923
+
Invalid names: Kallokibotion magnificum Nopcsa 1923 [synonym], Thalassodromeus sebesensis Grellet-Tinner and Codrea 2015 [synonym]
Invalid names: Kallokibotion bajazidi Nopcsa 1923 [valid]
Unr. †Mesochelydia Joyce 2017
+
G. †Condorchelys Sterli 2008
+
†Condorchelys antiqua Sterli 2008
G. †Eileanchelys Anquetin et al. 2009
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†Eileanchelys waldmani Anquetin et al. 2009
G. †Heckerochelys Sukhanov 2006
+
†Heckerochelys romani Sukhanov 2006
G. †Indochelys Datta et al. 2000
+
†Indochelys spatulata Datta et al. 2000
G. †Kayentachelys Gaffney et al. 1987
+
†Kayentachelys aprix Gaffney et al. 1987
G. †Palaeochersis Rougier et al. 1995
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†Palaeochersis talampayensis Rougier et al. 1995
Unr. †Paracryptodira Gaffney 1975
+
G. †Arundelemys Lipka et al. 2006
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†Arundelemys dardeni Lipka et al. 2006
Fm. †Baenidae Cope 1882
+
Unr. †Baenodda Gaffney and Meylan 1988
+
G. †Arvinachelys Lively 2015
+
†Arvinachelys goldeni Lively 2015
Tr. †Baenini Cope 1882
+
Invalid names: Eubaenina Williams 1950 [empty]
Unr. †Eubaeninae Williams 1950
+
G. †Baena Leidy 1870
G. †Boremys Lambe 1906
G. †Chisternon Leidy 1872
G. †Denazinemys Lucas and Sullivan 2006
G. †Goleremys Hutchison 2004
G. †Stygiochelys Gaffney and Hiatt 1971
Invalid names: Baena marshi Hay 1904 [nomen dubium], Chisternon interpositum Hay 1909 [nomen dubium]
Unr. †Palatobaeninae Gaffney 1972
+
G. †Cedrobaena Lyson and Joyce 2009
G. †Gamerabaena Lyson and Joyce 2010
G. †Palatobaena Gaffney 1972
G. †Plesiobaena Gaffney 1972
G. †Saxochelys Lyson et al. 2019
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†Saxochelys gilberti Lyson et al. 2019
G. †Scabremys Sullivan et al. 2012
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†Scabremys ornata Gilmore 1935
G. †Edowa Adrian et al. 2023
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†Edowa zuniensis Adrian et al. 2023
G. †Eubaena Hay 1908
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†Eubaena cephalica Hay 1904
†Eubaena hatcheri Hay 1901
G. †Gehennachelys Adrian et al. 2023
+
†Gehennachelys maini Adrian et al. 2019
G. †Hayemys Gaffney 1972
+
†Hayemys latifrons Hay 1908
G. †Lakotemys Joyce et al. 2020
+
†Lakotemys australodakotensis Joyce et al. 2020
G. †Neurankylus Lambe 1902
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†Neurankylus baueri Gilmore 1916
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Invalid names: Neurankylus hutchisoni Lively 2016 [synonym], Neurankylus utahensis Lively 2016 [synonym]
†Neurankylus eximius Lambe 1902
+
Invalid names: Baena fluviatilis Parks 1933 [synonym], Charitemys captans Hay 1908 [synonym]
†Neurankylus lithographicus Larson et al. 2013
†Neurankylus notos Lichtig and Lucas 2016
†Neurankylus torrejonensis Lyson et al. 2016
Invalid names: Charitemys Hay 1908 [synonym]
G. †Peckemys Lyson and Joyce 2009
G. †Probaena Hay 1903
G. †Protobaena Joyce and Lyson 2015
+
†Protobaena wyomingensis Gilmore 1919
G. †Stygiochelys Gaffney and Hiatt 1971
G. †Thescelus Hay 1908
+
†Thescelus insiliens Hay 1908
+
Invalid names: Baena longicauda Russell 1934 [synonym]
†Thescelus rapiens Hay 1908
G. †Trinitichelys Gaffney 1972
+
†Trinitichelys hiatti Gaffney 1972
Invalid names: Baena platyplastra Gilmore 1915 [nomen dubium], Baeninae Cope 1882 [empty], Hayemydinae Gaffney 1972 [empty], Thescelus hemispherica Gilmore 1935 [nomen dubium], Trinitychelyinae Gaffney 1972 [empty]
Superfm. †Baenoidea
+
Invalid names: Eubaenidae [empty], Neurankylidae [empty]
Fm. †Compsemydidae Pérez-García et al. 2015
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G. †Berruchelus Pérez-García 2012
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†Berruchelus russelli Pérez-García 2012
G. †Peltochelys Dollo 1884
+
G. †Tongemys Joyce et al. 2022
+
†Tongemys enigmatica Joyce et al. 2022
G. †Compsemys Leidy 1856
+
†Compsemys victa Leidy 1856
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Invalid names: Compsemys parva Hay 1910 [synonym], Compsemys puercensis Gilmore 1919 [synonym], Compsemys torrejonensis Gilmore 1919 [synonym], Compsemys vafer Hay 1910 [synonym]
Invalid names: Emys obscurus Leidy 1856 [nomen dubium], Glyptops depressus Hay 1908 [nomen dubium]
Fm. †Pleurosternidae Cope 1868
+
G. †Ballerstedtia Karl et al. 2012
+
†Ballerstedtia typocardia Seeley 1869
G. †Desmemys Wegner 1911
G. †Dinochelys Gaffney 1979
+
†Dinochelys whitei Gaffney 1979
G. †Dorsetochelys Evans and Kemp 1976
+
+
Invalid names: Ballerstedtia bueckebergensis Karl et al. 2012 [synonym], Dorsetochelys delairi Evans and Kemp 1976 [synonym], Thalassemys ruetimeyeri Lydekker 1889 [synonym]
Invalid names: Dorsetochelys buzzops Bakker 1998 [nomen dubium]
G. †Glyptops Marsh 1890
+
†Glyptops ornatus Marsh 1890
+
Invalid names: Glyptops utahensis Gilmore 1916 [synonym]
Invalid names: Glyptops plicatulus Cope 1877 [nomen dubium]
G. †Pleurosternon Owen 1853
+
+
Invalid names: Megasternon koenigii Gray 1870 [synonym], Mesochelys durlstonensis Evans and Kemp 1975 [synonym], Pleurosternon concinnum Owen 1853 [synonym], Pleurosternon emarginatum Owen 1853 [synonym], Pleurosternon ovatum Owen 1853 [synonym], Pleurosternon oweni Seeley 1869 [synonym], Pleurosternon sedgwicki Seeley 1869 [synonym], Pleurosternon vansittarti Seeley 1869 [synonym], Pleurosternum portlandicum Lydekker 1889 [synonym]
†Pleurosternon moncayensis Pérez-Garcia et al. 2021
Invalid names: Digerrhum Cope 1870 [synonym], Mesochelys Evans and Kemp 1975 [synonym]
G. †Riodevemys Pérez-García et al. 2015
+
†Riodevemys inumbragigas Pérez-García et al. 2015
G. †Selenemys Pérez-García and Ortega 2011
+
†Selenemys lusitanica Pérez-García and Ortega 2011
G. †Toremys Pérez-García et al. 2015
+
†Toremys cassiopeia Pérez-García et al. 2015
G. †Uluops Carpenter and Bakker 1990
+
†Uluops uluops Carpenter and Bakker 1990
Invalid names: Desmemydinae Nopcsa 1928 [empty]
Invalid names: Desmemys bertelmanni Wegner 1911 [nomen dubium], Glyptops caelatus Hay 1908 [nomen dubium], Glyptops pervicax Hay 1908 [nomen dubium], Polythorax Cope 1876 [nomen dubium], Polythorax missuriensis Cope 1876 [nomen dubium], Probaena sculpta Hay 1903 [nomen dubium]
Unr. †Perichelydia Joyce 2017
+
Unr. †Meiolaniformes Sterli and de la Fuente 2013
+
G. †Chubutemys Gaffney et al. 2007
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†Chubutemys copelloi Gaffney et al. 2007
Fm. †Meiolaniidae Lydekker 1887
+
G. †Gaffneylania Sterli et al. 2015
+
†Gaffneylania auricularis Sterli et al. 2015
G. †Meiolania Owen 1886
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†Meiolania brevicollis Megirian 1992
†Meiolania mackayi Anderson 1925
†Meiolania platyceps Owen 1886
+
Invalid names: Ceratochelys sthenurus Huxley 1887 [objective synonym], Ceratochelys sthenurus Huxley 1887 [objective synonym], Meiolania minor Owen 1886 [synonym], Meiolania minor Owen 1886 [synonym]
G. †Ninjemys Gaffney 1992
+
†Ninjemys oweni Woodward 1888
G. †Niolamia Ameghino 1899
+
†Niolamia argentina Ameghino 1899
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Invalid names: Miolania argentina Woodward 1901 [synonym]
Invalid names: Crossochelys Simpson 1937 [synonym]
G. †Warkalania Gaffney et al. 1992
+
†Warkalania carinaminor Gaffney et al. 1992
Invalid names: Gaffneylania Sterli et al. 2015 [valid], Meiolania Owen 1886 [valid], Miolaniidae [synonym], Miolaniidae [synonym], Ninjemys Gaffney 1992 [valid], Niolamia Ameghino 1899 [valid], Warkalania Gaffney et al. 1992 [valid]
G. †Otwayemys Gaffney et al. 1998
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†Otwayemys cunicularius Gaffney et al. 1998
G. †Patagoniaemys Sterli and de la Fuente 2011
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†Patagoniaemys gasparinae Sterli and de la Fuente 2011
G. †Peligrochelys Sterli and de la Fuente 2013
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†Peligrochelys walshae Sterli and de la Fuente 2013
G. †Trapalcochelys Sterli et al. 2013
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†Trapalcochelys sulcata Sterli et al. 2013
Unr. †Sichuanchelyidae Tong et al. 2012
+
G. †Laurasichersis Pérez-García 2020
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†Laurasichersis relicta Pérez-García 2020
G. †Mongolochelys Khosatzky 1997
+
†Mongolochelys efremovi Khosatzky 1997
G. †Sichuanchelys Ye and Pi 1997
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†Sichuanchelys chowi Ye and Pi 1997
