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Perissodactyla (odd-toed ungulate)
Taxonomy
Perissodactyla was named by Owen (1848) [Owen does not give a rank in his original description of Perissodactyla.
Owen does not give a rank in his original description of Perissodactyla.
Owen does not give a rank in his original description of Perissodactyla.]. It is extant.
It was reranked as the suborder Perissodactyla by Gadow (1898), Hay (1902), Lambe (1908) and Gregory (1910); it was synonymized subjectively with Mesodactyli by Jaekel (1911); it was reranked as the order Perissodactyla by Leidy (1873), Flower (1883), Ferrusquia-Villafranca (1969), McKenna (1975), Thurmond and Jones (1981), Schoch (1984), Kumar and Sahni (1985), Hooker (1986), MacFadden (1986), Carroll (1988), Prothero and Schoch (1989), Nowak (1991), Madsen et al. (2001), Wilson and Reeder (2005), Prothero (2005), Symeonidis et al. (2006), Hulbert and Whitmore (2006), MacFadden (2009), Asher and Helgen (2010), Cooper et al. (2014), Ferrusquía-Villafranca et al. (2014), Bai et al. (2017) and Rose et al. (2020).
It was assigned to Diplarthra by Cope (1889) and Cope (1891); to Ungulata by Owen (1848), Flower (1883) and Ameghino (1894); to Ungulata by Lydekker (1876), Flower and Lydekker (1891), Gadow (1898), Hay (1902) and Lambe (1908); to Mesaxonia by Gregory (1910); to Phenacodonta by McKenna (1975); to Eutheria by Hooker (1986); to Altungulata by Prothero and Schoch (1989); to Eutheria by Carroll (1988) and Nowak (1991); to Zooamata by Waddell et al. (1999); to Laurasiatheria by Madsen et al. (2001); to Euungulata by Waddell et al. (2001) and Asher and Helgen (2010); to Placentalia by Cooper et al. (2014); to Mammalia by Leidy (1873), Ferrusquia-Villafranca (1969), Thurmond and Jones (1981), Schoch (1984), Kumar and Sahni (1985), MacFadden (1986), Wilson and Reeder (2005), Prothero (2005), Symeonidis et al. (2006), Hulbert and Whitmore (2006), MacFadden (2009), Ferrusquía-Villafranca et al. (2014) and Bai et al. (2017); and to Perissodactylamorpha by Rose et al. (2020).
It was reranked as the suborder Perissodactyla by Gadow (1898), Hay (1902), Lambe (1908) and Gregory (1910); it was synonymized subjectively with Mesodactyli by Jaekel (1911); it was reranked as the order Perissodactyla by Leidy (1873), Flower (1883), Ferrusquia-Villafranca (1969), McKenna (1975), Thurmond and Jones (1981), Schoch (1984), Kumar and Sahni (1985), Hooker (1986), MacFadden (1986), Carroll (1988), Prothero and Schoch (1989), Nowak (1991), Madsen et al. (2001), Wilson and Reeder (2005), Prothero (2005), Symeonidis et al. (2006), Hulbert and Whitmore (2006), MacFadden (2009), Asher and Helgen (2010), Cooper et al. (2014), Ferrusquía-Villafranca et al. (2014), Bai et al. (2017) and Rose et al. (2020).
It was assigned to Diplarthra by Cope (1889) and Cope (1891); to Ungulata by Owen (1848), Flower (1883) and Ameghino (1894); to Ungulata by Lydekker (1876), Flower and Lydekker (1891), Gadow (1898), Hay (1902) and Lambe (1908); to Mesaxonia by Gregory (1910); to Phenacodonta by McKenna (1975); to Eutheria by Hooker (1986); to Altungulata by Prothero and Schoch (1989); to Eutheria by Carroll (1988) and Nowak (1991); to Zooamata by Waddell et al. (1999); to Laurasiatheria by Madsen et al. (2001); to Euungulata by Waddell et al. (2001) and Asher and Helgen (2010); to Placentalia by Cooper et al. (2014); to Mammalia by Leidy (1873), Ferrusquia-Villafranca (1969), Thurmond and Jones (1981), Schoch (1984), Kumar and Sahni (1985), MacFadden (1986), Wilson and Reeder (2005), Prothero (2005), Symeonidis et al. (2006), Hulbert and Whitmore (2006), MacFadden (2009), Ferrusquía-Villafranca et al. (2014) and Bai et al. (2017); and to Perissodactylamorpha by Rose et al. (2020).
