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Scrotifera

Mammalia

Taxonomy
Scrotifera was named by Waddell et al. (1999). It is extant.

It was assigned to Laurasiatheria by Waddell et al. (1999), Asher and Helgen (2010).

Subtaxa
Acreodi, Bulbulodentata, Euungulata (syn. Ungulata, Eparctocyona), Ferae (syn. Ostentoria), Fereuungulata
Sister taxa
Bulbulodentata, Euungulata (syn. Ungulata, Eparctocyona), Ferae (syn. Ostentoria), Fereuungulata

Synonyms
Synonymy list
YearName and author
1999Scrotifera Waddell et al. p. cover
2010Scrotifera Asher and Helgen p. 3 figs. Figure 1

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
—Osteichthyes()
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
—Tetrapoda
—Reptiliomorpha
—Anthracosauria
subclassAmphibiosauriaKuhn 1967
—Cotylosauria()
—Amniota
RankNameAuthor
subclassSynapsida
—Therapsida()
infraorderCynodontia()
—Mammaliamorpha
—Mammaliaformes
classMammalia
—Cladotheria
—Zatheria
subclassTribosphenida()
subclassTheria
—Eutheria()
—Placentalia
—Boreoeutheria
—Laurasiatheria
—Scrotifera
—Scrotifera

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Unr. Scrotifera Waddell et al. 1999
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Or. †Acreodi Matthew 1909
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G. †Andrewsarchus Osborn 1924
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†Andrewsarchus crassum Ding et al. 1977
Unr. †Mesonychia [mesonychian]
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Fm. †Hapalodectidae Szalay and Gould 1966 [condylarth]
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G. †Hapalodectes Matthew 1909
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†Hapalodectes anthracinus Zhou and Gingerich 1991
†Hapalodectes auctus Matthew and Granger 1925
†Hapalodectes compressus Matthew 1909
†Hapalodectes dux Lopatin 2001
†Hapalodectes hetangensis Ting and Li 1987
†Hapalodectes huanghaiensis Tong and Wang 2006
†Hapalodectes leptognathus Osborn and Wortman 1892
†Hapalodectes lopatini Solé et al. 2017
†Hapalodectes paleocenus Beard et al. 2010
†Hapalodectes paradux Lopatin 2023
†Hapalodectes serus Matthew and Granger 1925
G. †Hapalorestes Gunnell and Gingerich 1996
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†Hapalorestes lovei Gunnell and Gingerich 1996
Fm. †Mesonychidae Cope 1875 [condylarth]
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G. †Dissacus Cope 1881
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†Dissacus argenteus O'Leary and Rose 1995
†Dissacus blayaci Stehlin 1926
†Dissacus bohaiensis Tong and Wang 2006
†Dissacus europaeus Lemoine 1891
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Invalid names: Hyaenodictis gaudryi Lemione 1891 [synonym]
†Dissacus filholi Lemoine 1891
†Dissacus indigenus Dashzeveg 1976
†Dissacus magushanensis Yan and Tang 1976
†Dissacus navajovius Cope 1881
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Invalid names: Dissacus carnifex Cope 1882 [synonym]
†Dissacus praenuntius Matthew and Granger 1915
†Dissacus progressus Crusafont-Pairo and Golpe-Posse 1968
†Dissacus raslanloubatieri Solé et al. 2018
†Dissacus rotundus Wang 1975
†Dissacus rougierae Solé et al. 2018
†Dissacus saurognathus Wortman 1897
†Dissacus serior O'Leary and Rose 1995
†Dissacus serratus Chow and Qi 1978
†Dissacus shanghoensis Chow et al. 1973
†Dissacus willwoodensis O'Leary and Rose 1995
†Dissacus zanabazari Geisler and McKenna 2007
†Dissacus zengi Ting 2004
Invalid names: Hyaenodictis Lemoine 1891 [synonym], Hyoenodictis Lemoine 1880 [synonym], Plesidissacus Lemoine 1894 [synonym]
G. †Dissacusium Chow et al. 1977
G. †Harpagolestes Wortman 1901
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†Harpagolestes brevipes Thorpe 1923
†Harpagolestes immanis Matthew 1909
†Harpagolestes koreanicus Shikama 1943
†Harpagolestes leei Jin 2005
†Harpagolestes leotensis Peterson 1931
†Harpagolestes orientalis Szalay and Gould 1966
G. †Hessolestes Peterson 1931
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†Hessolestes ultimus Peterson 1931
G. †Honanodon Chow 1965
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†Honanodon hebetis Chow 1965
†Honanodon macrodontus Chow 1965
G. †Jiangxia Zhang et al. 1979
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†Jiangxia chaotoensis Zhang et al. 1979
G. †Lohoodon Chow et al. 1973
Subfm. †Mesonychinae Cope 1880
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G. †Ankalagon Van Valen 1980
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Invalid names: Ancalagon Van Valen 1978 [synonym]
G. †Guilestes Zheng and Chi 1978
G. †Hukoutherium Chow et al. 1973
G. †Mesonyx Cope 1872
G. †Mongolestes Szalay and Gould 1966
G. †Mongolonyx Szalay and Gould 1966
G. †Yantanglestes Ideker and Yan 1980
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Invalid names: Lestes Yan and Tang 1976 [replaced]
G. †Metahapalodectes Dashzeveg 1976
G. †Olsenia Matthew and Granger 1925
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†Olsenia mira Matthew and Granger 1925
G. †Pachyaena Cope 1874
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†Pachyaena gigantea Osborn and Wortman 1892
†Pachyaena gracilis Matthew and Granger 1915
†Pachyaena nemegetica Dashzeveg 1976
†Pachyaena ossifraga Cope 1874
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Invalid names: Pachyaena intermedia Wortman 1899 [synonym]
G. †Plagiocristodon Chow and Qi 1978
G. †Sinonyx Zhou et al. 1995
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†Sinonyx jiashanensis Zhou et al. 1995
G. †Synoplotherium Cope 1872
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†Synoplotherium canius Cope 1872
†Synoplotherium lanius Cope 1872