†Sichuanchelys palatodentata Joyce et al. 2016
G. †Spoochelys Smith and Kear 2013
+
†Spoochelys ormondea Smith and Kear 2013
G. †Platycheloides Haughton 1928
G. †Proganochelys Baur 1887
+
+
Invalid names: Psammochelys keuperina Quenstedt 1889 [objective synonym], Stegochelys dux Jaekel 1914 [synonym]
†Proganochelys ruchae de Broin 1984
†Proganochelys tenertesta Joyce et al. 2009
Invalid names: Psammochelys Quenstedt 1889 [synonym]
Fm. †Proterochersidae Nopcsa 1928
+
G. †Keuperotesta Szczygielski and Sulej 2016
G. †Proterochersis Fraas 1913
+
†Proterochersis limendorsa Szczygielski and Sulej 2016
†Proterochersis porebensis Szczygielski and Sulej 2016
†Proterochersis robusta Fraas 1913
+
Invalid names: Chelytherium obscurum Meyer 1863 [synonym], Chelytherium obscurum Meyer 1863 [synonym], Murrhardtia staeschei Karl and Tichy 2000 [synonym], Murrhardtia staeschei Karl and Tichy 2000 [synonym], Proterochersis intermedia Fraas 1913 [synonym], Proterochersis intermedia Fraas 1913 [synonym]
†Proterochersis robusta Fraas 1913
+
Invalid names: Chelytherium obscurum Meyer 1863 [synonym], Chelytherium obscurum Meyer 1863 [synonym], Murrhardtia staeschei Karl and Tichy 2000 [synonym], Murrhardtia staeschei Karl and Tichy 2000 [synonym], Proterochersis intermedia Fraas 1913 [synonym], Proterochersis intermedia Fraas 1913 [synonym]
Invalid names: Chelytherium Meyer 1863 [synonym], Murrhardtia Karl and Tichy 2000 [synonym]
G. †Saltosauropus Bernier et al. 1984
+
†Saltosauropus latus Bernier et al. 1984
G. †Siamochelys Tong et al. 2002
+
†Siamochelys peninsularis Tong et al. 2002
Or. Testudines Batsch 1788 [turtle]
+
Subor. †Amphichelydia Lydekker 1889
+
Superfm. †Pleurosternoidea Hay 1930
+
Fm. †Kallokibotiidae Nopcsa 1923
+
G. †Kallokibotion Nopcsa 1923
+
†Kallokibotion bajazidi Nopcsa 1923
+
Invalid names: Kallokibotion magnificum Nopcsa 1923 [synonym], Thalassodromeus sebesensis Grellet-Tinner and Codrea 2015 [synonym]
Invalid names: Kallokibotion bajazidi Nopcsa 1923 [valid]
Invalid names: Apertotemporalidae [empty]
Superfm. †Proganochelyoidea Baur 1887
+
Invalid names: Archaeochelydiidae Bergounioux 1938 [empty]
G. †Asturichelys Bergounioux 1957
G. †Chelonipus von Lilienstern 1939
+
†Chelonipus cuneiformis von Lilienstern 1939
†Chelonipus plieningeri Haubold 1971
†Chelonipus torquatus von Lilienstern 1939
Subor. Cryptodira Cope 1868
+
Fm. †Bashuchelyidae Tong et al. 2012
+
G. †Bashuchelys Tong et al. 2012
+
†Bashuchelys youngi Tong et al. 2012
G. †Chuannanchelys Tong et al. 2012
+
Fm. Chelonemydidae Rütimeyer 1872
+
Invalid names: Propleuridae Cope 1870 [synonym]
Superfm. Chelonioidea Baur 1893 [sea turtle]
+
Fm. Cheloniidae Bonaparte 1832
+
Subfm. †Allopleuroninae
G. †Ashleychelys Weems and Sanders 2014
G. †Cabindachelys Myers et al. 2018
Subfm. Carettinae Gray 1825
G. Chelone Linnaeus 1758
Subfm. Cheloniinae Oppel 1811
G. †Chelyopsis Smets 1887
G. †Dollochelys Zangerl 1971
G. †Eochelone Dollo 1903
G. †Erquelinnesia Dollo 1887
G. †Gigantatypus Kaddumi 2006
G. †Itilochelys Danilov et al. 2010
G. †Lembonax Cope 1870
G. Lepidochelys Fitzinger 1843 [Ridley sea turtle]
+
Invalid names: Cephalochelys Gray 1873 [synonym]
G. †Lusochelys Pérez-García and Antunes 2024
G. †Lytoloma Cope 1870
G. †Miocaretta Deraniyagala 1967
G. †Osonachelus de Lapparent de Broin et al. 2014
G. †Pacifichelys Parham and Pyenson 2010
G. †Procolpochelys Hay 1908
G. †Protrachyaspis Zvonok et al. 2024
G. †Puppigerus Cope 1870
+
Invalid names: Eochelys Moody 1968 [objective synonym]
Subfm. †Syllominae
G. †Syllomus Cope 1896
G. †Tasbacka Nesov 1986
G. †Trachyaspis von Meyer 1843
G. Zangerlchelys Hirayama 2006
Invalid names: Chelone acuticostata Bergounioux 1954 [nomen vanum], Chelone bellunensis Bergounioux 1954 [nomen vanum], Chelone gastaldii Portis 1879 [nomen vanum], Chelone murua de Vis 1905 [nomen dubium], Chelone simonellii Porta 1898 [nomen vanum], Chelone sismondai Portis 1879 [nomen vanum], Chelone sordellii Porta 1898 [nomen vanum], Chelone strobeli Porta 1898 [nomen vanum], Eocheloniinae [empty], Eochelyinae Moody 1967 [empty], Eretmochelyinae Deranyiagala 1953 [empty], Pliochelys derelicta Portis 1890 [nomen vanum], Progonosaurus pertinax Portis 1890 [nomen vanum]
G. †Cimochelys Zangerl 1960
+
†Cimochelys benstedi Mantell 1841
Fm. Dermochelyidae Lydekker 1889 [leatherback]
+
Subfm. †Dermochelyinae
G. Dermochelys de Blainville 1816 [Leatherback sea turtle]
+
Invalid names: Chelyra Rafinesque 1832 [synonym], Coriudo Fleming 1822 [objective synonym], Dermatochelys Wagler 1830 [synonym], Dermochelis Cuvier 1829 [synonym], Scytina Wagler 1828 [synonym], Sphargis Merrem [synonym]
G. †Egyptemys Wood et al. 1996
G. †Eosphargis Lydekker 1889
+
Invalid names: Anglocetus Tarlo 1963 [synonym]
G. †Maorichelys Karl and Tichy 2007
G. †Mesodermochelys Hirayama and Chitoku 1996
G. †Natemys Wood et al. 1996
G. †Psephophorus von Meyer 1847
G. †Turgaiscapha Averianov 2002
Invalid names: Chelonia californiensis Gilmore 1937 [invalid subgroup], Psephophoridae Goette 1899 [synonym], Sphargidae Gray 1825 [synonym]
Unr. †Dermochelyoidae Williams 1950
+
G. †Alienochelys de Lapparent de Broin et al. 2014
G. †Corsochelys Zangerl 1960
G. †Ocepechelon Bardet et al. 2013
Fm. †Desmatochelyidae Williston 1894
+
G. †Atlantochelys Agassiz 1849
G. †Neptunochelys Wieland 1900
G. †Kimurachelys Brinkman et al. 2015
+
†Kimurachelys slobodae Brinkman et al. 2015
G. †Leviathanochelys Castillo-Visa et al. 2022
+
†Leviathanochelys aenigmatica Castillo-Visa et al. 2022
G. †Nichollsemys Brinkman et al. 2006
+
†Nichollsemys baieri Brinkman et al. 2006
Fm. †Protostegidae Cope 1873
+
G. †Archelon Wieland 1896
G. †Bouliachelys Kear and Lee 2006
Subfm. †Chelospharginae Zangerl 1953
G. †Chelosphargis Zangerl 1953
G. †Cratochelone Longman 1915
G. †Desmatochelys Williston 1894
G. †Notochelone Lydekker 1889
+
Invalid names: Notochelys Owen 1882 [replaced]
G. †Pneumatoarthrus Cope 1870
G. †Protostega Cope 1871
Subfm. †Protosteginae Wieland
G. †Rhinochelys Seeley 1869
G. †Santanachelys Hirayama 1998
G. †Terlinguachelys Lehman and Tomlinson 2004
Fm. †Toxochelyidae Baur 1895
+
G. †Cynocercus Cope 1872
Subfm. †Lophochelyinae Zangerl 1953
Subfm. †Macrobaeninae Nessov 1984
Subfm. †Osteopyginae Zangerl 1953
+
Invalid names: Lytolominae [synonym]
G. †Porthochelys Williston 1901
Subfm. †Toxochelyinae Zangerl 1953
Invalid names: Glyptops belviderensis Cragin 1902 [nomen dubium]
Invalid names: Pandermochelys Joyce 2004 [empty], Pliochelys Portis 1890 [nomen vanum], Progonosaurus Portis 1890 [nomen vanum]
Fm. Chelydridae Swainson 1839
+
G. Chelydra Schweigger 1812 [Snapping turtle]
+
Chelydra acutirostris Peters 1862 [South American snapping turtle]
Chelydra rossignonii Bocourt 1868 [Central American snapping turtle]
Chelydra serpentina Linnaeus 1758 [Common snapping turtle]
+
Invalid names: Chelydra emarginata Agassiz 1857 [synonym], Chelydra lacertina Schweigger 1812 [synonym], Chelydra laticarinata Hay 1916 [synonym], Chelydra osceola Stejneger 1918 [synonym], Chelydra sculpta Hay 1916 [synonym], Devisia mythodes Ogilby 1905 [synonym], Macrochelys floridana Hay 1907 [synonym], Testudo longicauda Shaw in Gray 1831 [synonym], Testudo serpentaria Wiedemann 1802 [synonym], Testudo serrata Pennant in Gray 1830 [synonym]
Invalid names: Cheliurus Rafinesque 1815 [nomen nudum], Chelonura Fleming 1822 [synonym], Devisia Ogilby 1905 [synonym], Emysaurus Dumeril and Bibron 1835 [synonym], Ophichelone Jarocki 1822 [synonym], Rapara Gray 1825 [synonym], Saurochelys Latreille 1825 [synonym]
G. †Chelydrops Peters 1868
Subfm. †Chelydropsinae Mlynarski 1980
+
G. †Chelydrasia Ckhikvadze 1999
G. †Chelydropsis Peters 1868
+
†Chelydropsis apellanizi Murelaga et al. 1999
†Chelydropsis aubasi Joyce et al. 2022
†Chelydropsis decheni Meyer 1852
+
Invalid names: Chelydropsis sanctihenrici Broin 1977 [synonym]
†Chelydropsis heweneggensis Pappa et al. 2024
†Chelydropsis kusnetzovi Chkhikvadze 1985
†Chelydropsis manuascensis Bergounioux 1936
+
Invalid names: Chelydra allingensis Fuchs 1939 [synonym], Chelydropsis carinata Peters 1868 [synonym], Chelydropsis murchisoni staeschei Mlynarski 1980 [synonym], Chelydropsis staeschei Mlynarski 1980 [synonym], Trionyx sansaniensis Bergounioux 1935 [synonym]
†Chelydropsis pontica Pidoplichko and Tarashchuk 1960
†Chelydropsis sansaniensis Bergounioux 1935
Invalid names: Macrocephalochelys Pidoplichko and Tarashchuk 1960 [synonym]
G. Macrochelys Gray 1856 [alligator snapping turtle]
+
†Macrochelys auffenbergi Dobie 1968 [Auffenberg's snapping turtle]