Subtaxa
Anthracobunidae, Brontotheriidae (syn. Titanotheriidae), Cambaytheriidae, Cambaytherium (syn. Indobune), Chelodactyla, Desmostylia (syn. Desmostyliformes), Equina, Equoidea, Ghazijhippus, Hallensia, Hippopotamus iravaticus, Hyracotheriidae, Lophodontomorpha, Mesaxonia, Mesolambdolophus, Nakusia, Paschatherium, Perissobune, Rhinocerotida, Selenida, Skopaiolophus, Solidungula, Stereopterna, Tapiromorpha (syn. Moropomorpha), Teilhardimys (syn. Microhyus), Titanotheriomorpha
Synonyms
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Synonymy list
Year | Name and author |
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1848 | Perissodactyla Owen p. 131 |
1873 | Perissodactyla Leidy p. 321 |
1876 | Perissodactyla Lydekker |
1883 | Perissodactyla Flower p. 185 |
1889 | Perissodactyla Cope p. 876 |
1891 | Perissodactyla Cope p. 8 |
1891 | Perissodactyla Flower and Lydekker p. 90 |
1894 | Perissodactyla Ameghino p. 13 |
1897 | Hippopotamus iravaticus Noetling |
1898 | Perissodactyla Gadow p. 46 |
1902 | Perissodactyla Hay p. 607 |
1908 | Perissodactyla Lambe |
1910 | Perissodactyla Gregory p. 466 |
1969 | Perissodactyla Ferrusquia-Villafranca |
1975 | Perissodactyla McKenna p. 43 |
1981 | Perissodactyla Thurmond and Jones p. 182 |
1984 | Perissodactyla Schoch p. 3 |
1985 | Perissodactyla Kumar and Sahni p. 153 |
1986 | Perissodactyla Hooker p. 342 |
1986 | Perissodactyla MacFadden p. 467 |
1988 | Perissodactyla Carroll |
1989 | Perissodactyla Prothero and Schoch p. 530 |
1991 | Perissodactyla Nowak |
1997 | Selenida McKenna and Bell |
1999 | Perissodactyla Waddell et al. p. 3 |
2001 | Perissodactyla Madsen et al. p. 610 |
2001 | Perissodactyla Waddell et al. p. 148 |
2005 | Perissodactyla Prothero |
2005 | Perissodactyla Wilson and Reeder |
2006 | Perissodactyla Hulbert and Whitmore p. 13 |
2006 | Perissodactyla Symeonidis et al. |
2009 | Perissodactyla MacFadden |
2010 | Perissodactyla Asher and Helgen p. 3 figs. Figure 1 |
2014 | Perissodactyla Cooper et al. p. 3 fig. 3 |
2014 | Perissodactyla Ferrusquía-Villafranca et al. p. 204 |
2017 | Perissodactyla Bai et al. |
2020 | Perissodactyla Rose et al. p. 101 fig. 81 |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Or. Perissodactyla Owen 1848 [odd-toed ungulate]
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Fm. †Anthracobunidae Wells and Gingerich 1983
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G. †Anthracobune Pilgrim 1940
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†Anthracobune aijiensis Kumar 1991
†Anthracobune daviesi Pilgrim 1940
†Anthracobune pinfoldi Pilgrim 1940
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Invalid names: Pilgrimella pilgrimi Dehm and Oettingen-Spielberg 1958 [synonym]
†Anthracobune wardi Pilgrim 1940
G. †Lammidhania Gingerich 1977
G. †Pilgrimella Dehm and Oettingen-Spielberg 1958
Fm. †Brontotheriidae Marsh 1873 [brontothere]
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Subfm. †Brontotheriinae Marsh 1873
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Tr. †Brontotheriini Marsh 1873
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Subtr. †Brontotheriina Marsh 1873
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Unr. †Brontotheriita Marsh 1873
G. †Diplacodon Marsh 1875
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Invalid names: Eotitanotherium Peterson 1914 [synonym], Pseudodiplacodon Mader 2000 [synonym]
G. †Epimanteoceras Granger and Gregory 1943
G. †Pachytitan Granger and Gregory 1943
G. †Protitan Granger and Gregory 1943
G. †Rhinotitan Granger and Gregory 1943
G. †Microtitan Granger and Gregory 1943
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†Microtitan mongoliensis Osborn 1925
Invalid names: Microtitan elongatus Qi 1987 [nomen dubium]
G. †Nanotitanops Qi and Beard 1998
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†Nanotitanops shanghuangensis Qi and Beard 1998
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Invalid names: Nanotitan shanghuangensis Qi and Beard 1996 [replaced]
Invalid names: Nanotitan Qi and Beard 1996 [synonym]
Subtr. †Telmatheriina Osborn 1914
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G. †Qufutitan Wang and Wang 1997
G. †Wickia Mihlbachler 2008
Invalid names: Metatelmatheriinae Granger and Gregory 1943 [synonym]