†Synoplotherium vorax Marsh 1876
Invalid names: Dromocyon Marsh 1876 [synonym]
Invalid names: Hapalodectinae Szalay and Gould 1966 [empty]
G. †Simidectes Stock 1933
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†Simidectes magnus Peterson 1919
†Simidectes medius Peterson 1919
†Simidectes merriami Stock 1933
Invalid names: Pleurocyon Peterson 1919 [synonym]
G. †Wyolestes Gingerich 1981
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†Wyolestes apheles Gingerich 1981
†Wyolestes dioctes Gingerich 1982
†Wyolestes iglesius Novacek et al. 1991
Invalid names: Andrewsarchidae Szalay and Gould 1966 [empty], Wyolestidae [empty]
G. †Mondegodon Tabuce et al. 2011
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†Mondegodon eutrigonus Tabuce et al. 2011
Fm. †Triisodontidae Scott 1892
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G. †Eoconodon Matthew and Granger 1921
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†Eoconodon copanus Van Valen 1978
†Eoconodon coryphaeus Cope 1885
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Invalid names: Mioclaenus bathygnathus Cope 1888 [synonym], Triisodon biculminatus Cope 1888 [synonym]
†Eoconodon gaudrianus Cope 1888
†Eoconodon ginibitohia Clemens and Williamson 2005
†Eoconodon hutchisoni Clemens 2011
†Eoconodon nidhoggi Van Valen 1978
G. †Goniacodon Scott 1892
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†Goniacodon hiawathae Van Valen 1978
†Goniacodon levisanus Cope 1883
G. †Stelocyon Gingerich 1978
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†Stelocyon arctylos Gingerich 1978
G. †Triisodon Cope 1881
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Invalid names: Triisodon rusticus Cope 1884 [synonym]
†Triisodon quivirensis Cope 1881
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Invalid names: Sarcothraustes antiquus Cope 1882 [synonym], Triisodon conidens Cope 1882 [synonym]
Invalid names: Sarcothraustes Cope 1882 [synonym]
Invalid names: Oxyclaenidae Scott 1892 [invalid subgroup]
Unr. †Bulbulodentata Archibald 1998
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Fm. †Periptychidae Cope 1882 [condylarth]
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G. †Alticonus Eberle and Lillegraven 1998
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†Alticonus gazini Eberle and Lillegraven 1998
G. †Ampliconus Eberle and Lillegraven 1998
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†Ampliconus antoni Eberle and Lillegraven 1998
†Ampliconus browni Eberle and Lillegraven 1998
Subfm. †Anisonchinae Osborne and Earle 1895
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G. †Haploconus Cope 1882
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†Haploconus angustus Cope 1881
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Invalid names: Haploconus corniculatus Cope 1888 [synonym], Haploconus inopinatus Gazin 1939 [synonym], Haploconus xiphodon Cope 1882 [synonym]
†Haploconus elachistus Gazin 1941
†Haploconus entoconus Cope 1882
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Invalid names: Anisonchus coniferus Cope 1882 [synonym]
†Haploconus lineatus Cope 1882
G. †Pseudanisonchus Zhang et al. 1979
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†Pseudanisonchus antelios Zhang et al. 1979
G. †Anisonchus Cope 1881
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†Anisonchus athelas Van Valen 1978
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Invalid names: Anisonchus eowynae Van Valen 1978 [synonym]
†Anisonchus fortunatus Simpson 1932
†Anisonchus oligistus Gazin 1941
†Anisonchus onostus Gazin 1939
†Anisonchus sectorius Cope 1881
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Invalid names: Anisonchus dracus Gazin 1939 [synonym], Mioclaenus mandibularis Cope 1881 [synonym], Zetodon gracilis Cope 1883 [synonym]
†Anisonchus willeyi Rigby, Jr. 1980
Invalid names: Zetodon Cope 1883 [synonym]
G. †Auraria Middleton and Dewar 2004
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†Auraria urbana Middleton and Dewar 2004
G. †Beornus Atteberry and Eberle 2021
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†Beornus honeyi Atteberry and Eberle 2021
G. †Conacodon Matthew 1897
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†Conacodon cophater Cope 1884
†Conacodon delphae Eberle and Lillegraven 1998
†Conacodon harbourae Eberle and Lillegraven 1998
†Conacodon hettingeri Atteberry and Eberle 2021
†Conacodon kohlbergeri Archibald et al. 1983
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Invalid names: Conacodon utahensis Robison 1986 [synonym]
†Conacodon matthewi Middleton and Dewar 2004
Subfm. †Conacodontinae Archibald et al. 1983
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G. †Militocodon Weaver et al. 2024
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†Militocodon lydae Weaver et al. 2024
G. †Escatepos Reynolds 1936
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†Escatepos campi Reynolds 1936
G. †Goleroconus McKenna and Honey 2008
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†Goleroconus alfi McKenna and Honey 2008
G. †Hemithlaeus Cope 1882
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†Hemithlaeus josephi Van Valen 1978
G. †Miniconus Atteberry and Eberle 2021
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†Miniconus archibaldi Middleton and Dewar 2004
†Miniconus jeanninae Atteberry and Eberle 2021
G. †Mithrandir Cifelli 1983
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†Mithrandir gillianus Cope 1882
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Invalid names: Anisonchus apiculatus Cope 1884 [synonym]
Invalid names: Gillisonchus Rigby 1981 [synonym]
G. †Oxyacodon Osborn and Earle 1895