†Macrochelys schmidti Zangerl 1945
†Macrochelys stricta Matthew 1924
Macrochelys suwanniensis Thomas et al. 2014 [Suwannee alligator snapping turtle]
Macrochelys temminckii Troost 1835 [Alligator snapping turtle]
+
Invalid names: Gypochelys lacertina Agassiz 1857 [synonym], Macrochelys apalachicolae Thomas et al. 2014 [synonym]
Invalid names: Gypochelys Agassiz 1857 [synonym]
G. †Planiplastron Ckhikvadze 1971
Invalid names: Broilia Bergounioux and Crouzel 1965 [nomen dubium], Broilia robusta Bergounioux and Crouzel 1965 [nomen dubium], Chelydra argillarum Laube 1900 [nomen dubium], Chelydra strausi Schmidt 1966 [nomen dubium], Chelydrinae Lydekker 1889 [empty], Chelydropsis minax Ckhikvadze 1971 [nomen dubium], Emys sansaniensis Lartet 1851 [nomen dubium], Emysaurus meilheuratiae Pomel 1846 [nomen dubium], Hydraspis oeningensis Fitzinger 1835 [nomen nudum], Leptochelys Bergounioux and Crouzel 1965 [nomen dubium], Leptochelys braneti Bergounioux and Crouzel 1965 [nomen dubium], Protochelydrinae Gaffney 1975 [empty], Testudo grandis Macarovici and Vancea 1959 [nomen dubium]
Unr. Chelydroidea Baur 1893
+
Unr. †Panchelydridae Joyce et al. 2004
+
G. †Denverus Hutchison and Holroyd 2003
G. †Protochelydra Erickson 1973
Invalid names: Hoplochelys caelata Hay 1909 [nomen dubium]
Unr. †Pankinosternoidea Joyce et al. 2004
+
G. †Emarginachelys Whetstone 1978
G. †Lutemys Lyson et al. 2017
G. †Planetochelys Weems 1988
G. †Tullochelys Hutchison 2013
G. †Chitracephalus Dollo 1884
+
†Chitracephalus dumonii Dollo 1885
+
Invalid names: Salasemys pulcherrima Fuentes et al. 2003 [synonym]
Invalid names: Salasemys Fuentes et al. 2003 [synonym]
Superfm. †Dermochelyoidea
+
G. †Pygmaeochelys Laube 1896
Invalid names: Chelone regularis Fritsch 1905 [nomen dubium], Pygmaeochelys michelobana Laube 1896 [nomen dubium]
Unr. Durocryptodira Danilov and Parham 2008
+
G. †Toxochelys Cope 1873
+
†Toxochelys elkader Hay 1908
†Toxochelys latiremis Cope 1873
+
Invalid names: Porthochelys browni Hay 1905 [synonym], Toxochelys barberi Schmidt 1944 [synonym], Toxochelys brachyrhinus Case 1898 [synonym], Toxochelys serrifer Cope 1875 [synonym], Toxochelys weeksi Collins 1951 [synonym]
†Toxochelys moorevillensis Zangerl 1953
†Toxochelys procax Hay 1905
Unr. Eucryptodira Gaffney 1975
+
G. †Borealochelys Brinkman and Tarduno 2005
+
†Borealochelys axelheibergensis Brinkman and Tarduno 2005
G. †Enaliochelys Seeley 1869
+
†Enaliochelys chelonia Seeley 1869
G. †Hoyasemys Pérez-García et al. 2012
+
†Hoyasemys jimenezi Pérez-García et al. 2012
Fm. †Sandownidae Tong and Meylan 2013
+
G. †Angolachelys Mateus et al. 2009
G. †Brachyopsemys Tong and Meylan 2013
G. †Leyvachelys Cadena 2015
G. †Sandownia Meylan et al. 2000
Fm. †Sinemydidae Yeh 1963
+
G. †Dracochelys Gaffney and Ye 1992
G. †Hongkongochelys Ye 1999
G. †Jeholochelys Shao et al. 2018
G. †Liaochelys Zhou 2010
G. †Manchurochelys Endo and Shikama 1942
G. †Ordosemys Brinkman and Peng 1993
G. †Sinemys Wiman 1930
G. †Wuguia Matzke et al. 2004
G. †Xiaochelys Zhou and Rabi 2015
G. †Yumenemys Bohlin 1953
Invalid names: Kirgizemyinae Nessov and Khozatskiy 1973 [empty]
Fm. †Sinochelyidae Chkhikvadze 1970
+
G. †Sinochelys Wiman 1930
+
Invalid names: Heishanemys Bohlin 1953 [synonym], Peishanemys Bohlin 1953 [synonym], Scutemys Wiman 1930 [synonym]
Invalid names: Peishanemydidae Nessov 1981 [synonym]
G. †Tienfucheloides Nessov 1978
+
†Tienfucheloides undatus Nessov 1978
Fm. †Xinjiangchelyidae Nessov 1990
+
G. †Annemys Sukhanov and Narmandakh 2006
+
Invalid names: Annemys Sukhanov and Narmandakh 2000 [replaced]
G. †Brodiechelys Nopcsa 1928
G. †Camerochelys Pérez-García and Murelaga 2013
G. †Kalasinemys Tong et al. 2019
G. †Phunoichelys Tong et al. 2014
G. †Protoxinjiangchelys Tong et al. 2012
G. †Shartegemys Sukhanov and Narmandakh 2006
G. †Tienfuchelys Young and Chow 1953
G. †Undjulemys Sukhanov and Narmandakh 2006
G. †Xinjiangchelys Yeh 1986
Invalid names: Shartegemys Sukhanov and Narmandakh 2000 [nomen nudum], Undjulemys Sukhanov and Narmandakh 2000 [nomen nudum]
Invalid names: Angolachelonia Mateus et al. 2009 [empty], Centrocryptodira Gaffney and Meylan 1988 [empty], Polycryptodira Gaffney and Meylan 1988 [empty]
G. †Galvechelone Pérez-García and Murelaga 2012
+
†Galvechelone lopezmartinezae Pérez-García and Murelaga 2012
Fm. Kinosternoidea Hay 1892 [mud turtle]
+
G. Aromochelys Gray
G. Cinosternum von Spix 1824 [mud turtle]
Fm. Kinosternidae Hay 1892
Unr. †Pandermatemys Joyce et al. 2004
G. Sternotherus Gray 1825 [musk turtle]
Invalid names: Aromochelys Gray [valid], Cinosternum von Spix 1824 [valid], Kinosternidae Hay 1892 [valid], Pandermatemys Joyce et al. 2004 [valid], Sternotherus Gray 1825 [valid]
Fm. †Nanhsiungchelyidae Yeh 1966
+
G. †Anomalochelys Hirayama et al. 2001
+
†Anomalochelys angulata Hirayama et al. 2001
G. †Basilemys Hay 1902
+
†Basilemys gaffneyi Sullivan et al. 2012
†Basilemys imbricarius Cope 1876
†Basilemys morrinensis Mallon and Brinkman 2018
†Basilemys nobilis Hay 1911
†Basilemys ogmius Cope 1875
†Basilemys praeclara Hay 1911
†Basilemys sinuosa Riggs 1906
†Basilemys variolosa Cope 1876
G. †Bulganemys Sukhanov and Narmandakh 2006
+
Invalid names: Bulganemys jaganchobili Sukhanov and Narmandakh 2000 [nomen nudum]
G. †Hanbogdemys Sukhanov and Narmandakh 2006
+
†Hanbogdemys jaganchobili Sukhanov and Narmandakh 2006
†Hanbogdemys orientalis Sukhanov and Narmandakh 1975
G. †Jiangxichelys Tong and Mo 2010
+
†Jiangxichelys ganzhouensis Tong and Mo 2010
†Jiangxichelys neimongolensis Brinkman and Peng 1996
G. †Kharakhutulia Sukhanov et al. 2008
+
†Kharakhutulia kalandadzei Sukhanov et al. 2008
G. †Nanhsiungchelys Yeh 1966
+
†Nanhsiungchelys yangi Ke et al. 2023
G. †Xianyuechelys Ke et al. 2024
+
†Xianyuechelys yingliangi Ke et al. 2024
G. †Yuchelys Tong et al. 2012
+
†Yuchelys nanyangensis Tong et al. 2012
G. †Zangerlia Mlynarski 1972
+
†Zangerlia dzamynchondi Sukhanov and Narmandakh 2006
†Zangerlia testudinimorpha Mlynarski 1972
†Zangerlia ukhaachelys Joyce and Norell 2005
Invalid names: Zangerlia dzamynchondi Sukhanov and Narmandakh 2000 [nomen nudum]
Invalid names: Bulganemys Sukhanov and Narmandakh 2000 [nomen nudum], Charitonyx Chkhikvadze 1980 [nomen dubium], Hanbogdemysa Sukhanov 2000 [nomen nudum]
Unr. Panchelonioidea Joyce et al. 2004
+
G. †Mexichelys Parham and Pyenson 2010
Unr. Pancheloniidae Joyce et al. 2004
+
G. †Argillochelys Lydekker 1889
G. †Australobaena Karl and Tichy 2002
G. †Carolinochelys Hay 1923
Fm. †Ctenochelyidae Gentry 2018
G. †Euclastes Cope 1870
+
Invalid names: Catapleura Cope 1870 [synonym], Glossochelys Seeley 1871 [synonym], Osteopygoides Karl et al. 1998 [synonym], Pampaemys de la Fuente and Casadio 2000 [synonym]
G. †Peritresius Cope 1870
G. †Prionochelys Zangerl 1953
Invalid names: Sphenophyllum colombianum Huertas 2003 [nomen vanum]
Unr. †Pantestudinoidea Joyce et al. 2004
+
Superfm. Testudinoidea Gray 1825
+
G. †Bergouniouxchelys Hervet 2004
G. †Borkenia Schleich 1994
G. †Cucullemys Hervet 2004
G. †Cuvierichelys Hervet 2004
Fm. Emydidae Rafinesque 1815
+
Invalid names: Cistudinidae Cope 1882 [synonym]
G. †Euroemys Hervet 2004
G. †Francellia Hervet 2004
Fm. Geoemydidae Theobald 1868
G. †Landreatchelys Hervet 2004
Fm. †Lindholmemydidae Chkhikvadze 1970
G. †Owenemys Hervet 2004
G. †Palaeomauremys Hervet 2004
Unr. †Panemydidae Joyce et al. 2004
Unr. †Pangeoemydidae Joyce 2007
Unr. †Pantestudinidae Joyce et al. 2004
G. †Promalacoclemmys Reinach 1900
G. †Provencemys Hervet 2004
Fm. Testudinidae Batsch 1788
G. †Wutuchelys Tong et al. 2017
Invalid names: Chrysemys (Pseudemys) hibbardi Preston 1979 [nomen dubium], Clemmys hesperia Hay 1903 [nomen dubium], Clemmys percrassus Cope 1899 [nomen dubium], Emys euglypha Leidy 1889 [nomen dubium], Emys euthnetus Cope 1873 [nomen dubium], Emys latilabiatus Cope 1871 [nomen dubium], Emys megaulax Cope 1873 [nomen dubium], Emys pachylomus Cope 1873 [nomen dubium], Emys petrolei Leidy 1868 [nomen dubium], Eupachemys obtusus Leidy 1877 [nomen dubium], Hadrianus allabiatus Cope 1871 [nomen dubium], Hybemys arenarius Leidy 1871 [nomen dubium], Notomorpha testudinea Cope 1872 [nomen dubium], Rhinoclemmys nicoyana Acuña-Mesen and Laurito-Mora 1996 [nomen dubium], Testudo culbratus Cope 1873 [nomen dubium], Testudo distans Hay 1916 [nomen dubium], Testudo isis Andrews 1906 [nomen dubium], Testudo munda Hay 1920 [nomen dubium], Trachemys delicata Hay 1916 [nomen dubium], Trachemys nuchocarinata Hay 1916 [nomen dubium]
G. †Polysternum Bergounioux 1935
Unr. †Selmacryptodira Gaffney et al.