Tr. †Dolichorhinini Riggs 1912
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G. †Sphenocoelus Osborn 1895
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†Sphenocoelus hyognathus Osborn 1889
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Invalid names: Dolichorhinus longiceps Douglass 1909 [synonym], Sphenocoelus cornutum Osborn 1895 [synonym]
†Sphenocoelus intermedius Osborn 1908
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Invalid names: Dolichorhinus fluminalis Riggs 1912 [synonym], Dolichorhinus heterodon Douglass 1909 [synonym], Mesatirhinus superior Riggs 1912 [synonym]
†Sphenocoelus uintensis Osborn 1895
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Invalid names: Sphenocoelus blairi Cook 1926 [synonym], Sphenocoelus bridgeri Cook 1926 [synonym], Sphenocoelus harundivoras Cook 1926 [synonym]
Invalid names: Dolichorhinoides Granger and Gregory 1943 [synonym], Dolichorhinus Hatcher 1895 [synonym], Tanyorhinus Cook 1926 [synonym]
Invalid names: Rhadinorhininae Osborn 1929 [synonym]
Unr. †Embolotheriita Osborn 1929
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G. †Brachydiastematherium Bockh and Maty 1876
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†Brachydiastematherium transylvanicum Bockh and Maty 1876
G. †Embolotherium Osborn 1929
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†Embolotherium andrewsi Osborn 1929
†Embolotherium grangeri Osborn 1929
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Invalid names: Embolotherium insigne Yanovskaya 1980 [synonym], Embolotherium loucksii Osborn 1929 [synonym], Titanodectes ingens Granger and Gregory 1943 [synonym], Titanodectes minor Granger and Gregory 1943 [synonym]
†Embolotherium ultimum Granger and Gregory 1943
G. †Metatitan Granger and Gregory 1943
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†Metatitan khaitshinus Yanovskaya 1954
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Invalid names: Metatitan reshetovi Yanovskaya 1980 [synonym]
†Metatitan primus Granger and Gregory 1943
†Metatitan relictus Granger and Gregory 1943
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Invalid names: Protitan khaitshinus Yanovskaya 1980 [synonym]
G. †Metarhinus Osborn 1908
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†Metarhinus abbotti Riggs 1912
†Metarhinus pater Stock 1937
Invalid names: Heterotitanops Peterson 1914 [synonym], Metarhinus cristatus Riggs 1912 [nomen dubium], Metarhinus earlei Osborn 1908 [nomen dubium], Rhadinorhinus Riggs 1912 [synonym], Telmatherium accola Cook 1926 [nomen dubium], Telmatherium advocata Cook 1926 [nomen dubium]
G. †Protitanotherium Osborn 1908
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†Protitanotherium emarginatum Hatcher 1895
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Invalid names: Protitanotherium superbum Osborn 1908 [synonym], Sthenodectes australis Wilson 1977 [synonym]
Invalid names: Protitanotherium koreanicum Takai 1939 [nomen dubium]
G. †Telmatherium Marsh 1872
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†Telmatherium megarhinum Earle 1891
†Telmatherium validum Marsh 1872
†Telmatherium validus Marsh 1872
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Invalid names: Telmatherium cultridens Osborn et al. 1878 [synonym], Telmatherium manteoceras Osborn in Matthew 1899 [synonym], Telmatherium washakiensis Osborn 1908 [synonym]
Invalid names: Leurocephalus Osborn et al. 1878 [synonym], Manteoceras Hatcher 1895 [synonym], Telmatotherium Marsh 1872 [synonym]
Subfm. †Eotitanopinae Osborn 1913
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G. †Eotitanops Osborn 1907
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†Eotitanops borealis Cope 1880
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Invalid names: Eotitanops brownianus Cope 1881 [synonym], Eotitanops major Osborn 1913 [synonym], Eotitanops princeps Osborn 1913 [synonym]
†Eotitanops dayi Dehm and Oettingen-Spielberg 1958
†Eotitanops pakistanensis Missiaen et al. 2011
G. †Lambdotherium Cope 1880
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†Lambdotherium popoagicum Cope 1880
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Invalid names: Lambdotherium magnum Osborn 1919 [synonym], Lambdotherium primaevum Loomis 1907 [synonym], Lambdotherium priscum Osborn 1913 [synonym], Lambdotherium progressum Osborn 1913 [synonym]