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†Oxyacodon agapetillus Cope 1884
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Invalid names: Fimbrethil ambaronae Van Valen 1978 [synonym]
†Oxyacodon apiculatus Osborn and Earle 1895
†Oxyacodon ferronensis Archibald et al. 1983
†Oxyacodon marshater Van Valen 1978
†Oxyacodon priscilla Matthew 1937
Invalid names: Fimbrethil Van Valen 1978 [synonym]
G. †Paleoungulatum Kelly 2014
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†Paleoungulatum hooleyi Kelly 2014
Subfm. †Periptychinae Osborn and Earle 1895
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G. †Ectoconus Cope 1884
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†Ectoconus cavigellii Eberle and Lillegraven 1998
†Ectoconus ditrigonus Cope 1882
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Invalid names: Ectoconus majusculus Matthew 1937 [synonym]
†Ectoconus symbolus Gazin 1941
G. †Periptychus Cope 1881
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Invalid names: Catathlaeus rhabdodon Cope 1881 [synonym], Eohyus robustus Marsh 1894 [synonym], Periptychus gilmorei Gazin 1938 [synonym], Periptychus superstes Matthew 1935 [synonym]
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Invalid names: Periptychus brabensis Cope 1888 [synonym], Periptychus hamaxitus Gazin 1941 [synonym], Periptychus matthewi Simpson 1936 [synonym], Plagioptychus matthewi Simpson 1936 [synonym]
Invalid names: Carsioptychus Simpson 1936 [synonym], Catathlaeus Cope 1881 [synonym], Plagioptychus Matthew 1936 [synonym]
G. †Tinuviel Van Valen 1978
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†Tinuviel eurydice Van Valen 1978
Unr. Euungulata Waddell et al. 2001
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Unr. Artiodactylamorpha Spaulding et al. 2009
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Unr. Artiodactyla Owen 1848 [even-toed ungulate]
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Subfm. †Antiacodontinae
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G. †Sarcolemur Cope 1875
Superfm. †Cainotherioidea Camp and VanDerHoof 1940
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Fm. †Cainotheriidae Cope 1881
Fm. †Robiacinidae Weppe et al. 2019
Superfm. †Choeropotamoidea
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Fm. †Choeropotamidae Owen 1845
Fm. †Climacoceratidae Hamilton 1978
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G. †Climacoceras MacInnes 1936
G. †Prolibytherium Arambourg 1961
Invalid names: Climacoceratinae Hamilton 1978 [empty], Sperrgebietomerycinae Morales et al. 1999 [empty]
Fm. †Diacodexeidae Krishtalka and Stucky 1985
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G. †Bunophorus Sinclair 1914
G. †Diacodexis Cope 1882
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Invalid names: Trigonolestes Cope 1894 [synonym], Wasatchia Sinclair 1914 [synonym]
G. †Gujaratia Bajpai et al. 2005
G. †Jiangsudon Metais et al. 2008
G. †Magwetherium Ducrocq et al. 2016
G. †Neodiacodexis West and Atkins 1970
G. †Simpsonodus Krishtalka and Stucky 1986
G. †Tapochoerus McKenna 1959
Invalid names: Bunophorinae Stucky and Krishtalka 1990 [empty]
Superfm. †Dichobunoidea Gill 1872
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G. †Acoessus Cope 1881
Fm. †Cebochoeridae Lydekker
Fm. †Dichobunidae Gill 1872
Fm. †Helohyidae Marsh 1877
Fm. †Homacodontidae Marsh 1849
Fm. †Leptochoeridae Marsh 1894
Fm. †Mixtotheriidae Pearson 1927
Fm. †Raoellidae Sahni et al. 1981
G. †Eolantianus Averianov 1996
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†Eolantianus russelli Averianov 1996
G. †Erlianhyus Li and Li 2022
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†Erlianhyus primitivus Li and Li 2022
G. †Hsanotherium Ducrocq et al. 2000
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†Hsanotherium parvum Ducrocq et al. 2000
G. †Myanmarius Tsubamoto et al. 2013
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†Myanmarius chitseini Tsubamoto et al. 2013
Subor. Omnivora
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G. †Hadrohyus Leidy 1872
G. †Raphenacodus Gabunia 1971
Unr. Ruminantiamorpha Spaulding et al. 2009
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Fm. †Protoceratidae Marsh 1891 [protoceratid]
Unr. Ruminantia Scopoli 1777 [ruminant]
Subor. Suiformes Jaeckel 1911
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Invalid names: Anthracotherioidea Gill 1872 [empty], Palaeodonta Matthew [synonym]
Superfm. Suoidea Gray 1821
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G. †Doliochoerus Filhol 1882
G. †Egatochoerus Ducrocq 1994
G. †Odoichoerus Tong and Zhao 1986
G. †Palaeochoerus Pomel 1847
G. †Schizochoerus Crusafont-Pairo and Lavocat 1954
Fm. Suidae Gray 1821 [pig]
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Invalid names: Phacochoeridae Gray 1868 [synonym]
G. †Taucanamo Simpson 1945
Invalid names: Palaeochoeridae Matthew 1924 [invalid subgroup], Sanitheriidae Simpson 1945 [empty], Suina Gray 1868 [invalid subgroup], Suinamorpha Spaulding et al. 2009 [synonym]
Fm. †Tapirulidae Cope 1879
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G. †Obotherium Bai et al. 2023
G. †Tapiruloides Bai et al. 2023
G. †Tapirulus Gervais 1850
G. †Tragulohyus Gervais 1874
Subor. Tylopoda Illiger 1811
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Fm. †Amphimerycidae Stehlin 1910
Superfm. †Anoplotherioidea Bonaparte 1850
Fm. †Oromerycidae Gazin 1955
Invalid names: Camelidamorpha Spaulding et al. 2009 [synonym], Cameloidea [empty], Oreodontoidea Gill 1872 [empty], Xiphodontoidea Flower 1883 [empty]