+
Invalid names: Kallokibotionidae Gaffney and Meylan 1992 [empty]
Subor. Trionychia
+
Unr. Carettochelyidae Gill 1889
+
G. †Akrochelys Bergounioux 1935
G. †Anosteira Leidy 1871
Subfm. †Anosteirinae Lydekker 1889
Unr. †Carettochelyinae Boulenger 1887
G. Carettochelys Ramsay 1886 [pig-nose turtle]
G. †Kizylkumemys Nessov 1977
G. †Pseudanosteira Clark 1932
Invalid names: Allaeochelyinae Bergounioux 1952 [empty], Anosteira radulina Cope 1872 [nomen dubium], Apholidemys granosa Pomel 1847 [nomen nudum], Apholidemys sublaevis Pomel 1847 [nomen nudum], Chorlakkichelys de Broin 1987 [nomen dubium], Chorlakkichelys shahi de Broin 1987 [nomen dubium]
G. †Kappachelys Hirayama et al. 2013
+
†Kappachelys okurai Hirayama et al. 2013
Unr. †Pantrionychidae Joyce et al. 2004
+
G. †Drazinderetes Head et al. 1999
G. †Khunnuchelys Brinkman et al. 1993
G. †Murgonemys White 2001
Unr. †Plastomenidae Hay 1902
Invalid names: Amyda egregia Hay 1908 [nomen dubium], Amyda salebrosa Hay 1908 [nomen dubium], Anostira molopina Cope 1872 [nomen dubium], Anostira oedemia Cope 1872 [nomen dubium], Anostira trionychoides Cope 1872 [nomen dubium], Aspideretes amnigenus Hay 1910 [nomen dubium], Aspideretes perplexus Gilmore 1919 [nomen dubium], Plastomenus catenatus Cope 1875 [nomen dubium], Plastomenus coalescens Cope 1875 [nomen dubium], Plastomenus communis Cope 1875 [nomen dubium], Plastomenus corrugatus Cope 1875 [nomen dubium], Plastomenus costatus Cope 1875 [nomen dubium], Plastomenus fractus Cope 1875 [nomen dubium], Plastomenus insignis Cope 1874 [nomen dubium], Plastomenus lachrymalis Cope 1875 [nomen dubium], Plastomenus multifoveatus Cope 1873 [nomen dubium], Plastomenus punctulatus Cope 1874 [nomen dubium], Plastomenus serialis Cope 1877 [nomen dubium], Plastomenus torrejonensis Gilmore 1919 [nomen dubium], Platypeltis amnicola Hay 1907 [nomen dubium], Platypeltis antiqua Hay 1907 [nomen dubium], Platypeltis trepida Hay 1907 [nomen dubium], Procyclanorbis sardus Portis 1901 [nomen dubium], Trionyx buiei Cope 1870 [nomen dubium], Trionyx cariosus Cope 1875 [nomen dubium], Trionyx cellulosus Cope 1868 [nomen dubium], Trionyx concentricus Cope 1872 [nomen dubium], Trionyx guttatus Leidy 1869 [nomen dubium], Trionyx halophilus Cope 1869 [nomen dubium], Trionyx heteroglyptus Cope 1873 [nomen dubium], Trionyx leptomitus Cope 1875 [nomen dubium], Trionyx pennatus Cope 1869 [nomen dubium], Trionyx priscus Cope 1869 [nomen dubium], Trionyx punctiger Cope 1891 [nomen dubium], Trionyx quinni Holman 1982 [nomen dubium], Trionyx radulus Cope 1875 [nomen dubium], Trionyx vagans Cope 1874 [nomen dubium], Trionyx ventricosus Cope 1877 [nomen dubium], Trionyx virginiana Clark 1895 [nomen dubium]
Fm. Trionychidae Gray 1825 [softshell turtle]
+
G. Apalone Rafinesque 1832 [softshell turtle]
+
Subg. Apalone (Platypeltis) Fitzinger 1835
†Apalone amorense Valdes et al. 2017
Apalone ferox Schneider 1783 [Florida softshell turtle]
†Apalone lima Cope 1869
Apalone mutica LeSueur 1827 [smooth softshell turtle]
Apalone spinifera LeSueur 1827 [spiny softshell turtle]
†Apalone trionychoides Cope 1872
G. Aspideretes Hay 1904
+
Aspideretes gangeticus Cuvier 1825 [Indian softshell]
Aspideretes hurum Gray 1831 [Indian peacock softshell]
Aspideretes leithii Gray 1872 [Leith's softshell]
Aspideretes nigricans Anderson 1875 [Black softshell]
†Aspideretes sinuosus Chow and Yeh 1958
G. Aspidonectes Wagler 1830
Subfm. †Chitrainae Jasinski et al. 2022
+
G. Chitra Gray 1844
G. Pelochelys Gray 1864
G. †Conchochelys Hay 1905
G. †Crassithecachelys Chkhikvadze 2000
Subfm. Cyclanorbinae Hummel 1929
+
G. Cyclanorbis Gray 1854
G. Cycloderma Peters 1854
G. Lissemys Smith 1931
G. †Nemegtemys Danilov et al. 2014
G. †Eugenechelys Chkhikvadze 2000
G. †Hummelichelys Chkhikvadze 2000
Subfm. †Kuhnemydinae Jasinski et al. 2022
+
G. †Kuhnemys Chkhikvadze 1999
G. †Perochelys Li et al. 2015
G. †Petrochelys Vitek et al. 2017
G. †Paleotrionyx Schmidt 1945
G. †Striatochelys Massonne et al. 2023
+
†Striatochelys baba Massonne et al. 2023
G. †Temnotrionyx Hay 1908
Subfm. Trionychinae Gray 1825 [softshell turtle]
+
G. †Aulacochelys Lydekker 1889
G. †Axestemys Hay 1899
+
Invalid names: Axestus Cope 1872 [synonym], Eurycephalochelys Moody and Walker 1970 [synonym]
Tr. Chitrini Gray 1870
G. Dogania Gray 1844
G. Nilssonia Gray 1869 [Burmese peacock softshells]
G. Palea Siebenrock 1906 [Wattleneck softshells]
G. Pelodiscus Fitzinger 1835
G. Rafetus Gray 1864
Tr. Trionychini Gray 1825
G. Trionyx Geoffroy Saint-Hilaire 1809
+
Invalid names: Francedebroinella Chkhikvadze 1999 [synonym], Sinamyda Chkhikvadze 2000 [synonym]
Invalid names: Trionyx (Amyda) subangularis Bergounioux 1954 [nomen dubium], Trionyx bambolii Ristori 1891 [nomen dubium], Trionyx blayaci Bergounioux 1933 [nomen dubium], Trionyx desmostyli Matsumoto 1918 [nomen dubium], Trionyx dieupentalensis Bergounioux 1935 [nomen dubium], Trionyx harmati Szalai 1934 [nomen dubium], Trionyx irregularis Bergounioux 1954 [nomen dubium], Trionyx pliopedemontana Sacco 1889 [nomen dubium], Trionyx stadleri Teppner 1913 [nomen dubium], Trionyx stiriacus Peters 1855 [nomen dubium]
G. †Zangerlichelys Chkhikvadze 2000
Invalid names: Altaytrionyx Chkhikvadze 2008 [nomen dubium], Altaytrionyx burtschaki Chkhikvadze 2008 [nomen dubium], Altaytrionyx devjatkini Chkhikvadze 2008 [nomen dubium], Altaytrionyx phiruzae Chkhikvadze 2008 [nomen dubium], Amyda menneri Chkhikvadze 1988 [nomen dubium], Aspideretes jaxarticus Riabinin 1938 [nomen nudum], Aspideretes muyuensis Lei and Ye 1985 [nomen dubium], Aspilus Portis 1885 [nomen dubium], Aspilus cortesii Portis 1885 [nomen dubium], Chitra minor Jaekel 1911 [nomen dubium], Lissemydinae Williams 1950 [empty], Paleotrionyx jimenezfuentesi Chkhikvadze 1995 [nomen dubium], Plastomenus gabunii Chkhikvadze 1984 [nomen dubium], Plastomenus jaxarticus Riabinin 1938 [nomen nudum], Plastomenus mlynarskii Chkhikvadze 1970 [nomen dubium], Platypeltis subcircularis Chow and Yeh 1957 [nomen dubium], Procyclanorbis Portis 1901 [nomen dubium], Rafetini Chkhikvadze 2000 [empty], Rafetus karkhualexandri Chkhikvadze 2000 [nomen nudum], Rafetus yexiangkuii Chkhikvadze 1999 [nomen dubium], Trionyx acutiformis Bergounioux 1935 [nomen dubium], Trionyx amansii Gray 1831 [nomen dubium], Trionyx anthracotheriorum Portis 1883 [nomen dubium], Trionyx aquitanicus Delfortrie 1869 [nomen dubium], Trionyx aspidiformis Laube 1900 [nomen dubium], Trionyx australiensis De Vis 1894 [nomen dubium], Trionyx austriacus Peters 1859 [nomen dubium], Trionyx borkenensis Gramann 1956 [nomen dubium], Trionyx bowerbanki Lydekker 1889 [nomen dubium], Trionyx bruxelliensis Winkler 1869 [nomen dubium], Trionyx burdigalensis Bergounioux 1935 [nomen dubium], Trionyx capellinii bulgaricus Khosatzky et al. 1983 [nomen dubium], Trionyx chaubeti Bergounioux 1935 [nomen dubium], Trionyx ciryi Bergounioux 1935 [nomen dubium], Trionyx cliftii Fitzinger 1835 [nomen nudum], Trionyx croaticus Koch 1915 [nomen dubium], Trionyx danovi Chkhikvadze 1989 [nomen dubium], Trionyx dodunii Gray 1831 [nomen nudum], Trionyx elongatus Liebus 1930 [nomen dubium], Trionyx erquelinnensis Dollo 1909 [nomen nudum], Trionyx hilberi Hoernes 1892 [nomen dubium], Trionyx ishiharaensis Miura and Uyama 1987 [nomen dubium], Trionyx italicus Schauroth 1865 [nomen dubium], Trionyx jakhimovitchae Chkhikvadze 1989 [nomen dubium], Trionyx kazusensis Otsuka 1969 [nomen dubium], Trionyx khosatzkyi Chkhikvadze 1983 [nomen dubium], Trionyx laurillardii Gray 1831 [nomen dubium], Trionyx lockardi Gray 1831 [nomen nudum], Trionyx lorioli Portis 1882 [nomen dubium], Trionyx manouri Gray 1831 [nomen dubium], Trionyx marini Hernández Sampelayo and Bataller 1944 [nomen dubium], Trionyx moldaviensis Khosatzky 1986 [nomen dubium], Trionyx mourieri Bergounioux 1935 [nomen dubium], Trionyx munzenbergensis Hummel 1927 [nomen nudum], Trionyx oligocenica Negri 1892 [nomen nudum], Trionyx oweni Reinach 1900 [nomen dubium], Trionyx parisiensis Gray 1831 [nomen dubium], Trionyx partschii Peters 1855 [nomen dubium], Trionyx pedemontana Portis 1879 [nomen dubium], Trionyx petersi trifailensis Teppner 1914 [nomen dubium], Trionyx pliocaenicus Reinach 1903 [nomen dubium], Trionyx pompignanensis Bergounioux 1933 [nomen dubium], Trionyx pontanus Laube 1895 [nomen dubium], Trionyx portisi Ristori 1891 [nomen dubium], Trionyx preschenensis Laube 1900 [nomen dubium], Trionyx propinquus Ristori 1891 [nomen dubium], Trionyx pseudovindobonensis Szalai 1934 [nomen dubium], Trionyx pustulatus Owen and Bell 1849 [nomen dubium], Trionyx ragusensis De Gregorio 1883 [nomen dubium], Trionyx reticulatus Rieppel 1979 [nomen dubium], Trionyx rocchettiana Portis 1882 [nomen dubium], Trionyx roncensis Harrassowitz 1919 [nomen dubium], Trionyx rotundiformis Bergounioux 1933 [nomen dubium], Trionyx sculptus Gilmore 1931 [nomen dubium], Trionyx senckenbergianus Reinach 1903 [nomen dubium], Trionyx senensis Ristori 1891 [nomen dubium], Trionyx siegeri Heritsch 1909 [nomen dubium], Trionyx sophiae Heritsch 1909 [nomen dubium], Trionyx stormsi Delheid 1899 [nomen dubium], Trionyx ubeensis Chitani 1925 [nomen dubium], Trionyx valdensis Portis 1882 [nomen dubium], Trionyx zakhidovi Khosatzky 1966 [nomen dubium], Zaisanonyx Chkhikvadze 2008 [nomen dubium]