G. †Metatelmatherium Granger and Gregory 1938
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†Metatelmatherium cristatum Granger and Gregory 1938
†Metatelmatherium ultimum Osborn 1908
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Invalid names: Manteoceras uintensis Douglass 1909 [synonym]
Invalid names: Metatelmatherium lahirii Pilgrim 1925 [nomen dubium], Metatelmatherium parvum Granger and Gregory 1943 [nomen dubium]
G. †Mulkrajanops Kumar and Sahni 1985
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†Mulkrajanops moghliensis Kumar and Sahni 1985
†Mulkrajanops mogliensis Kumar and Sahni 1985
Subfm. †Palaeosyopinae Steinmann and Döderlein 1890
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G. †Palaeosyops Leidy 1870
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†Palaeosyops junius Leidy 1873
†Palaeosyops manteoceras Osborn in Matthew 1899
†Palaeosyops paludosus Leidy 1870
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Invalid names: Canis marshii Hay 1899 [synonym], Limnohyops matthewi Osborn 1908 [synonym], Palaeosyops humilis Leidy 1872 [synonym], Palaeosyops laevidens Cope 1873 [synonym], Palaeosyops longirostris Earle 1892 [synonym], Palaeosyops major Leidy 1871 [synonym], Palaeosyops minor Marsh 1871 [synonym], Palaeosyops minor Earle 1891 [synonym], Palaeosyops priscus Osborn 1908 [synonym]
†Palaeosyops robustus Marsh 1872
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Invalid names: Palaeosyops copei Osborn 1908 [synonym], Palaeosyops diaconus Cope 1873 [synonym], Palaeosyops grangeri Osborn 1908 [synonym], Palaeosyops laticeps Leidy 1872 [synonym], Palaeosyops leidyi Osborn 1908 [synonym]
Invalid names: Eometarhinus Osborn 1919 [synonym], Limnohyops Marsh 1890 [synonym], Limnohyus Marsh 1872 [synonym], Palaeosyops ultimus Matthew 1897 [nomen nudum]
G. †Sivatitanops Pilgrim 1925
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†Sivatitanops cotteri Pilgrim 1925
Invalid names: Sivatitanops birmanicum Pilgrim and Cotter 1916 [nomen dubium]
G. †Sthenodectes Gregory 1912
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Invalid names: Brontopinae Osborn 1914 [invalid subgroup], Epimanteoceratinae Granger and Gregory 1943 [empty], Menodontidae Cope 1881 [invalid subgroup], Titanotheriidae [synonym]
G. †Cambaytherium Bajpai et al. 2005
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†Cambaytherium gracilis Smith et al. 2016
†Cambaytherium marinus Bajpai et al. 2006
†Cambaytherium thewissi Bajpai et al. 2005
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Invalid names: Cambaytherium bidens Bajpai et al. 2005 [synonym], Cambaytherium minor Bajpai et al. 2005 [synonym], Indobune vastanensis Rose et al. 2006 [synonym]
Invalid names: Indobune Rose et al. 2006 [synonym]
Or. †Desmostylia Reinhart 1953 [desmostylian]
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G. †Behemotops Domning et al. 1986
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†Behemotops katsuiei Inuzuka 2000
†Behemotops proteus Domning et al. 1986
Or. †Desmostyloidea Abel 1933
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Fm. †Desmostylidae Osborn 1905
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G. †Desmostylus Marsh 1888
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†Desmostylus hesperus Marsh 1888
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Invalid names: Desmostylella typica Nagao 1937 [synonym], Desmostylus californicus Hay 1923 [synonym], Desmostylus cymatias Hannibal 1922 [synonym], Desmostylus minor Nagao 1937 [synonym], Desmostylus mirabilis Nagao 1935 [synonym], Desmostylus watasei Hay 1915 [synonym]
†Desmostylus japonicus Tokunaga and Iwasaki 1914
Invalid names: Desmostylella Nagao 1937 [synonym]
Fm. †Paleoparadoxiidae Reinhart 1959
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G. †Neoparadoxia Barnes 2013
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†Neoparadoxia cecilialina Barnes 2013
†Neoparadoxia repenningi Domning and Barnes 2007
G. †Paleoparadoxia Reinhart 1959
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†Paleoparadoxia media Inuzuka 2005
†Paleoparadoxia tabatai Tokunaga 1939
Invalid names: Behemotopsinae Inuzuka 1987 [empty], Paleoparadoxiinae Reinhart 1959 [empty]