Invalid names: Achaenodontidae [empty], Ancodonta Matthew [empty], Antiacodontidae Krishtalka and Stucky 1985 [empty], Booidea [empty], Bunomerycidae Gentry and Hooker 1988 [empty], Cetartiodactyla Montgelard et al. 1997 [synonym], Eurytheriidae Cope 1889 [empty], Haplobunodontidae Sudre 1978 [empty], Hyodonta Matthew 1929 [empty], Leptochoerinae [empty], Merycoidodontoidea [empty], Neoselenodontia Webb and Taylor 1980 [empty], Nonruminantia Gregory 1910 [empty], Trigonolestidae [empty]
Unr. †Panperissodactyla Welker et al. 2015
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Or. †Panameriungulata de Muizon and Cifelli 2000
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Fm. †Didolodontidae Simpson 1934
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G. †Archaeohyracotherium Ameghino 1906
G. †Argyrolambda Ameghino 1904
G. †Depaulacoutoia Cifelli and Ortiz-Jaureguizar 2014
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Invalid names: Paulacoutoia Cifelli 1983 [replaced]
G. †Didolodus Ameghino 1897
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Invalid names: Cephanodus Ameghino 1902 [synonym], Lonchoconus Ameghino 1901 [synonym], Nephacodus Ameghino 1902 [synonym]
G. †Enneoconus Ameghino 1901
G. †Ernestokokenia Ameghino 1901
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Invalid names: Notoprotogonia Ameghino 1904 [synonym]
G. †Escribania Bonaparte et al. 1993
G. †Lamegoia Paula Couto 1952
G. †Paulogervaisia Ameghino 1901
G. †Proectocion Ameghino 1904
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Invalid names: Oxybunotherium Pascual 1965 [synonym]
G. †Raulvaccia Bonaparte et al. 1993
G. †Ricardocifellia Mones 2015
G. †Salladolodus Soria and Hoffstetter 1983
G. †Saltaodus Gelfo et al. 2020
G. †Umayodus Gelfo and Sigé 2011
Invalid names: Bunolitopternidae Schlosser 1823 [synonym], Didolodidae Scott 1913 [replaced]
Subor. †Litopterna Ameghino 1889
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Fm. †Adianthidae Ameghino 1891
G. †Eolicaphrium Ameghino 1902
Subor. †Lopholipterna Cifelli 1983
Fm. †Macraucheniidae Ameghino 1889
Fm. †Mesorhinidae Ameghino 1891
Fm. †Notonychopidae Soria 1989
Fm. †Proterotheriidae Ameghino 1887
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Invalid names: Bunodontheridae Mercerat 1891 [synonym]
Fm. †Protolipternidae Cifelli 1983
Fm. †Sparnotheriodontidae Soria 1980
G. †Thoatheriopsis Soria 2001
Invalid names: Eolitopterna Soria 2001 [empty], Lambdaconus elegans Roth 1904 [nomen dubium], Paulogervaisia celata Ameghino 1901 [nomen dubium], Proacrodon Roth 1899 [nomen vanum], Proacrodon transformatus Roth 1899 [nomen vanum], Proadiantus gibbus Ameghino 1901 [nomen vanum]
Superor. Perissodactylamorpha Rose et al. 2020
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Or. Perissodactyla Owen 1848 [odd-toed ungulate]
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Fm. †Anthracobunidae Wells and Gingerich 1983
Fm. †Brontotheriidae Marsh 1873 [brontothere]
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Invalid names: Titanotheriidae [synonym]
Fm. †Cambaytheriidae Bajpai et al. 2005
G. †Cambaytherium Bajpai et al. 2005
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Invalid names: Indobune Rose et al. 2006 [synonym]
Or. †Desmostylia Reinhart 1953 [desmostylian]
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Invalid names: Desmostyliformes Hay 1923 [synonym]
Superfm. Equoidea Hay 1902
G. †Ghazijhippus Missiaen and Gingerich 2014
G. †Hallensia Franzen and Haubold 1986
Subor. †Lophodontomorpha Hooker and Dashzeveg 2004
G. †Mesolambdolophus Holbrook and Lapergola 2011
G. †Nakusia Ginsburg et al. 1999
G. †Paschatherium Russell 1963
G. †Perissobune Missiaen and Gingerich 2014
G. †Skopaiolophus Métais et al. 2006
Subor. †Solidungula Cuvier
Unr. Tapiromorpha Haeckel 1873
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Invalid names: Moropomorpha [synonym]
G. †Teilhardimys Kretzoi and Kretzoi 2000
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Invalid names: Microhyus Teilhard de Chardin 1927 [replaced]
Invalid names: Chelodactyla Scott 1913 [empty], Equina Bonaparte 1850 [empty], Hippopotamus iravaticus Noetling 1897 [nomen dubium], Hyracotheriidae Cope 1881 [empty], Mesaxonia Marsh 1884 [empty], Rhinocerotida Owen 1845 [empty], Selenida McKenna and Bell 1997 [invalid subgroup], Stereopterna Ameghino 1889 [empty], Triplopodidae Cope 1881 [empty]
G. †Radinskya McKenna et al. 1989
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†Radinskya yupingae McKenna et al. 1989
Invalid names: Anthracobunia Ginsburg et al. 1999 [empty]
Unr. Phenacodonta McKenna 1975
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G. †Olbitherium Tong et al. 2004
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†Olbitherium millenariusum Tong et al. 2004
Fm. †Phenacodontidae Cope 1881 [condylarth]
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G. †Almogaver Crusafont-Pairo and Villalta 1954
G. †Copecion Gingerich 1989
G. †Eodesmatodon Zheng and Chi 1978
G. †Lophocion Wang and Tong 1997
Subfm. †Meniscotheriinae Cope 1882
G. †Phenacodus Cope 1873
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Invalid names: Eohyus Marsh 1894 [synonym], Opisthotomus Cope 1875 [synonym], Trispondylus Cope 1884 [replaced]
G. †Tetraclaenodon Scott 1892
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Invalid names: Euprotogonia Cope 1893 [synonym], Protogonia Cope 1881 [synonym]
Invalid names: Eparctocyona McKenna 1975 [synonym], Ungulata Linnaeus 1766 [synonym]
Unr. Ferae Linnaeus 1758
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Unr. Carnivoramorpha Wyss and Flynn 1993
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Unr. †Carnivoraformes Flynn et al.