Superfm. Trionychoidea Fitzinger 1826
+
Fm. †Adocidae Cope 1869
+
G. †Adocoides Sukhanov and Narmandakh 2006
G. †Adocus Cope 1868
+
Invalid names: Alamosemys Hay 1908 [synonym], Homorophus Cope 1869 [synonym], Shineusemys Sukhanov and Narmandakh 2006 [synonym], Zygoramma Cope 1869 [synonym]
G. †Ferganemys Nessov and Yulinen 1977
G. †Isanemys Tong et al. 2006
Subfm. †Shachemydinae Khosatzky 1977
G. †Shachemys Kuznetsov 1976
G. †Shineusemys Sukhanov and Narmandakh 2006
Invalid names: Adocinae Cope 1869 [empty], Adocoides Narmandakh 1985 [nomen nudum], Adocus askari Nesov and Krasovskaya 1984 [nomen dubium]
G. †Basilochelys Tong et al. 2009
+
†Basilochelys macrobios Tong et al. 2009
Fm. Kinosternoidea Hay 1892 [mud turtle]
+
G. Aromochelys Gray
G. Cinosternum von Spix 1824 [mud turtle]
Fm. Kinosternidae Hay 1892
Unr. †Pandermatemys Joyce et al. 2004
G. Sternotherus Gray 1825 [musk turtle]
Invalid names: Aromochelys Gray [valid], Cinosternum von Spix 1824 [valid], Kinosternidae Hay 1892 [valid], Pandermatemys Joyce et al. 2004 [valid], Sternotherus Gray 1825 [valid]
G. †Sinaspideretes Young and Chow 1953
+
†Sinaspideretes wimani Young and Chow 1953
Invalid names: Yehguia Danilov and Parham 2006 [synonym]
Invalid names: Peltochelyidae Seeley 1880 [empty], Planetochelyidae Hutchison 2013 [empty]
Invalid names: Adocusia Danilov and Parham 2006 [empty], Carettochelyoidea Romer 1956 [empty], Charitonyx tajanikolaevae Chkhikvadze 1980 [nomen dubium], Chersidae Zittel 1890 [empty], Cryptoderinea Vaillant 1894 [empty], Dermatochelydidae [empty], Glyptopidae [empty], Glyptopsidae Marsh 1890 [empty], Indochelyidae Datta et al. 2000 [empty], Kallokibotioninae Nopsca 1923 [empty], Kayentachelyidae Gaffney et al. 1987 [empty], Kinosternia Gaffney and Meylan 1988 [empty], Kinosternoidae Holroyd and Hutchison 2002 [empty], Meiolaniidae Lydekker 1887 [empty], Pantrionychia Joyce et al. 2004 [empty], Platysternoidea Baur 1893 [empty], Tritosternidae Lydekker 1889 [empty]
G. †Entoplastrites Huene 1956
+
Fm. †Haichemydidae Sukhanov and Narmandakh 2006
+
G. †Haichemys Sukhanov and Narmandakh 2006
+
†Haichemys ulensis Sukhanov and Narmandakh 2006
G. †Larachelus Pérez-García and Murelaga 2012
+
†Larachelus morla Pérez-García and Murelaga 2012
G. †Odontochelys Li et al. 2008
+
†Odontochelys semitestacea Li et al. 2008
Unr. Pancryptodira Joyce et al. 2004
+
Unr. †Macrobaenidae Sukhanov 1964
+
G. †Anatolemys Khozatsky and Nessov 1979
+
†Anatolemys maximus Khozatskii and Nessov 1979
†Anatolemys oxensis Khozatskii and Nessov 1977
G. †Appalachemys Gentry et al. 2023
+
†Appalachemys ebersolei Gentry et al. 2023
G. †Asiachelys Sukhanov and Narmandakh 2006
+
Invalid names: Asiachelys perforata Sukhanov and Narmandakh 2000 [nomen nudum]
G. †Aurorachelys Vandermark et al. 2009
+
†Aurorachelys gaffneyi Vandermark et al. 2009
G. †Changmachelys Brinkman et al. 2013
+
†Changmachelys bohlini Brinkman et al. 2013
G. †Gallica Pérez-García 2020
+
†Gallica lapparentiana Pérez-García 2020
G. †Hangaiemys Sukhanov and Narmandakh 1974
+
†Hangaiemys acutus Bohlin 1953
†Hangaiemys hoburensis Sukhanov and Narmandakh 1974
†Hangaiemys kansuensis Bohlin 1953
†Hangaiemys latilimbata Bohlin 1953
†Hangaiemys leptis Sukhanov and Narmandakh 2006
Invalid names: Hangaiemys leptis Sukhanov and Narmandakh 2000 [nomen nudum]
G. †Judithemys Parham and Hutchison 2003
+
†Judithemys sukhanovi Parham and Hutchison 2003
G. †Kirgizemys Nessov and Khozatskiy 1973
+
†Kirgizemys dmitrievi Nessov and Khozatskii 1981
†Kirgizemys exaratus Nessov and Khozatskiy 1973
G. †Macrobaena Tatarinov 1959
+
†Macrobaena mongolica Tatarinov 1959
G. †Osteopygis Cope 1868
+
†Osteopygis backmani Russell 1934
†Osteopygis emarginata Cope 1868
+
Invalid names: Amblypeza entellus Hay 1908 [synonym], Catapleura chelydrina Cope 1875 [synonym], Osteopygis borealis Wieland 1904 [synonym], Osteopygis chelydrinus Cope 1868 [synonym], Osteopygis erosus Cope 1875 [synonym], Osteopygis gibbi Wieland 1904 [synonym], Osteopygis platylomus Cope 1870 [synonym], Osteopygis robustus Hay 1908 [synonym]
†Osteopygis kranzi Weems 2014
†Osteopygis russelli Brinkman 2015
G. †Oxemys Nessov 1977
+
†Oxemys gutta Nessov 1977
G. †Yakemys Tong et al. 2021
+
†Yakemys multiporcata Tong et al. 2021
Invalid names: Asiachelys Sukhanov 2000 [nomen nudum], Batoremys Narmandakh 2000 [nomen nudum]
G. †Owadowia Szczygielski et al. 2017
+
†Owadowia borsukbialynickae Szczygielski et al. 2017
Fm. †Thalassemydidae Zittel 1889
+
G. †Changisaurus Young 1959
+
†Changisaurus wutuensis Young 1961
G. †Phyllemys Schmidt 1944
G. †Sontiochelys Stache 1905
+
†Sontiochelys cretacea Stache 1905
G. †Yaxartemys Riabinin 1948
+
†Yaxartemys longicauda Riabinin 1948
Invalid names: Acichelyidae Lydekker 1889 [synonym], Emys beaugrandi Sauvage 1872 [nomen dubium], Trionyx primoevus Bergounioux 1937 [nomen dubium]
Unr. †Panpleurodira Danilov and Parham 2008
+
G. †Caribemys Fuente and Iturralde-Vinet 2001
Unr. †Dortokidae Lapparent de Broin and Murelaga 1996
+
G. †Dortoka Lapparent de Broin and Murelaga 1996
+
†Dortoka botanica Gheerbrant et al. 1999
†Dortoka vasconica Lapparent de Broin and Murelaga 1996
†Dortoka vremiri Augustin et al. 2021
Invalid names: Ronella Gheerbrant et al. 1999 [synonym]
G. †Eodortoka Pérez-García et al. 2014
+
†Eodortoka morellana Pérez-García et al. 2014
Invalid names: Muehlbachia Vremir and Codrea 2009 [nomen nudum], Muehlbachia nopcsai Vremir and Codrea 2009 [nomen nudum]
G. Pelomedusa Wagler 1830 [African helmeted turtle]
+
†Pelomedusa senutpickfordina de Lapparent de Broin 2008
Pelomedusa subrufa Bonaterre 1789 [Helmeted turtle]
+
G. †Notoemys Cattoi and Freiberg 1961
+
†Notoemys laticentralis Cattoi and Freiberg 1961
†Notoemys oxfordiensis de la Fuente and Iturralde-Vinent 2001
†Notoemys tlaxiacoensis Lopez-Conde et al. 2017
†Notoemys zapatocaensis Cadena Rueda and Gaffney 2005
G. †Platychelys Wagner 1853
+
†Platychelys oberndorferi Wagner 1853
Invalid names: Helemys Rutimeyer 1859 [nomen nudum], Helemys serrata Rutimeyer 1859 [nomen nudum], Platychelys courrenti Bergounioux 1935 [nomen nudum]
Subor. Pleurodira Cope 1865 [sideneck turtle]
+
Unr. †Bothremydidae Baur 1891
+
G. †Arenila Lapparent de Broin and Werner 1998
+
†Arenila krebsi Lapparent de Broin and Werner 1998
G. †Atolchelys Romano et al. 2014
+
†Atolchelys lepida Romano et al. 2014
G. †Azabbaremys Gaffney et al. 2001
+
†Azabbaremys moragjonesi Gaffney et al. 2001
Subfm. †Bothremydinae Gaffney 1975
+
G. †Paiutemys Joyce et al. 2016
Invalid names: Cearachelyodda Gaffney et al. 2006 [empty]
Tr. †Bothremydini Gaffney et al. 2006
+
Subtr. †Bothremydina Gaffney et al. 2006
Subtr. †Foxemydina Gaffney et al. 2006
G. †Khargachelys AbdelGawad et al. 2023
G. †Palauchelys López-Conde et al. 2021
G. †Puentemys Cadena et al. 2012
Unr. †Bothremydodda Gaffney et al. 2006
+
G. †Abalakemys Pérez-García 2023
G. †Carteremys Williams 1953
+
†Carteremys leithii Carter 1852
Tr. †Cearachelyini Gaffney et al. 2006
+
G. †Elkanemys Maniel et al. 2021
G. †Galianemys Gaffney et al. 2002
G. †Cearachelys Gaffney et al. 2001
+
†Cearachelys placidoi Gaffney et al. 2001
G. †Chedighaii Gaffney et al. 2006
+
†Chedighaii barberi Schmidt 1940
+
Invalid names: Podocnemis alabamae Zangerl 1948 [synonym]
†Chedighaii hutchisoni Gaffney et al. 2006
G. †Elochelys Nopsca 1931
+
†Elochelys perfecta Nopsca 1931
G. †Inaechelys de Araujo Carvalho et al. 2016
Tr. †Kurmademydini Gaffney et al. 2006
+
G. †Jainemys Joyce and Bandyopadhyay 2020
G. †Kinkonychelys Gaffney et al. 2009
G. †Kurmademys Gaffney et al. 2001
+
†Kurmademys kallamedensis Gaffney et al. 2001
G. †Labrostochelys Gaffney et al. 2006
+
†Labrostochelys galkini Gaffney et al. 2006
G. †Nigeremys Broin 1977
+
†Nigeremys gigantea Bergounoux and Crouzel 1968
Invalid names: Potamochelys Fitzinger 1843 [synonym]
G. †Palemys Gray 1870
+
†Palemys bowerbankii Owen 1842
+
Invalid names: Dacochelys delabechei Bell 1849 [synonym], Emys conybearii Owen 1858 [synonym], Emys delabechii Owen and Bell 1849 [synonym], Emys laevis Owen and Bell 1849 [synonym]
Invalid names: Dacochelys Lydekker 1889 [synonym]
G. †Phosphatochelys Gaffney and Tong 2003
+
†Phosphatochelys tedfordi Gaffney and Tong 2003
G. †Pleurochayah Adrian et al. 2021
+
†Pleurochayah appalachius Adrian et al. 2021
G. †Rhothonemys Gaffney et al. 2006
+
†Rhothonemys brinkmani Gaffney et al. 2006
G. †Rosasia Costa 1940
+
†Rosasia soutoi Carrington da Costa 1940