G. †Jamilcotatus Aranda-Manteca and Barnes 1998
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†Jamilcotatus boreios Aranda-Manteca and Barnes 1998
†Jamilcotatus kumiaica Aranda-Manteca and Barnes 1998
Invalid names: Cornwalliidae Shikama 1966 [empty], Desmostyliformes Hay 1923 [synonym]
Superfm. Equoidea Hay 1902
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G. †Cymbalophus Hooker 1980
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†Cymbalophus cuniculus Owen 1842
†Cymbalophus hookeri Godinot et al. 1987
Fm. Equidae Gray 1821 [horse]
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Subfm. †Anchitheriinae Leidy 1869 [anchitheriine horse]
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G. †Anchitherium von Meyer 1844
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Subg. †Anchitherium (Hypohippus) Leidy 1858
Subg. †Anchitherium (Parahippus) Leidy 1858
†Anchitherium aurelianense Cuvier 1825
†Anchitherium clarencei Simpson 1932
†Anchitherium hypohippoides Matsumoto 1924
†Anchitherium navasotae Hay 1924
†Anchitherium zitteli Schlosser 1903
G. †Archaeohippus Gidley 1906
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†Archaeohippus blackbergi Hay 1924
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Invalid names: Archaeohippus nanus Simpson 1932 [synonym], Parahippus minutalis Hay 1924 [synonym]
†Archaeohippus mannulus O'Sullivan 2003
†Archaeohippus minimus Douglass 1899
†Archaeohippus mourningi Merriam 1913
†Archaeohippus penultimus Matthew 1924
†Archaeohippus stenolophus Lambe 1905
†Archaeohippus ultimus Cope 1886
G. †Desmatippus Scott 1893
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†Desmatippus crenidens Scott 1893
†Desmatippus integer Matthew 1924
†Desmatippus nebrascensis Peterson 1907
†Desmatippus texanus Leidy 1868
†Desmatippus tyleri Loomis 1908
Invalid names: Anchippus Leidy 1868 [synonym]
G. †Hypohippus Leidy 1858
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†Hypohippus affinis Leidy 1858
†Hypohippus chico MacFadden 1982
†Hypohippus equinus Scott 1893
†Hypohippus nevadensis Merriam 1913
†Hypohippus osborni Gidley 1907
†Hypohippus pertinax Matthew 1918
Invalid names: Hypohippus (Drymohippus) Merriam 1913 [nomen dubium]
G. †Kalobatippus Osborn 1915
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†Kalobatippus agatensis Osborn 1918
†Kalobatippus australis Leidy 1873
†Kalobatippus avus Schlaikjer 1935
G. †Megahippus McGrew 1938
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†Megahippus matthewi Barbour 1914
†Megahippus mckennai Tedford and Alf 1962
G. †Mesohippus Marsh 1875
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†Mesohippus bairdi Leidy 1850
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Invalid names: Mesohippus hypostylus Osborn 1904 [synonym], Mesohippus proteulophus Osborn 1904 [synonym], Mesohippus viejensis Clark and Beerbower 1967 [synonym]
†Mesohippus barbouri Schlaikjer 1931
†Mesohippus celer Marsh 1874
†Mesohippus latidens Douglass 1903
†Mesohippus propinquus Lambe 1905
†Mesohippus texanus McGrew 1971
†Mesohippus westoni Cope 1889
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Invalid names: Mesohippus montanensis Osborn 1904 [synonym], Mesohippus praecocidens Lambe 1905 [synonym]
Invalid names: Mesohippus cuneatus Cope 1873 [nomen dubium]
G. †Miohippus Marsh 1874
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†Miohippus anceps Marsh 1874
†Miohippus condoni Leidy 1870
†Miohippus equiceps Cope 1878
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Invalid names: Anchitherium brachylophum Cope 1878 [synonym], Mesohippus acutidens Sinclair 1905 [synonym]
†Miohippus equinanus Osborn 1918
†Miohippus gemmarosae Osborn 1918
†Miohippus gidleyi Osborn 1904
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Invalid names: Mesohippus grallipes Sinclair 1925 [synonym], Miohippus validus Osborn 1904 [synonym]
†Miohippus grandis Clark and Beerbower 1967
†Miohippus intermedius Osborn and Wortman 1895
†Miohippus obliquidens Osborn 1904
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Invalid names: Mesohippus brachystylus Osborn 1904 [synonym], Mesohippus eulophus Osborn 1904 [synonym], Mesohippus meteulophus Osborn 1904 [synonym], Pediohippus antiquus Schlaikjer 1935 [synonym]