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G. †Africtis Mattingly et al. 2020
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†Africtis sirtensis Mattingly et al. 2020
Or. Carnivora Bowdich 1821 [carnivoran]
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G. †Adracon Filhol 1884
Fm. Canidae Fischer de Waldheim 1817 [canid]
Subor. Caniformia Kretzoi 1943
Subor. Feliformia Kretzoi 1945
Subor. Fissipedia Blumenbach 1791
Superfm. †Miacoidea Cope
G. †Monachiopsis Kretzoi 1941
Fm. †Percrocutidae Werdelin and Solounias 1991
G. †Pristinictis Fox and Youzwyshyn 1994
G. †Ravenictis Fox and Youzwyshyn 1994
Invalid names: Eosictis Scott 1945 [nomen dubium], Felidi Jaekel 1911 [empty], Phocidi Jaekel 1911 [empty], Protelidi Jaekel 1911 [empty], Protoferi Jaekel 1911 [empty], Ursina Gray 1825 [empty], Ursocanidi Jaekel 1911 [empty], Viverra memphitis Linnaeus 1758 [nomen vanum]
G. †Dormaalocyon Solé et al. 2014
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†Dormaalocyon latouri Quinet 1966
G. †Gracilocyon Smith and Smith 2010
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†Gracilocyon igniculus Beard and Dawson 2009
†Gracilocyon rosei Heinrich et al. 2008
†Gracilocyon rundlei Hooker 2010
†Gracilocyon solei Smith and Smith 2010
†Gracilocyon winkleri Gingerich 1983
G. Lycarion Matthew 1909
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†Lycarion medius Matthew 1909
G. †Miacis Cope 1872
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†Miacis bathygnathus Scott 1888
†Miacis boqinghensis Huang et al. 1999
†Miacis deutschi Gingerich 1983
†Miacis edax Leidy 1872
†Miacis exiguus Matthew and Granger 1915
†Miacis gracilis Clark 1939
†Miacis hookwayi Stock 1934
†Miacis invictus Matthew and Granger 1925
†Miacis latidens Matthew and Granger 1915
†Miacis lushiensis Chow 1975
†Miacis parvivorus Cope 1872
†Miacis petilus Gingerich 1983
†Miacis rundlei Hooker 2010
†Miacis vorax Leidy 1872
†Miacis vulpinus Scott and Osborn 1887
Invalid names: Harpalodon Marsh 1872 [synonym], Prodaphaenus Wortman and Matthew 1899 [synonym]
G. †Neovulpavus Wortman 1901
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†Neovulpavus mccarrolli Tomiya et al. 2021
†Neovulpavus washakius Wortman 1901
G. †Palaearctonyx Matthew 1909
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†Palaearctonyx meadi Matthew 1909
G. †Paramiacis Filhol 1876
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†Paramiacis exilis Filhol 1876
†Paramiacis teilhardi Mathis 1987
G. †Procynodictis Wortman and Matthew 1899
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†Procynodictis vulpiceps Wortman and Matthew 1899
Invalid names: Plesiomiacis Simpson 1945 [synonym]
G. †Quercygale Kretzoi 1945
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†Quercygale angustidens Filhol 1872
†Quercygale hastingsae Davies 1884
†Quercygale helvetica Rütimeyer 1862
†Quercygale smithi Solé 2014
Invalid names: Humbertia De Beaumont 1965 [objective synonym]
G. †Uintacyon Leidy 1873
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†Uintacyon acutus Thorpe 1923
†Uintacyon asodes Gazin 1952
†Uintacyon gingerichi Heinrich et al. 2008
†Uintacyon hookeri Solé 2014
†Uintacyon jugulans Matthew 1909
†Uintacyon rudis Rose 1981
G. †Vassacyon Matthew 1909
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†Vassacyon bowni Heinrich et al. 2008
†Vassacyon prieuri Solé et al. 2016
†Vassacyon promicrodon Wortman and Matthew 1899
†Vassacyon taxidiotis Solé et al. 2013
G. †Vulpavus Marsh 1871
+
Subg. †Vulpavus (Phlaodectes) Matthew 1909
†Vulpavus australis Matthew and Granger 1915
+
Invalid names: Vulpavus asius Gazin 1952 [synonym]
†Vulpavus canavus Cope 1881
+
Invalid names: Miacis brevirostris Cope 1881 [synonym]
†Vulpavus completus Matthew 1909
†Vulpavus farsonensis Gunnell 1998
†Vulpavus hargeri Wortman 1901
†Vulpavus ovatus Matthew 1909
†Vulpavus palustris Marsh 1871
†Vulpavus profectus Matthew 1909
G. †Xinyuictis Zheng et al. 1975
+
†Xinyuictis tenuis Zheng et al. 1975
G. †Zodiocyon Tong and Wang 2006
+
†Zodiocyon zetesios Tong and Wang 2006
G. †Dawsonicyon Spaulding et al. 2010
+
†Dawsonicyon isami Spaulding et al. 2010
G. †Oodectes Wortman 1901
+
†Oodectes herpestoides Wortman 1901
+
Invalid names: Oodectes proximus Matthew 1909 [synonym], Paeneprolimnocyon amissadomus Guthrie 1967 [synonym], Triacodon nanus Marsh 1872 [synonym]
†Oodectes jepseni Guthrie 1967
+
Invalid names: Paeneprolimnocyon iudei Guthrie 1967 [synonym]
†Oodectes pugnax Wortman and Matthew 1899
Invalid names: Paeneprolimnocyon Guthrie 1967 [synonym]
G. †Tapocyon Stock 1934
+
†Tapocyon dawsonae Wesley and Flynn 2003
†Tapocyon robustus Peterson 1919
+
Invalid names: Tapocyon occidentalis Stock 1934 [synonym]
Fm. †Viverravidae Wortman and Matthew 1899
+
G. †Didymictis Cope 1875
+
†Didymictis altidens Cope 1880
†Didymictis dellensis Dorr 1952
†Didymictis leptomylus Cope 1880
†Didymictis protenus Cope 1874
+
Invalid names: Didymictis curtidens Cope 1882 [synonym]
†Didymictis proteus Polly 1997
†Didymictis vancleveae Robinson 1966
G. †Orientictis Huang and Zheng 2005
+
†Orientictis spanios Huang and Zheng 2005
G. †Preonictis Tong and Wang 2006
+
†Preonictis youngi Tong and Wang 2006
G. †Protictis Matthew
+
Subg. †Protictis (Bryanictis) MacIntyre 1966
Subg. †Protictis (Protictis) Matthew 1937
Subg. †Protictis (Protictoides) Flynn and Galiano 1982
†Protictis agastor Gingerich and Winkler 1985
†Protictis haydenianus Cope 1882
+
Invalid names: Didymictis primus Cope 1884 [synonym], Prolimnocyon macfaddeni Rigby, Jr. 1980 [synonym]
†Protictis minor Meehan and Wilson 2002
†Protictis paralus Holtzman 1978
†Protictis simpsoni Meehan and Wilson 2002
G. †Simpsonictis MacIntyre 1962
+