Invalid names: Inaechelys pernambucensis de Araujo Carvalho et al. 2016 [nomen dubium]
G. †Sankuchemys Gaffney et al. 2003
+
†Sankuchemys sethnai Gaffney et al. 2003
G. †Sindhochelys de Lapparent de Broin et al. 2021
+
†Sindhochelys ragei de Lapparent de Broin et al. 2021
Tr. †Taphrosphyini Gaffney et al. 2006
+
G. †Chupacabrachelys Lehman and Wick 2010
G. †Ilatardia Pérez-García 2019
Subtr. †Taphrosphyina Gaffney et al. 2006
Invalid names: Crassachelys Moody 1972 [nomen dubium], Eusarkia Bergounioux 1952 [nomen dubium], Gafsachelys Stefano 1903 [nomen dubium], Nigeremydina Gaffney et al. 2006 [empty]
G. †Taphrosphys Cope 1869
+
†Taphrosphys ippolitoi Gaffney et al. 2006
†Taphrosphys nodosus Cope 1870
†Taphrosphys strenuus Cope 1870
+
Invalid names: Prochonias princeps Cope 1869 [synonym]
†Taphrosphys sulcatus Leidy 1856
+
Invalid names: Amblypeza antellius Hay 1908 [synonym], Prochonias longinuchus Cope 1870 [synonym], Taphrosphys enodis Cope 1870 [synonym], Taphrosphys leslianus Cope 1870 [synonym], Taphrosphys molops Cope 1870 [synonym]
Invalid names: Amblypeza Hay 1908 [synonym], Bantuchelys Dollo 1924 [synonym], Prochonias Cope 1869 [synonym]
G. †Ummulisani Gaffney et al. 2006
+
†Ummulisani rutgersensis Gaffney et al. 2006
Invalid names: Kurmademydinae Gaffney et al. 2006 [empty], Podocnemis somaliensis Walker 1966 [invalid subgroup]
Fm. Chelidae Gray 1825
+
G. Acanthochelys Gray 1873
+
†Acanthochelys cosquinensis de la Fuente 1992
Acanthochelys macrocephala Rhodin et al. 1984 [Pantanal swamp turtle]
Acanthochelys pallidipectoris Freiberg 1945 [Chaco sideneck turtle]
Acanthochelys radiolata Mikan 1820 [Radiolated Brazilian swamp turtle]
Acanthochelys spixii Duméril and Bibron 1837 [black spine-neck swamp turtle]
G. †Baalemys Sarda and Maniel 2024
+
†Baalemys mansillai Sarda and Maniel 2024
G. †Birlimarr Megirian and Murray 1999
+
†Birlimarr gaffneyi Megirian and Murray 1999
Subfm. Chelinae Gray 1825
+
G. Mesoclemmys Gray 1873
+
Invalid names: Batrachemys Stejneger 1909 [synonym], Bufocephala McCord et al. 2001 [synonym], Ranacephala McCord et al. 2001 [synonym]
G. Rhinemys Wagler 1830
G. Chelodina Fitzinger 1826
+
Subg. Chelodina (Chelodina) Fitzinger 1826
Subg. Chelodina (Macrochelodina) Wells and Wellington 1985
Subg. Chelodina (Macrodiremys) McCord and Joseph-Ouni 2007
†Chelodina alanrixi Lapparent de Broin and Molnar 2001
Chelodina expansa Gray 1857 [Giant snakeneck turtle]
†Chelodina insculpta De Vis 1897
Chelodina mccordi Rhodin 1994 [McCord's snakeneck turtle]
†Chelodina murrayi Yates 2013
Chelodina novaeguinae Boulenger 1888 [New Guinea snakeneck turtle]
Chelodina oblonga Gray 1841 [Narrow-breasted snakeneck turtle]
+
Invalid names: Chelodina colliei Gray 1856 [synonym]
Chelodina parkeri Rhodin and Mittermeter 1976 [Parker's snakeneck turtle]
Chelodina pritchardi Rhodin 1994 [Pritchard's snakeneck turtle]
Chelodina reimanni Philippen and Grossmann 1990 [Reimann's snakeneck turtle]
Chelodina rugosa Ogilby 1890 [Northern Australian snakeneck turtle]
Chelodina siebenrocki Werner 1901 [Siebenrock's snakeneck turtle]
Chelodina steindachneri Siebenrock 1914 [Steindachner's snakeneck turtle]
Subfm. Chelodininae Baur 1893
+
G. Elusor Cann and Legler 1994
G. Flaviemys Le et al. 2013 [Manning River sawshelled turtle]
G. Chelus Duméril 1806
+
†Chelus colombianus Wood 1976 [Chelus colombiana]
+
Invalid names: Chelus lewisi Wood 1976 [synonym]
Chelus fimbriata Schneider 1783 [Mata mata]
Chelus fimbriatus Schneider 1783 [Mata mata]
G. Elseya Gray 1867
+
Elseya albagula Thomson et al. 2006 [White-throated snapping turtle]
Elseya branderhorsti Ouwens 1914 [White-bellied snapping turtle]
Elseya dentata Gray 1863 [Northern Australia snapping turtle]
Elseya irwini Cann 1997 [Irwins snapping turtle]
Elseya macrurus Cann and Legler 1994 [Cann's snakeneck turtle]
†Elseya nadibajagu Thomson and Mackness 1999
Elseya novaeguineae Meyer 1874 [New Guinea snapping turtle]
Elseya oneiros Joseph-Ouni et al. 2020
Elseya schultzei Vogt 1911 [Schultzes snapping turtle]
†Elseya uberrima De Vis 1897
Invalid names: Pelecomastes De Vis 1897 [synonym]
G. Emydura Bonaparte 1836
+
Emydura australis Gray 1841 [Australian bighead turtle]
Emydura krefftii Gray 1871 [Krefft's river turtle]
Emydura macquarii Gray 1831 [Murray River turtle]
Emydura signata Ahl 1932 [Brisbane shortneck turtle]
Emydura subglobosa Krefft 1876 [Redbelly shortneck turtle]
Emydura tanybaraga Cann 1997 [Northern yellow-faced turtle]
Emydura victoriae Gray 1842 [Victoria shortneck turtle]
G. Hydromedusa Wagler 1830
+
†Hydromedusa casamayorensis de la Fuente and Bona 2002
Hydromedusa maximiliani Mikan 1820 [Brazilian snakeneck turtle]
Hydromedusa tectifera Cope 1869 [Argentine snake-necked turtle]
+
Invalid names: Platemys antiqua Ameghino 1882 [synonym], Platemys fossilis Ameghino 1882 [synonym], Platemys laevis Ameghino 1882 [synonym], Platemys robusta Ameghino 1882 [synonym]
Subfm. Hydromedusinae Baur 1893
+
G. †Iaremys Agnolín et al. 2024
G. †Mendozachelys de la Fuente et al. 2017
+
†Mendozachelys wichmanni de la Fuente et al. 2017
G. Myuchelys Thomson and Georges 2009
+
Myuchelys bellii Gray 1844
Myuchelys georgesi Georges and Adams 1992
Myuchelys latisternum Gray 1867 [Serrated snapping turtle]
Myuchelys purvisi Wells and Wellington 1985
G. Phrynops Wagler 1830
+
Phrynops geoffroanus Schweigger 1812 [side-necked turtle of Geoffroy]
Phrynops hilarii Dumeril and Bibron 1835 [side-necked turtle of Saint Hillaire]
†Phrynops paranaensis Wieland 1923
Phrynops tuberosus Peters 1870 [Guianan Shield side-necked turtle]
Phrynops williamsi Rhodin and Mittermeier 1983 [Williams side-necked turtle]
Phrynops zuliae Pritchard and Trebbau 1984 [Zulia toadhead turtle]
Invalid names: Parahydraspis Wieland 1923 [synonym]
G. Platemys Wagner 1830
+
Platemys platycephala Schneider 1792 [Twistneck turtle]
G. †Prochelidella Lapparent de Broin and de la Fuente 2001
+
†Prochelidella argentinae Lapparent de Broin and de la Fuente 2001
†Prochelidella buitreraensis Maniel et al. 2020
†Prochelidella cerrobarcinae de la Fuente et al. 2011
†Prochelidella palomoi Maniel et al. 2022
†Prochelidella portezuelae de la Fuente 2003
G. Pseudemydura Siebenrock 1901
+
Pseudemydura umbrina Siebenrock 1901 [Western swamp turtle]
G. Rheodytes Legler and Cann 1980
+
†Rheodytes devisi Thomson 2000
Rheodytes leukops Legler and Cann 1980 [Fitzroy River turtle]
G. †Rionegrochelys De la Fuente et al. 2017
+
†Rionegrochelys caldieroi De la Fuente et al. 2017
G. †Yaminuechelys de la Fuente et al. 2001
+
†Yaminuechelys gasparinii de la Fuente et al. 2001
†Yaminuechelys maior Staesche 1929
+
Invalid names: Osteopygis sculptus Staesche 1929 [synonym]
†Yaminuechelys sulcipeculiaris Oriozabala et al. 2020
Invalid names: Chelydidae Gray 1869 [synonym], Chelymys antiqua De Vis 1897 [nomen dubium], Chelymys arata De Vis 1897 [nomen dubium], Hydraspididae Cope 1868 [synonym], Hydraspis arenarius Rusconi 1934 [nomen dubium], Naiadochelys patagonica Staesche 1929 [nomen dubium], Pelecomastes ampla De Vis 1897 [nomen dubium], Platemys bergii Ambrosetti 1893 [nomen dubium], Platemys entreriana Ambrosetti 1893 [nomen dubium], Platemys holmbergii Ambrosetti 1893 [nomen dubium], Platemys mesopotamica Ambrosetti 1893 [nomen dubium], Platemys robusta Ambrosetti 1893 [nomen dubium], Platemys torrentium Burmeister 1885 [nomen dubium], Trionyx argentina Ameghino 1889 [nomen dubium]
Unr. Eupleurodira Gaffney and Meylan 1988
+
Invalid names: Bothremydinura Gaffney et al. 2006 [empty], Cheloides Gray 1825 [empty]
Unr. †Panchelidae Joyce et al. 2004
+
G. †Bonapartemys Lapparent de Broin and de la Fuente 2001
+
†Bonapartemys bajobarrealis Lapparent de Broin and de la Fuente 2001
G. †Linderochelys de la Fuente et al. 2007
+
†Linderochelys rinconensis de la Fuente et al. 2007
G. †Lomalatachelys Lapparent de Broin and de la Fuente 2001
+
†Lomalatachelys neuquina Lapparent de Broin and de la Fuente 2001
G. †Palaeophrynops Lapparent de Broin and de la Fuente 2001
+
†Palaeophrynops patagonicus Lapparent de Broin and de la Fuente 2001
G. †Salamanchelys Bona 2006
+
Invalid names: Platemys guaranitica Ameghino 1889 [nomen dubium], Platemys patagonica Ameghino 1889 [nomen dubium]
Unr. †Pelomedusoides Broin 1988
+
G. †Apodichelys Price 1954
+
†Apodichelys lucianoi Price 1954
Fm. †Araripemydidae Price 1973
+
G. †Araripemys Price 1973
G. †Laganemys Sereno and ElShafie 2013
G. †Taquetochelys Broin 1980
G. †Caririemys Oliveira and Kellner 2007
+
†Caririemys violetae Oliveira and Kellner 2007
G. †Francemys Pérez-García 2019
+
†Francemys gadoufaouaensis Pérez-García 2019
G. †Itapecuruemys Batista et al. 2021
+
†Itapecuruemys amazonensis Batista et al. 2021
G. †Palaeaspis Gray 1870
+
†Palaeaspis bowerbanki Owen 1842
Fm. †Peiropemydidae Gaffney et al. 2011
+
G. †Pricemys Gaffney et al. 2011
G. †Yuraramirim Ferreira et al. 2018