†Miohippus primus Osborn 1918
†Miohippus quartus Osborn 1918
†Miohippus taxus Douglass 1908
Invalid names: Altippus Douglass 1908 [synonym], Anchitherium longicristis Cope 1878 [nomen dubium], Pediohippus Schlaikjer 1935 [synonym]
G. †Parahippus Leidy 1858
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†Parahippus agrestis Leidy 1873
†Parahippus atavus Osborn 1918
†Parahippus cognatus Leidy 1858
†Parahippus coloradensis Gidley 1907
†Parahippus leonensis Sellards 1916
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Invalid names: Merychippus socius Hay 1924 [synonym], Parahippus barbouri White 1942 [synonym], Parahippus vellicans Hay 1924 [synonym]
†Parahippus maxsoni Jahns 1940
†Parahippus pawniensis Gidley 1907
†Parahippus pristinus Osborn 1918
†Parahippus wyomingensis Schlaikjer 1935
Invalid names: Anchitherium australe Leidy 1873 [nomen dubium]
G. †Eohippus Marsh 1876
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†Eohippus angustidens Cope 1875
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Invalid names: Eohippus validus Marsh 1876 [synonym], Hyracotherium cuspidatum Cope 1875 [synonym], Hyracotherium seekinsi Morris 1968 [synonym]
†Eohippus borealis Granger 1908
†Eohippus cristonensis Cope 1877
†Eohippus resartus Granger 1908
G. †Epihippus Marsh 1878
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Subg. †Epihippus (Duchesnehippus) Peterson 1932
Subg. †Epihippus (Epihippus) Stirton 1940
†Epihippus uintensis Marsh 1875
Subfm. Equinae Steinmann and Döderlein 1890
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G. †Acritohippus Kelly 1995
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†Acritohippus quinni Kelly 1995
†Acritohippus stylodontus Merriam 1919
†Acritohippus tertius Osborn 1918
Tr. Equini Quinn 1955 [horse]
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G. †Allohippus Kretzoi 1938
G. †Boreohippidion Dos Santos Avilla et al. 2015
G. †Calippus Matthew and Stirton 1930
G. †Dinohippus Quinn 1955
G. Equus Linnaeus 1758 [horse]
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Invalid names: Allozebra Trumler 1961 [synonym], Asinus Gray 1822 [synonym], Dolichohippus Heller 1912 [synonym], Haringtonhippus Heintzman et al. 2017 [synonym], Hippotigris Hamilton-Smith 1841 [synonym], Kolpohippus van Hoepen 1930 [synonym], Kraterohippus van Hoepen 1930 [synonym], Neohippus Abel 1913 [synonym], Onager Brisson 1762 [synonym], Sterrohippus van Hopen 1930 [synonym], Tomolabis Cope 1892 [synonym]
G. †Heteropliohippus Kelly 1995
G. †Hippidion Owen 1869
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Invalid names: Onohippidium Moreno 1891 [synonym], Parahipparion Ameghino 1904 [synonym]
G. †Parapliohippus Kelly 1995
G. †Plesippus Matthew 1924
Subtr. Pliohippina Prado and Alberdi 1996
G. †Pliohippus Marsh 1874
Invalid names: Equus proversus Merriam 1916 [nomen dubium], Protohippina Quinn 1955 [empty]
Tr. †Hipparionini Quinn 1955 [hipparionine horse]
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G. †Cormohipparion Skinner and MacFadden 1977
G. †Hipparion de Christol 1832
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Invalid names: Hemihipparion Wehrli 1941 [synonym], Hipparitherium Christol 1847 [synonym], Stylohipparion van Hoepen 1932 [synonym]
G. †Hippotherium Kaup 1832
G. †Nannippus Matthew 1926
Invalid names: Hipparion anthonyi Merriam 1916 [nomen dubium], Hippotherium sinclairi Wortman 1882 [nomen dubium]
G. †Merychippus Leidy 1856
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Subg. †Merychippus (Merychippus) Stirton 1940
Subg. †Merychippus (Protohippus) Leidy 1858
†Merychippus brevidontus Bode 1935
†Merychippus calamarius Cope 1875
†Merychippus californicus Merriam 1915
†Merychippus eohipparion Osborn 1918
†Merychippus eoplacidus Osborn 1918
†Merychippus gunteri Simpson 1930
†Merychippus labrosus Cope 1874
†Merychippus missouriensis Douglass 1908
†Merychippus patruus Osborn 1918
†Merychippus primus Osborn 1918
†Merychippus proparvulus Osborn 1918
†Merychippus quintus Osborn 1918
†Merychippus relictus Cope 1889
†Merychippus republicanus Osborn 1918
†Merychippus sejunctus Cope 1874
†Merychippus seversus Cope 1878
†Merychippus stevensi Dougherty 1940
Invalid names: Merychippus isonesus secundus Osborn 1918 [nomen vanum], Merychippus quartus Osborn 1918 [nomen vanum]