†Simpsonictis jaynanneae Rigby, Jr. 1980
†Simpsonictis pegus Gingerich and Winkler 1985
†Simpsonictis tenuis Simpson 1935
G. †Variviverra Tong and Wang 2006
+
†Variviverra vegetatus Tong and Wang 2006
G. †Viverravus Marsh 1872
+
†Viverravus acutus Matthew and Granger 1915
†Viverravus gracilis Marsh 1872
+
Invalid names: Didymictis dawkinsianus Cope 1881 [synonym], Harpalodon vulpinus Marsh 1872 [synonym], Triacodon fallax Marsh 1871 [synonym], Ziphacodon rugatus Marsh 1872 [synonym]
†Viverravus lawsoni Hooker 2010
†Viverravus laytoni Gingerich and Winkler 1985
+
Invalid names: Viverravus bowni Gingerich 1987 [synonym]
†Viverravus lutosus Gazin 1952
†Viverravus minutus Wortman 1901
†Viverravus nitidus Marsh 1872
†Viverravus politus Matthew and Granger 1915
†Viverravus rosei Polly 1997
†Viverravus schaffi Gingerich and Winkler 1985
†Viverravus sicarius Matthew 1909
Invalid names: Ziphacodon Marsh 1872 [synonym]
G. †Viverriscus Beard and Dawson 2009
+
†Viverriscus omnivorus Beard and Dawson 2009
Invalid names: Deltatherium durini Van Valen 1978 [nomen dubium], Didymictidae Flynn and Galiano 1982 [synonym]
Or. †Creodonta Cope 1875 [creodont]
+
G. †Altacreodus Fox 2015
+
†Altacreodus magnus Clemens and Russell 1965
Or. †Hyaenodonta Van Valen 1967
+
G. †Eoproviverra Solé et al. 2014
+
†Eoproviverra eisenmanni Godinot 1981
Superfm. †Hyaenodontoidea Leidy 1869
+
Fm. †Hyaenodontidae Leidy 1869
Subfm. †Proviverrinae Schlosser 1886
Superfm. †Hyainailouroidea Borths et al. 2016
+
Unr. †Hyainailouridae Solé et al. 2015
G. †Kyawdawia Egi et al. 2005
G. †Metasinopa Osborn 1909
G. †Paratritemnodon Ranga Rao 1973
Unr. †Teratodontinae Savage 1965
Subfm. †Limnocyoninae Wortman 1902
+
G. †Iridodon Morlo and Gunnell 2003
G. †Limnocyon Marsh 1872
+
Invalid names: Telmatocyon Marsh 1899 [synonym]
G. †Oxyaenodon Matthew 1899
G. †Prolaena Xu et al. 1979
G. †Prolimnocyon Matthew and Granger 1915
G. †Thinocyon Marsh 1872
Invalid names: Limnocyonini Wortman 1902 [empty], Machaeroidini Matthew 1909 [empty]
Unr. †Oxyaenodonta Van Valen 1971
+
Fm. †Oxyaenidae Cope 1877
+
G. †Dipsalidictis Matthew and Granger 1915
+
Invalid names: Dipsalidictides Denison 1938 [synonym]
G. †Dipsalodon Jepsen 1930
G. †Dormaalodon Lange-Badré 1987
G. †Galethylax Gervais 1850
G. †Machaeroides Matthew 1909
Subfm. †Machaeroidinae
Subfm. †Oxyaeninae Trouessart 1885
Subfm. †Palaeonictinae Denison 1938
G. †Sarkastodon Granger 1938
G. †Tytthaena Gingerich 1980
Invalid names: Ambloctonidae Cope 1877 [empty], Amblyctonidae Cope 1877 [empty], Arctocyonoidae Trouessart 1885 [empty], Hyaenodontida Leidy 1869 [empty], Limnocyonidae [empty], Oxyaenoidae Cope 1877 [empty], Uintacyonidae Hay 1902 [empty]
Unr. †Pholidotamorpha Gaudin et al. 2009
+
Subor. †Palaeanodonta Matthew 1918
+
G. †Amelotabes Rose 1978
+
†Amelotabes simpsoni Rose 1978
G. †Arcticanodon Rose et al. 2004
+
†Arcticanodon dawsonae Rose et al. 2004
Fm. †Epoicotheriidae Simpson 1927
+
G. †Alocodontulum Rose et al. 1978
G. †Auroratherium Tong and Wang 1997
G. †Dipassalus Rose et al. 1991
G. †Epoicotherium Simpson 1927
+
Invalid names: Pseudochrysochloris Turnbull and Reed 1967 [synonym], Xenotherium Douglass 1905 [replaced]
G. †Tetrapassalus Simpson 1959
G. †Tubulodon Jepsen 1932
+
Invalid names: Alocodon Rose et al. 1977 [synonym], Pentapassalus Gazin 1952 [synonym]
G. †Xenocranium Colbert 1942
G. †Ernanodon Ding 1979
+
†Ernanodon antelios Ding 1979
Fm. †Escavadodontidae Rose and Lucas 2000
+
G. †Escavadodon Rose and Lucas 2000
G. †Melaniella Fox 1984
+
†Melaniella timosa Fox 1984
Fm. †Metacheiromyidae Wortman 1903
+
G. †Brachianodon Gunnell and Gingerich 1993
G. †Metacheiromys Wortman 1903
G. †Mylanodon Secord et al. 2002
G. †Palaeanodon Matthew 1918
G. †Propalaeanodon Rose 1979
Or. Pholidota Weber 1904 [pangolin]
+
Unr. †Eupholidota Gaudin et al. 2009
+
G. †Eomanis Storch 1978
Unr. †Manoidea Gaudin et al.
G. †Eurotamandua Storch 1981
+
Eurotamandua joresi Storch 1981
G. †Leptomanis Filhol 1894
+
†Leptomanis edwardsi Filhol 1894
Invalid names: Eomanidae Storch 2003 [empty], Squamata Huxley 1872 [synonym]
Invalid names: Ostentoria [synonym]
Unr. Fereuungulata Waddell et al. 1999
hide
Invalid names: Zooamata Waddell et al. 1999 [empty]
Invalid names: Variamana [synonym]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: phosphaticsubp
Maximum body mass: 1000 kgg
Minimum body mass: 300 gs
Environment: terrestrialsubc
Locomotion: actively mobilec
Life habit: scansorialsubc
Diet: insectivoresubc
Reproduction: viviparoussubc
Created: 2005-09-06 20:43:42
Modified: 2005-09-06 22:44:32
Source: s = species, g = genus, subc = subclass, c = class, subp = subphylum
References: Benton 1996, Legendre 1988, Carroll 1988, Ji et al. 2002, Lillegraven 1979, Hendy et al. 2009

Age range

Maximum range based only on fossils: base of the Middle Turonian to the top of the Holocene or 93.50000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 89.3 Ma

Collections (15728 total)

Oldest occurrences

Time interval Ma Country or state Original ID and collection number
Middle Turonian - Late/Upper Turonian93.5 - 89.3Uzbekistan (Navoi) Periptychidae indet. (58168)
Maastrichtian72.1 - 66.0India (Madhya Pradesh) Kharmerungulatum vanvaleni (75493)
Lancian70.6 - 66.0USA (Wyoming) Cimolestes magnus (14519 14598) Protungulatum sp. (14514)
Lancian70.6 - 66.0USA (Montana) Cimolestes magnus (14516 14532)
Lancian70.6 - 66.0Canada (Saskatchewan) Cimolestes magnus (14558)
Lancian70.6 - 66.0Canada (Alberta) Cimolestes magnus (14630 14633 14635 14637)
Late/Upper Maastrichtian70.6 - 66.0USA (North Dakota) Cimolestes magnus (45144)
Late/Upper Maastrichtian70.6 - 66.0USA (Montana) Paleoungulatum hooleyi, Baioconodon nordicus (183109) Protungulatum coombsi (110464) Protungulatum gorgun (14674)