Unr. †Pelomedusera Gaffney et al. 2006
+
Fm. Pelomedusidae Cope 1868
+
Invalid names: Eusarkiidae Bergounioux 1952 [synonym], Peltocephalidae Gray 1870 [synonym], Sternothaeridae Cope 1868 [synonym]
+
Superfm. †Euraxemydoidea Gaffney et al. 2006
G. †Teneremys Broin 1980
Invalid names: Platycheloides nyasae Haughton 1928 [nomen dubium]
Superfm. Podocnemidoidae Cope 1868
+
G. †Amabilis Hermanson et al. 2020
G. †Roxochelys Price 1953
Invalid names: Brasilemydidae Lapparent de Broin 2000 [empty]
Superfm. †Podocnemidoidea Gaffney et al. 2006
+
G. †Nostimochelone Georgalis et al. 2013
Fm. Podocnemididae Cope 1868
+
G. †Albertwoodemys Gaffney et al. 2011
+
†Albertwoodemys testudinum Gaffney et al. 2011
G. †Bairdemys Gaffney and Wood 2002
+
†Bairdemys hartsteini Gaffney and Wood 2002
†Bairdemys healeyorum Weems and Knight 2013
†Bairdemys miocenica Collins and Lynn 1936
†Bairdemys sanchezi Gaffney et al. 2008
†Bairdemys thalassica Ferreira et al. 2015
†Bairdemys venezuelensis Wood and Diaz 1971
†Bairdemys winklerae Gaffney et al. 2008
G. †Bauruemys Suarez 1969
+
†Bauruemys elegans Suarez 1969
Invalid names: Bauruemys brasiliensis Staesche 1937 [nomen dubium]
G. †Brontochelys Gaffney et al. 2011
+
†Brontochelys gaffneyi Wood 1970
G. †Cambaremys França and Langer 2005
G. †Carbonemys Cadena et al. 2012
+
†Carbonemys cofrinii Cadena et al. 2012
G. †Cerrejonemys Cadena et al. 2010
+
†Cerrejonemys wayuunaiki Cadena et al. 2010
G. †Cordichelys Gaffney et al. 2011
+
†Cordichelys antiqua Andrews 1903
G. †Dacquemys Williams 1954
+
†Dacquemys paleomorpha Williams 1954
Subfm. †Erymnochelyinae Broin 1988
+
G. †Andrewsemys Pérez-García 2019
G. †Caninemys Meylan et al. 2009
G. †Eocenochelus Pérez-García et al. 2017
Tr. †Erymnochelyini de Broin 1988
G. †Neochelys Bergounioux 1954
G. †Ragechelus de Lapparent de Broin et al. 2020
G. †Stupendemys Wood 1976
G. Erymnochelys Baur 1888
+
G. †Gestemys de la Fuente et al. 2022
+
†Gestemys powelli de la Fuente et al. 2022
G. †Kenyemys Wood 1983
+
†Kenyemys williamsi Wood 1983
G. †Lapparentemys Gaffney et al. 2011
+
†Lapparentemys vilavilensis de Broin 1971
G. †Latentemys Gaffney et al. 2011
+
†Latentemys plowdeni Gaffney et al. 2011
G. †Lemurchelys Gaffney et al. 2011
+
†Lemurchelys diasphax Gaffney et al. 2011
G. †Mogharemys Gaffney et al. 2011
+
†Mogharemys blanckenhorni Dacque 1912
G. †Papoulemys Tong 1998
G. †Peiropemys Gaffney et al. 2011
+
†Peiropemys mezzalirai Gaffney et al. 2011
G. Peltocephalus Duméril and Bibron 1835
+
Peltocephalus dumerilianus Schweigger 1812 [Bighead Amazon River turtle]
†Peltocephalus maturin Ferreira et al. 2024
Subfm. Podocnemidinae Cope 1868
+
Unr. †Podocnemidodda Gaffney et al. 2011
G. Podocnemis Wagler 1830 [river turtle]
G. †Shweboemys Swinton 1939
+
†Shweboemys pilgrimi Swinton 1939
Subtr. †Stereogenyina Gaffney et al. 2011
+
Invalid names: Bairdemydita Gaffney et al. 2011 [empty]
Tr. †Stereogenyini Gaffney et al. 2011
+
Unr. †Stereogenyita Gaffney et al. 2011
Invalid names: Mogharemydina Gaffney et al. 2011 [empty]
G. †Stereogenys Andrews 1901
+
†Stereogenys cromeri Andrews 1901
Invalid names: Stereogenys podocnemoides Reinach 1903 [nomen dubium]
G. †Turkanemys Wood 2003
+
†Turkanemys pattersoni Wood 2003
Invalid names: Albertwoodemys Gaffney et al. 2011 [valid], Bairdemys Gaffney and Wood 2002 [valid], Bauruemydinae Gaffney et al. 2011 [empty], Bauruemydinae Gaffney et al. 2011 [empty], Bauruemys Suarez 1969 [valid], Brontochelys Gaffney et al. 2011 [valid], Cambaremys França and Langer 2005 [valid], Carbonemys Cadena et al. 2012 [valid], Cerrejonemys Cadena et al. 2010 [valid], Cordichelys Gaffney et al. 2011 [valid], Dacquemys Williams 1954 [valid], Erymnochelyinae Broin 1988 [valid], Erymnochelys Baur 1888 [valid], Gestemys de la Fuente et al. 2022 [valid], Kenyemys Wood 1983 [valid], Lapparentemys Gaffney et al. 2011 [valid], Latentemys Gaffney et al. 2011 [valid], Lemurchelys Gaffney et al. 2011 [valid], Mogharemys Gaffney et al. 2011 [valid], Papoulemys Tong 1998 [valid], Peiropemys Gaffney et al. 2011 [valid], Peltocephalus Duméril and Bibron 1835 [valid], Podocnemidinae Cope 1868 [valid], Shweboemys Swinton 1939 [valid], Stereogenyina Gaffney et al. 2011 [valid], Stereogenyini Gaffney et al. 2011 [valid], Stereogenys Andrews 1901 [valid], Turkanemys Wood 2003 [valid]
Unr. †Podocnemidinura Gaffney et al. 2006
Unr. †Sahonachelyidae Joyce et al. 2021
+
G. †Sahonachelys Joyce et al. 2021
G. †Sokatra Gaffney and Krause 2011
Invalid names: Chrysemys montolivensis Roman 1897 [invalid subgroup], Naiadochelys Hay 1908 [nomen dubium], Panpodocnemidae Joyce et al. 2004 [empty], Taphrosphys dares Hay 1908 [nomen dubium]
Fm. Podocnemididae Cope 1868
+
G. †Albertwoodemys Gaffney et al. 2011
+
†Albertwoodemys testudinum Gaffney et al. 2011
G. †Bairdemys Gaffney and Wood 2002
+
†Bairdemys hartsteini Gaffney and Wood 2002
†Bairdemys healeyorum Weems and Knight 2013
†Bairdemys miocenica Collins and Lynn 1936
†Bairdemys sanchezi Gaffney et al. 2008
†Bairdemys thalassica Ferreira et al. 2015
†Bairdemys venezuelensis Wood and Diaz 1971
†Bairdemys winklerae Gaffney et al. 2008
G. †Bauruemys Suarez 1969
+
†Bauruemys elegans Suarez 1969
Invalid names: Bauruemys brasiliensis Staesche 1937 [nomen dubium]
G. †Brontochelys Gaffney et al. 2011
+
†Brontochelys gaffneyi Wood 1970
G. †Cambaremys França and Langer 2005
G. †Carbonemys Cadena et al. 2012
+
†Carbonemys cofrinii Cadena et al. 2012
G. †Cerrejonemys Cadena et al. 2010
+
†Cerrejonemys wayuunaiki Cadena et al. 2010
G. †Cordichelys Gaffney et al. 2011
+
†Cordichelys antiqua Andrews 1903
G. †Dacquemys Williams 1954
+
†Dacquemys paleomorpha Williams 1954
Subfm. †Erymnochelyinae Broin 1988
+
G. †Andrewsemys Pérez-García 2019
G. †Caninemys Meylan et al. 2009
G. †Eocenochelus Pérez-García et al. 2017
Tr. †Erymnochelyini de Broin 1988
G. †Neochelys Bergounioux 1954
G. †Ragechelus de Lapparent de Broin et al. 2020
G. †Stupendemys Wood 1976
G. Erymnochelys Baur 1888
+
G. †Gestemys de la Fuente et al. 2022
+
†Gestemys powelli de la Fuente et al. 2022
G. †Kenyemys Wood 1983
+
†Kenyemys williamsi Wood 1983
G. †Lapparentemys Gaffney et al. 2011
+
†Lapparentemys vilavilensis de Broin 1971
G. †Latentemys Gaffney et al. 2011
+
†Latentemys plowdeni Gaffney et al. 2011
G. †Lemurchelys Gaffney et al. 2011
+
†Lemurchelys diasphax Gaffney et al. 2011
G. †Mogharemys Gaffney et al. 2011
+
†Mogharemys blanckenhorni Dacque 1912
G. †Papoulemys Tong 1998
G. †Peiropemys Gaffney et al. 2011
+
†Peiropemys mezzalirai Gaffney et al. 2011
G. Peltocephalus Duméril and Bibron 1835
+
Peltocephalus dumerilianus Schweigger 1812 [Bighead Amazon River turtle]
†Peltocephalus maturin Ferreira et al. 2024
Subfm. Podocnemidinae Cope 1868
+
Unr. †Podocnemidodda Gaffney et al. 2011
G. Podocnemis Wagler 1830 [river turtle]
G. †Shweboemys Swinton 1939
+
†Shweboemys pilgrimi Swinton 1939
Subtr. †Stereogenyina Gaffney et al. 2011
+
Invalid names: Bairdemydita Gaffney et al. 2011 [empty]
Tr. †Stereogenyini Gaffney et al. 2011
+
Unr. †Stereogenyita Gaffney et al. 2011
Invalid names: Mogharemydina Gaffney et al. 2011 [empty]
G. †Stereogenys Andrews 1901
+
†Stereogenys cromeri Andrews 1901
Invalid names: Stereogenys podocnemoides Reinach 1903 [nomen dubium]
G. †Turkanemys Wood 2003
+
†Turkanemys pattersoni Wood 2003
Invalid names: Albertwoodemys Gaffney et al. 2011 [valid], Bairdemys Gaffney and Wood 2002 [valid], Bauruemydinae Gaffney et al. 2011 [empty], Bauruemydinae Gaffney et al. 2011 [empty], Bauruemys Suarez 1969 [valid], Brontochelys Gaffney et al. 2011 [valid], Cambaremys França and Langer 2005 [valid], Carbonemys Cadena et al. 2012 [valid], Cerrejonemys Cadena et al. 2010 [valid], Cordichelys Gaffney et al. 2011 [valid], Dacquemys Williams 1954 [valid], Erymnochelyinae Broin 1988 [valid], Erymnochelys Baur 1888 [valid], Gestemys de la Fuente et al. 2022 [valid], Kenyemys Wood 1983 [valid], Lapparentemys Gaffney et al. 2011 [valid], Latentemys Gaffney et al. 2011 [valid], Lemurchelys Gaffney et al. 2011 [valid], Mogharemys Gaffney et al. 2011 [valid], Papoulemys Tong 1998 [valid], Peiropemys Gaffney et al. 2011 [valid], Peltocephalus Duméril and Bibron 1835 [valid], Podocnemidinae Cope 1868 [valid], Shweboemys Swinton 1939 [valid], Stereogenyina Gaffney et al. 2011 [valid], Stereogenyini Gaffney et al. 2011 [valid], Stereogenys Andrews 1901 [valid], Turkanemys Wood 2003 [valid]
Invalid names: Amphichelydidae Hay [empty], Megapleurodira Gaffney et al. 2006 [empty], Minipleurodira Gaffney et al. 2006 [empty], Notoemydidae Fernandez and de la Fuente 1994 [empty], Panpelomedusoides Joyce et al. 2004 [empty], Platychelira Gaffney et al. 2006 [empty]
Subor. †Proganochelydia
+
+
Invalid names: Triassochelyidae Abel 1919 [synonym]
G. †Tacuarembemys Perea et al. 2014
+
†Tacuarembemys kusterae Perea et al. 2014
G. †Triassochelys Jaekel 1918