Invalid names: Calippini Quinn 1955 [empty], Hippodon Leidy 1854 [invalid subgroup], Hippotheriini Bonaparte 1850 [empty], Protohippini Quinn 1955 [empty], Stylonus Cope 1878 [nomen dubium]
G. †Gobihippus Dashzeveg 1979
G. †Hyperhippidium Sefve 1910
G. †Hypohippidium Fernandez de Alvarez 1957
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†Hypohippidium humahuaquense Fernandez de Alvarez 1957
G. †Lophiopus Ellenberger 1980
G. †Minippus Froehlich 2002
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†Minippus index Cope 1873
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Invalid names: Hyracotherium sulcatus Gingerich 1989 [synonym], Orotherium loevii Cope 1877 [synonym]
†Minippus jicarillai Froehlich 2002
G. †Orohippus Marsh 1872
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†Orohippus agilis Marsh 1873
†Orohippus major Marsh 1874
†Orohippus progressus Granger 1908
†Orohippus proteros Korth and Evander 1982
†Orohippus pumilus Marsh 1871
+
Invalid names: Helotherium procyoninum Cope 1871 [synonym], Hyracotherium osbornianum Cope 1883 [synonym], Lophiotherium ballardi Marsh 1871 [synonym], Oligotomus cinctus Cope 1873 [synonym], Orohippus atavus Granger 1908 [synonym], Orohippus typicus Granger 1908 [synonym]
Invalid names: Helohippus Marsh 1892 [synonym], Helotherium Cope 1872 [synonym], Oligotomus Cope 1873 [synonym], Orohippus (Aminippus) Granger 1908 [invalid subgroup]
G. †Plesiohipparion Qiu et al. 1987
G. †Pliolophus Owen 1858
+
†Pliolophus barnesi Hooker 2010
†Pliolophus quesnoyensis Bronnert et al. 2017
†Pliolophus vulpiceps Owen 1858
G. †Protorohippus Wortman 1896
+
†Protorohippus montanum Wortman 1896
†Protorohippus venticolum Cope 1881
G. †Sifrhippus Froehlich 2002
+
†Sifrhippus grangeri Gingerich 1989
†Sifrhippus sandrae Gingerich 1989
Invalid names: Arenahippus Froehlich 2002 [synonym]
G. †Xenicohippus Bown and Kihm 1981
+
†Xenicohippus craspedotum Cope 1880
†Xenicohippus grangeri Bown and Kihm 1981
†Xenicohippus osborni Bown and Kihm 1981
Invalid names: Protohippinae Gidley 1907 [empty]
Fm. †Palaeotheriidae Bonaparte 1850
+
G. †Cantabrotherium Casanova-Cladellas and Santafe-Llopis 1987
+
†Cantabrotherium casanovasae Cuesta 1993
†Cantabrotherium truyolsi Casanovas and Santafe 1987
G. †Franzenium Casanovas and Santafe 1989
+
†Franzenium durense Cuesta 1993
†Franzenium tetradactylum Casanovas and Santafe 1989
G. †Hyracotherium Owen 1841
+
†Hyracotherium leporinum Owen 1841
†Hyracotherium levei Hooker 1994
†Hyracotherium remyi Checa-Soler 1997
†Hyracotherium tapirinum Cope 1875
G. †Iberolophus Badiola and Cuesta 2008
+
†Iberolophus arabensis Badiola and Cuesta 2008
†Iberolophus jimenezi Badiola and Cuesta 2008
G. †Leptolophus Remy 1965
+
†Leptolophus cuestai Perales-Gogenola et al. 2021
†Leptolophus franzeni Perales-Gogenola et al. 2021
†Leptolophus magnus Remy 1998
†Leptolophus nouleti Remy 1998
†Leptolophus stehlini Remy 1965
G. †Orolophus Remy 2017
+
†Orolophus maldani Remy 2017
Invalid names: Orotherium Marsh 1872 [replaced]
Subfm. †Pachynolophinae Paulow 1877
+
G. †Lophiotherium Marsh 1871
+
†Lophiotherium cervulum Gervais 1849
†Lophiotherium geiseltalensis Matthes 1977
†Lophiotherium robiacense Deperet 1917
†Lophiotherium siderolithicum Pictet 1857
G. †Metanchilophus Remy 2012
+
†Metanchilophus castrensis Remy 2012
†Metanchilophus chaubeti Remy 2012
†Metanchilophus depereti Stehlin 1905
†Metanchilophus dumasi Gervais 1849
†Metanchilophus gaudini Pictet and Humbert 1869
†Metanchilophus radegondensis Gervais 1852
G. †Pachynolophus Pomel 1847
+
†Pachynolophus boixedatensis Crusafont Pairo and Remy 1970
†Pachynolophus bretovensis Remy 1988
†Pachynolophus cayluxi Filhol 1888
†Pachynolophus cesserasicus Gervais 1849
†Pachynolophus duvali Pomel 1847
†Pachynolophus eulaliensis Danilo et al. 2013
†Pachynolophus garimondi Remy 1967
†Pachynolophus gaytei Remy 2015
†Pachynolophus hookeri Godinot et al. 1987
†Pachynolophus lavocati Remy 1972
†Pachynolophus livinierensis Savage et al. 1965
†Pachynolophus molipontiensis Checa-Soler 1997
†Pachynolophus ruscassierensis Remy et al. 2019
†Pachynolophus zambranensis Badiola et al. 2005
G. †Propalaeotherium Gervais
+
†Propalaeotherium argentonicum Gervais 1853
†Propalaeotherium gaudryi Lemoine 1878
†Propalaeotherium helveticum Savage et al. 1965