Pu166.0 - 63.3USA (Montana) Mimatuta minuial, Protungulatum donnae (14762)
Pu366.0 - 63.3USA (Montana) Eoconodon hutchisoni (143205) Eoconodon hutchisoni, Eoconodon nidhoggi (218647) Oxyacodon ferronensis, Eoconodon hutchisoni, Eoconodon nidhoggi (14738)
Puercan66.0 - 63.3USA (New Mexico) Arctocyon ferox (74549 125183) Ectoconus ditrigonus, Periptychus coarctatus, Escatepos campi, Hemithlaeus kowalevskianus, Conacodon entoconus, Conacodon cophater, Conacodon kohlbergeri, Oxyacodon apiculatus, Oxyacodon priscilla, Eoconodon sp., Eoconodon ginibitohia, Eoconodon coryphaeus, Eoconodon gaudrianus, Oxyclaenus simplex, Oxyclaenus cuspidatus, Loxolophus kimbetovius, Mimotricentes mirielae, Fimbrethil ambaronae, Gillisonchus gillianus (14760) Ectoconus ditrigonus, Periptychus coarctatus, Hemithlaeus kowalevskianus, Conacodon entoconus, Conacodon kohlbergeri, Eoconodon coryphaeus, Eoconodon gaudrianus, Dissacus navajovius, Oxyclaenus simplex, Oxyclaenus cuspidatus, Loxolophus hyattianus, Loxolophus kimbetovius, Gillisonchus gillianus (14703) Ectoconus ditrigonus, Periptychus coarctatus, Hemithlaeus kowalevskianus, Conacodon entoconus, Eoconodon gaudrianus, Loxolophus hyattianus, Gillisonchus gillianus (14794) Hemithlaeus sp., Conacodon sp., Ectoconus sp., Anisonchus sp., Carsioptychus sp. (14661) Oxyacodon apiculatus, Eoconodon coryphaeus, Oxyclaenus simplex, Oxyclaenus cuspidatus, Loxolophus priscus, Loxolophus hyattianus, Loxolophus pentacus, Chriacus antiquus, Protogonodon grangeri, Conacodon sp., Ectoconus sp., Gillisonchus gillianus, Plagioptychus matthewi (14662) Periptychus coarctatus, Loxolophus priscus, Loxolophus hyattianus, Loxolophus pentacus, Gillisonchus gillianus (14795)
Puercan66.0 - 63.3USA (North Dakota) Oxyacodon priscilla, Protungulatum sp. (14765)
Puercan66.0 - 63.3USA (Utah) Ectoconus ditrigonus, Anisonchus athelas, Loxolophus sp., Conacodon utahensis (14739) Ectoconus ditrigonus, Ectoconus symbolus, Anisonchus oligistus, Anisonchus athelas, Oxyacodon ferronensis, Loxolophus pentacus, Periptychus hamaxitus (14736) Ectoconus ditrigonus, Ectoconus symbolus, Periptychus coarctatus, Haploconus sp., Haploconus elachistus, Anisonchus athelas, Anisonchus onostus, Oxyacodon apiculatus, Oxyclaenus pugnax, Loxolophus spiekeri (14692) Ectoconus sp., Ectoconus symbolus, Periptychus hamaxitus (14748) Ectoconus symbolus, Carsioptychus hamaxitus, Anisonchus oligistus, Anisonchus athelas, Oxyacodon marshater, Goniacodon hiawathae, Oxyclaenus sp., Loxolophus sp., Protogonodon sp., Periptychus hamaxitus, Conacodon utahensis, Haploconus sp. (14790) Ectoconus symbolus, Periptychus coarctatus, Oxyacodon sp., Oxyacodon apiculatus, Oxyacodon ferronensis, Oxyclaenus pugnax, Loxolophus pentacus (14670) Eoconodon sp., Oxyclaenus simplex (14791) Periptychidae indet., Ectoconus maiusculus, Anisonchus oligistus (224186)
Puercan66.0 - 63.3USA (Wyoming) Arctocyonidae indet. (14715 14735) Arctocyonidae indet., Oxyprimus galadrielae (14804) Baioconodon sp. (14712) Conacodon cophater (14727) Ectoconus ditrigonus (14717) Ectoconus ditrigonus, Periptychus coarctatus, Conacodon cophater, Arctocyonidae indet. (14716) Eoconodon sp., Oxyprimus sp. (14710) Maiorana ferrisensis, Mimatuta sp., Protungulatum donnae (14711) Mimatuta makpialutae, Ragnarok wovokae (14807) Mimatuta sp. (14725) Miniconus jeanninae, Beornus honeyi, Ampliconus antoni, Conacodon hettingeri, Conacodon harbourae, Oxyprimus sp., Maiorana noctiluca, Mimatuta sp., Sigynorum magnadivisus, Protungulatum donnae (174233) Mithrandir gillianus (14714 14718 14724 14726) Mithrandir gillianus, Conacodon cophater, Arctocyonidae indet., Oxyclaenus cuspidatus (14723) Mithrandir gillianus, Loxolophus faulkneri (14707) Mithrandir gillianus, Loxolophus hyattianus (14728) Mithrandir gillianus, Loxolophus sp., Loxolophus faulkneri, Ampliconus browni (14713) Mithrandir gillianus, Oxyacodon agapetillus, Eoconodon sp., Arctocyonidae indet., Loxolophus hyattianus (14731) Oxyacodon apiculatus (14734) Oxyclaenus simplex (14803) Oxyclaenus sp., Oxyclaenus cuspidatus, Baioconodon denverensis (14729) Oxyprimus putorius, Oxyprimus galadrielae, Maiorana noctiluca, Mimatuta minuial, Loxolophus nordicus, Eoconodon sp., Oxyacodon josephi (14808) Oxyprimus sp., Oxyprimus galadrielae, Mimatuta sp., Protungulatum sloani, Protungulatum donnae (14796) Periptychidae indet., Mithrandir gillianus, Ectoconus cavigellii, Ampliconus antoni, Alticonus gazini, Oxyacodon agapetillus, Oxyacodon priscilla, Baioconodon middletoni, Loxolophus priscus, Loxolophus hyattianus (14797) Periptychidae indet., Mithrandir gillianus, Ectoconus sp., Conacodon delphae, Oxyacodon agapetillus, Arctocyonidae indet., Protungulatum gorgun, Protungulatum donnae, Conacodon harbourae (14722) Periptychidae indet., Mithrandir gillianus, Periptychus coarctatus, Oxyacodon priscilla, Loxolophus faulkneri (14706) Periptychidae indet., Periptychus coarctatus (14719) Periptychus coarctatus (14721 14733) Periptychus coarctatus, Oxyacodon priscilla, Loxolophus priscus, Protungulatum donnae (14720) Protungulatum donnae (14708 14709) Triisodontinae indet., Arctocyonidae indet. (14805)