+
Invalid names: Stegochelys Jaekel 1914 [replaced]
Invalid names: Apertotemporalis Stromer 1934 [nomen dubium], Apholidemys Pomel 1847 [nomen dubium], Casichelydia Gaffney 1975 [empty], Chelone obovata Owen 1842 [nomen dubium], Chelone sopita Leidy 1865 [nomen vanum], Chelonia Latreille 1800 [synonym], Chelonii Latreille 1800 [synonym], Cryptrodiramorpha Lee 1995 [empty], Euclastochelys Bergounioux 1955 [nomen dubium], Euclastochelys interrupta Bergounioux 1955 [nomen dubium], Meiolania damelipi White et al. 2010 [nomen dubium], Platychelone Dollo 1909 [nomen nudum], Platychelone emarginata Dollo 1909 [nomen nudum], Plesiochelydiae [empty], Podocnemis lata Ristori 1895 [invalid subgroup], Senonemys suemegensis Bohn 1966 [nomen dubium], Sokotochelys Halstead 1979 [nomen dubium], Temnoclemmys cordiformis Bergounioux 1958 [nomen vanum], Trionyx fuchienensis Yeh 1974 [nomen dubium], Trionyx tshelkarensis Chkhikvadze 1973 [nomen nudum]
G. †Testudinites Weiss 1830
Fm. Testudofiexoolithidae Hirsch 1996
+
G. †Testudofiexoolithus Hirsch 1996
+
Fm. Testudoolithidae Hirsch 1996
+
G. †Testudoolithus Hirsch 1996
+
†Testudoolithus rigidus Hirsch 1996
G. †Tiantaioolithus Fang et al. 2003
+
†Tiantaioolithus jiangi Fang et al. 2003
Unr. †Thalassochelydia Anquetin et al. 2017
+
G. †Achelonia von Meyer 1860
+
†Achelonia formosa von Meyer 1860
Fm. †Eurysternidae Dollo 1886
+
G. †Chelonides Maack 1869
+
†Chelonides wittei Maack 1869
G. †Hydropelta Meyer 1850
+
†Hydropelta meyeri Tholliere 1851
G. †Idiochelys Meyer 1839
+
†Idiochelys fitzingeri Meyer 1839
+
Invalid names: Chelonemys ovata Jourdan 1862 [synonym], Chelonemys plana Jourdan 1862 [synonym], Idiochelys wagneri Meyer 1846 [synonym]
†Idiochelys wagneri Meyer 1846
G. †Palaeomedusa Meyer 1860
+
†Palaeomedusa testa von Meyer 1860
+
Invalid names: Eurysternum crassipes Wagner 1861 [objective synonym]
G. †Parachelys Meyer 1864
+
†Parachelys eichstaettensis von Meyer 1864
Invalid names: Plesiochelys minima Oertel 1915 [nomen dubium]
G. †Eurysternum Meyer 1839
+
†Eurysternum neuquinum Fernandez and de la Fuente 1988
†Eurysternum wagleri Meyer 1839
G. †Hylaeochelys Lydekker 1889
+
†Hylaeochelys belli Mantell 1844
+
Invalid names: Archaeochelys valdensis Lydekker 1889 [synonym], Hylaeochelys sollasi Nopcsa 1928 [synonym], Platemys dixoni Owen 1853 [synonym], Platemys mantelli Owen 1841 [synonym]
†Hylaeochelys kappa Pérez-García and Ortega 2014
+
Invalid names: Pleurosternon koeneni Grabbe 1884 [synonym]
Invalid names: Archaeochelys Lydekker 1889 [synonym]
G. †Jurassichelon Pérez-García 2015
+
†Jurassichelon moseri Bräm 1965
†Jurassichelon oleronensis Pérez-García 2015
G. †Neusticemys Fernandez and de la Fuente 1993
+
†Neusticemys neuquina Fernandez and de la Fuente 1988
Fm. †Plesiochelyidae Rütimeyer 1873
+
G. †Craspedochelys Rütimeyer 1873
+
†Craspedochelys jaccardi Pictet 1860
+
Invalid names: Plesiochelys choffati Sauvage 1898 [synonym]
†Craspedochelys passmorei Andrews 1921
†Craspedochelys picteti Rütimeyer 1873
+
Invalid names: Craspedochelys crassa Rütimeyer 1873 [synonym]
Invalid names: Emys dollfusii Lennier 1870 [nomen dubium]
G. †Globochelus de Lapparent de Broin et al. 2021
+
†Globochelus lennieri de Lapparent de Broin et al. 2021
G. †Plesiochelys Rütimeyer 1873
+
†Plesiochelys bigleri Puntener et al. 2017
†Plesiochelys etalloni Pictet and Humbert 1857
+
Invalid names: Plesiochelys langii Rütimeyer 1873 [synonym], Plesiochelys sanctaeverenae Rütimeyer 1873 [synonym], Plesiochelys solodurensis Rütimeyer 1873 [synonym], Stylemys hannoverana Maack 1869 [synonym], Stylemys lindenensis Maack 1869 [synonym]
†Plesiochelys normandicus Bergounioux 1937
Invalid names: Clemmys grayi Fitzinger 1835 [nomen oblitum], Emys hugi Gray 1831 [nomen oblitum], Emys hugii Gray 1831 [nomen oblitum], Emys jurensis Keferstein 1834 [nomen oblitum], Emys trionychoides Gray 1831 [nomen oblitum], Plesiochelys minor Portis 1878 [nomen dubium], Stegochelys Lydekker 1889 [synonym]
G. †Portlandemys Gaffney 1975
+
†Portlandemys gracilis Anquetin et al. 2015
†Portlandemys mcdowelli Gaffney 1975
G. †Tholemys Andrews 1921
G. †Yanduchelys Peng et al. 2005
+
†Yanduchelys delicatus Peng et al. 2005
Invalid names: Emys dutertrei Sauvage 1872 [nomen dubium], Hispaniachelys Slater et al. 2011 [nomen dubium], Hispaniachelys prebetica Slater et al. 2011 [nomen dubium], Stylemys lindenensis Maack 1869 [nomen dubium]
G. †Solnhofia Gaffney 1975
+
†Solnhofia brachyrhyncha Anquetin and Püntener 2020
†Solnhofia parsonsi Gaffney 1975
G. †Thalassemys Rütimeyer 1859
+
†Thalassemys bruntrutana Puntener et al. 2015
†Thalassemys carva Nessov 1984
†Thalassemys hugii Rütimeyer 1873
+
Invalid names: Eurysternum ignoratum Bräm 1965 [synonym], Thalassemys gresslyi Rütimeyer 1873 [synonym]
Invalid names: Parathalassemys Nessov and Krassevskaya 1984 [synonym]
G. †Tropidemys Rütimeyer 1873
+
†Pelobatochelys blakii Seeley 1875
†Tropidemys langii Rütimeyer 1873
+
Invalid names: Craspedochelys plana Rütimeyer 1873 [synonym], Tropidemys expansa Rütimeyer 1873 [synonym], Tropidemys gibba Rütimeyer 1873 [synonym]
†Tropidemys seebachi Portis 1878
Invalid names: Chelonia valanginiensis Pictet and Campiche 1862 [nomen dubium], Chelonides robusta Portis 1878 [nomen dubium]
Invalid names: Acichelys approximata Wagner 1856 [nomen dubium], Euryaspis radians Wagner 1861 [nomen dubium], Plesiochelys oblonga Oertel 1924 [nomen dubium], Plesiochelys pumilio Oertel 1924 [nomen dubium], Thalassemys heusseri Oertel 1924 [nomen dubium], Tropidemys morinica Sauvage 1880 [nomen dubium]
Subor. Thecophora Dollo
+
Invalid names: Chelonioidae Hay 1930 [empty], Chelydoidae Hay 1930 [empty], Emydoidae Hay 1930 [empty], Pleurosternoidae Hay 1930 [empty], Thalassemydoidae Hay 1930 [empty], Trionychoidae Hay 1930 [empty]
Invalid names: Aplax oberndorferi Meyer 1843 [nomen dubium], Athecae Cope 1871 [empty], Chelonipedidae Sarjeant and Langston 1994 [empty], Chelycarapookus Warren 1969 [nomen dubium], Chelycarapookus arcuatus Warren 1969 [nomen dubium], Chelys blakii Mackie 1863 [nomen dubium], Chengyuchelyidae Yeh 1990 [empty], Emys neumayri Seeley 1881 [nomen dubium], Emys paranensis Burmeister 1885 [nomen nudum], Heckerochelyidae Sukhanov 2006 [empty], Megasternon Gray 1842 [nomen nudum], Mongolochelyidae Sukhanov and Pozdnjakov [empty], Odontochelyidae Li et al. 2008 [empty], Platemys sehuensis Ameghino 1889 [nomen dubium], Protochelys Lydekker 1889 [nomen dubium], Testudo elata Gervais 1877 [nomen dubium], Testudo stricklandi Phillips 1871 [nomen dubium], Tretosternon punctatum Owen 1842 [nomen dubium], Trionyx mammillaris Cope 1877 [nomen nudum]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: phosphaticsubp
Environment: terrestrialuc
Locomotion: actively mobilec
Created: 2017-04-17 15:15:35
Modified: 2017-04-17 15:15:35
Source: c = class, subp = subphylum, uc = unranked clade
References: Carroll 1988, Hendy et al. 2009

Age range

Maximum range based only on fossils: base of the Olenekian to the top of the Holocene or 251.20000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 247.2 Ma

Collections (6236 total)

Oldest occurrences

Time interval Ma Country or state Original ID and collection number
Spathian251.3 - 247.2USA (Wyoming) Chelonipus sp. (135635 135636 135637)
Olenekian - Anisian251.2 - 242.0Germany (Thuringia) Chelonipus cuneiformis, Chelonipus torquatus (182300)
Tuvalian232.0 - 221.5China (Guizhou) Odontochelys semitestacea (85166)
Norian228.0 - 208.5Argentina (La Rioja) Palaeochersis talampayensis (105693)
Norian228.0 - 208.5Germany (Baden-Württemberg) Murrhardtia staeschei (138872)
Norian228.0 - 208.5Argentina (San Juan) Testudinata indet. (182613)
Norian228.0 - 208.5Poland (Opole) Proterochersis porebensis (140077)
Norian228.0 - 208.5Thailand (Khon Kaen) Chelonia indet. (106554) Proganochelys ruchae (106555)
Norian228.0 - 208.5Germany (Baden-Württemberg) Keuperotesta limendorsa, Proterochersis robusta (138886) Proganochelys quenstedti (22683) Proganochelys quenstedtii (138871)
Norian228.0 - 208.5Germany (Bayern) Reptilia indet. (48683)
Norian228.0 - 208.5USA (New Mexico) Chinlechelys tenertesta (48516) Testudines indet. (80926)
Norian228.0 - 208.5Thailand (Phetchabun) Chelonia indet. (106542)
Norian228.0 - 208.5Germany (Baden-Wurttemberg) Proganochelys quenstedtii (68038)
Norian228.0 - 208.5Switzerland (Aargau) Proganochelys quenstedtii (227790)
Norian228.0 - 208.5Germany (Baden-Württemburg) Proterochersis intermedia (138885)
Norian - Rhaetian228.0 - 201.3Poland (Śląskie) Testudinata indet., Proterochersis sp. (132933)
Alaunian215.56 - 212.0Greenland Proganochelys sp. (26997)
Rhaetian208.5 - 201.3Germany (Baden-Württemberg) Testudinata indet. (131422)
Rhaetian208.5 - 201.3Germany (Sachsen-Anhalt) Stegochelys dux, Triassochelys dux (22684)
Rhaetian208.5 - 201.3Italy (Lombardy) Emydidae indet. (38083)