Puercan66.0 - 63.3USA (Montana) Baioconodon nordicus (14693) Eoconodon hutchisoni (140079 218651 218655) Eoconodon nidhoggi (218650) Mimatuta minuial (14701) Mimatuta minuial, Protungulatum donnae (231047) Mimatuta minuial, Ragnarok engdahli (14694) Mimatuta morgoth (14675 108198 108199) Mimatuta morgoth, Baioconodon sp., Protungulatum sp., Protungulatum gorgun, Protungulatum donnae (14672) Mimatuta morgoth, Protungulatum gorgun, Ragnarok nordicum (14782) Mimatuta sp. (14801 14802) Mimatuta sp., Mimatuta morgoth, Protungulatum donnae (14771) Mimatuta sp., Mimatuta morgoth, Protungulatum gorgun, Protungulatum donnae, Ragnarok nordicum (14787) Mimatuta sp., Mimatuta morgoth, Protungulatum gorgun, Ragnarok nordicum (14763) Mimatuta sp., Protungulatum sp. (14770) Mimatuta sp., Ragnarok nordicum (14668 14669 14740 14789) Oxyacodon ferronensis (218654) Oxyacodon ferronensis, Eoconodon hutchisoni (14799) Oxyacodon ferronensis, Eoconodon hutchisoni, Eoconodon nidhoggi (218648) Oxyacodon ferronensis, Eoconodon nidhoggi (218649) Oxyprimus erikseni (14696 14702) Oxyprimus erikseni, Mimatuta minuial, Protungulatum donnae, Ragnarok nordicum (14786) Oxyprimus erikseni, Mimatuta morgoth, Protungulatum donnae (14671) Oxyprimus erikseni, Mimatuta morgoth, Protungulatum donnae, Ragnarok engdahli (14798) Oxyprimus erikseni, Mimatuta morgoth, Protungulatum donnae, Ragnarok nordicum (14749) Oxyprimus erikseni, Mimatuta morgoth, Protungulatum gorgun, Protungulatum donnae, Ragnarok nordicum (14699 14754) Oxyprimus erikseni, Mimatuta morgoth, Ragnarok engdahli (14742) Oxyprimus erikseni, Mimatuta morgoth, Ragnarok harbichti, Protungulatum sp., Protungulatum gorgun, Protungulatum donnae (14741) Oxyprimus erikseni, Mimatuta sp. (14767) Oxyprimus erikseni, Mimatuta sp., Baioconodon nordicum, Ragnarok harbichti (14743) Oxyprimus erikseni, Mimatuta sp., Mimatuta minuial, Baioconodon sp., Baioconodon engdahli (231045) Oxyprimus erikseni, Mimatuta sp., Mimatuta minuial, Protungulatum donnae (193844) Oxyprimus erikseni, Mimatuta sp., Mimatuta morgoth, Mimatuta minuial, Protungulatum donnae, Ragnarok nordicum (14777) Oxyprimus erikseni, Mimatuta sp., Mimatuta morgoth, Mimatuta minuial, Protungulatum gorgun, Protungulatum donnae, Ragnarok nordicum (14788) Oxyprimus erikseni, Mimatuta sp., Mimatuta morgoth, Protungulatum donnae, Ragnarok nordicum (14751) Oxyprimus erikseni, Mimatuta sp., Protungulatum gorgun (14776) Oxyprimus erikseni, Mimatuta sp., Protungulatum gorgun, Protungulatum donnae (14780) Oxyprimus erikseni, Mimatuta sp., Protungulatum gorgun, Protungulatum donnae, Ragnarok nordicum (14784 14785) Oxyprimus erikseni, Mimatuta sp., Ragnarok nordicum (14773) Oxyprimus erikseni, Protungulatum gorgun (14698) Oxyprimus erikseni, Ragnarok nordicum (14660) Oxyprimus sp. (14665 14673) Oxyprimus sp., Oxyprimus erikseni, Mimatuta morgoth, Mimatuta minuial, Baioconodon engdahli, Protungulatum gorgun, Protungulatum donnae, Baioconodon harbichti (14705) Periptychidae indet., Anisonchus sp., Oxyacodon sp., Oxyclaenus simplex, Oxyprimus sp., Baioconodon denverensis, Loxolophus sp., Carcinodon aquilonius, Protungulatum sp., Protogonodon pentacus (14809) Periptychidae indet., Mimatuta minuial, Ragnarok nordicum, Protungulatum mckeeveri (14761) Periptychidae indet., Mimatuta sp., Protungulatum donnae, Mimatuta sp. (14759) Protungulatum donnae (14663 14667 14753) Protungulatum donnae, Ragnarok nordicum (14779) Protungulatum gorgun (14755 14757) Protungulatum gorgun, Ragnarok nordicum (14676) Ragnarok nordicum (14697 14700 14752 14758 14778) Tinuviel eurydice, Anisonchus oligistus, Oxyclaenus pugnax, Protungulatum sp., Oxprimus albertensis, Tricentes sp., Eoconodon sp., Oxyacodon josephi, Eoconodon sp. (14766) Tinuviel eurydice, Oxyacodon apiculatus, Oxyacodon ferronensis, Eoconodon nidhoggi, Oxyclaenus sp., Baioconodon nordicus, Loxolophus sp., Loxolophus schizophrenus, Carcinodon sp. (14745) Tinuviel eurydice, Oxyacodon ferronensis, Baioconodon nordicus, Carcinodon sp. (14744)
Puercan66.0 - 63.3Canada (Saskatchewan) Anisonchus oligistus, Oxyacodon agapetillus, Eoconodon sp., Oxyclaenus corax, Baioconodon denverensis, Loxolophus schizophrenus, Carcinodon aquilonius, Carnivora indet., Ravenictis krausei (14768) Arctocyonidae indet., Oxyclaenus sp., Baioconodon denverensis, Loxolophus schizophrenus (14691) Oxyprimus erikseni, Mimatuta sp., Protungulatum donnae, Ragnarok sp. (14756) Periptychidae indet., Protungulatum donnae (14737)
Puercan66.0 - 63.3USA (Colorado) Ampliconus browni, Oxyprimus sp., Maiorana sp., Baioconodon nordicus, Protungulatum donnae (176863) Baioconodon denverensis, Tinuviel gazini (14774) Carsioptychus coarctatus (176748 205865 205867 205895 205900 205901 205903 205907 205908 205910 205914 205928 209872 209892 209904 209921 209924 209943 209946 209957 209964 209968 209976 210033) Carsioptychus coartatus (206072 206084) Carsioptychus sp. (206071) Conacodon delphae, Oxyclaenus cuspidatus (14689) Conacodon entoconus (205919) Conacodon harbourae (88207) Ectoconus ditrigonus (14684 14687 14688 205863 205864 205868 205871 205881 205886 209900 209983 209997) Ectoconus ditrigonus, Periptychus coarctatus (14679) Ectoconus ditrigonus, Periptychus coarctatus, Anisonchus athelas, Loxolophus hyattianus (14792) Eoconodon coryphaeus (205932 205934) Loxolophus sp. (209874 209879 209890 209965) Militocodon lydae (234689) Oxyacodon archibaldi, Auraria urbana, Ampliconus browni, Conacodon matthewi, Arctocyonidae indet., Oxyclaenus subbituminus, Oxyprimus galadrielae, Baioconodon denverensis, Baioconodon cannoni, Tinuviel gazini (14658) Periptychidae indet. (205873) Periptychus coarctatus (14659 14680 14681 14682 14685 14686 14746 14750 14793) Periptychus coarctatus, Conacodon delphae (14678) Periptychus coarctatus, Conacodon entoconus, Oxyclaenus cuspidatus, Loxolophus hyattianus (14683) Periptychus coarctatus, Loxolophus hyattianus (14677)
Early/Lower Paleocene66.0 - 61.7China (Anhui) Pappictidops orientalis (132943 146556)
Early/Lower Paleocene66.0 - 61.7China (Hunan) Dissacus rotundus (182815)
Early/Lower Paleocene66.0 - 61.7China (Shaanxi) Hukoutherium shimemensis (132651)