†Propalaeotherium rollinati Stehlin 1905
†Propalaeotherium sinense Zdansky 1930
†Propalaeotherium sudrei Remy et al. 2016
†Propalaeotherium voigti Matthes 1977
Subfm. †Palaeotheriinae Bonaparte 1850
+
G. †Palaeotherium Cuvier 1804
+
†Palaeotherium cartieri Stehlin 1904
†Palaeotherium castrense Noulet 1863
†Palaeotherium crassum Cuvier 1805
†Palaeotherium crusafonti Cassanovas-Cladellas 1975
†Palaeotherium curtum Cuvier 1812
†Palaeotherium duvali Pomel 1853
†Palaeotherium eocaenum Gervais 1875
†Palaeotherium giganteum Cuesta 1993
†Palaeotherium isselanum Cuvier 1824
†Palaeotherium lautricense Stehlin 1904
†Palaeotherium magnum Cuvier 1804
†Palaeotherium maximum Leidy 1852
†Palaeotherium medium Cuvier 1804
†Palaeotherium minus Cuvier 1804
†Palaeotherium muehlbergi Stehlin 1904
†Palaeotherium pomeli Franzen 1968
†Palaeotherium renevieri Stehlin 1904
†Palaeotherium robustum Franzen 1968
†Palaeotherium ruetimeyeri Franzen 1968
†Palaeotherium siderolithicum Pictet and Humbert 1869
†Palaeotherium tapiroides Cuvier 1812
G. †Plagiolophus Pomel 1847
+
Subg. †Plagiolophus (Plagiolophus) Pomel 1847
†Plagiolophus annectens Owen 1847
†Plagiolophus cartailhaci Stehlin 1904
†Plagiolophus codiciensis Gaudry 1865
†Plagiolophus curtisi Hooker 1986
†Plagiolophus mamertensis Remy 2004
†Plagiolophus markgrafi Lőrenthey 1907
†Plagiolophus mazateronensis Cuesta 1994
†Plagiolophus ministri Brunet and Jehenne 1989
†Plagiolophus minor Cuvier 1804
G. †Paraplagiolophus Depéret 1917
G. †Propachynolophus Lemoine 1891
+
†Propachynolophus duvali Pomel 1853
†Propachynolophus maldani Remy 2017
G. †Pseudopalaeotherium Franzen 1972
G. †Ghazijhippus Missiaen and Gingerich 2014
+
†Ghazijhippus talibhasani Missiaen and Gingerich 2014
G. †Hallensia Franzen and Haubold 1986
+
†Hallensia louisi Hooker 1994
†Hallensia matthesi Franzen and Haubold 1986
Subor. †Lophodontomorpha Hooker and Dashzeveg 2004
+
Infraor. †Euperissodactyla Hooker and Dashzeveg 2004
+
G. †Mesolambdolophus Holbrook and Lapergola 2011
+
†Mesolambdolophus setoni Holbrook and Lapergola 2011
G. †Paschatherium Russell 1963
+
†Paschatherium dolloi Teilhard de Chardin 1927
†Paschatherium levei Hooker and Russell 2012
†Paschatherium marianae Estravís and Russell 1992
†Paschatherium plaziati Marandat 1989
†Paschatherium russelli Godinot 1978
†Paschatherium yvettae Gheerbrant et al. 1997
G. †Perissobune Missiaen and Gingerich 2014
+
†Perissobune intizarkhani Missiaen and Gingerich 2014
†Perissobune munirulhaqi Missiaen and Gingerich 2014
Subor. †Solidungula Cuvier
+
Unr. Tapiromorpha Haeckel 1873
+
Unr. †Ancylopoda Cope 1889
+
Superfm. †Chalicotherioidea Gill 1872
+
Fm. †Chalicotheriidae Gill 1872 [chalicothere]
+
Subfm. †Chalicotheriinae
G. †Eomoropus Osborn 1913
G. †Limognitherium Filhol 1880
Subfm. †Macrotheriinae Holland and Peterson 1913
G. †Moropus Marsh 1877
G. †Pernatherium Gervais 1876
Subfm. †Schizotheriinae Holland and Peterson 1913
G. †Schizotherium Gervais 1876
Invalid names: Moropodinae Marsh 1877 [empty]
Fm. †Lophiodontidae Gill 1872
+
G. †Atalonodon Dal-Piaz 1929
G. †Lophiaspis Depéret 1910
G. †Lophiodochoerus Lemoine 1880
Subfm. †Lophiodontinae Cope 1881
G. †Paleomoropus Radinsky 1964
G. †Protomoropus Hooker and Dashzeveg 2004
+
†Protomoropus gabuniai Hooker and Dashzeveg 2004
†Protomoropus hengyangensis Young 1944
Invalid names: Artionychia Osborn and Wortman 1893 [empty], Lophiodontoidea [empty], Perissonychia Osborn and Wortman 1893 [empty]
G. †Cardiolophus Gingerich 1991
+
†Cardiolophus radinskyi Gingerich 1991
+
Invalid names: Homogalax planeticus Gingerich 1980 [synonym]
†Cardiolophus semihians Cope 1882
Unr. †Ceratomorpha Wood 1937
+
G. †Parabreviodon Reshetov 1975
Superfm. Rhinocerotoidea Gray 1821
+
Fm. †Amynodontidae Scott and Osborn 1883
+
G. †Amynodon Marsh 1877
G. †Amynodontopsis Stock 1933
G. †Andarakodon Averianov and Potapova 1996
Tr. †Cadurcodontini
Tr. †Metamynodontini Kretzoi 1942
G. †Orthocynodon Scott and Osborn 1882
G. †Rostriamynodon Wall and Manning 1986
Invalid names: Amynodontinae Scott and Osborn 1883 [empty], Cadurcotheriinae Kretzoi 1942 [invalid subgroup], Gigantamynodon Gromova 1954 [nomen dubium], Gigantamynodon cessator Gromova 1954 [nomen dubium], Gigantamynodon giganteus Xu 1961 [nomen dubium], Gigantamynodon maguanensis Qi 1992 [nomen dubium], Paramynodontinae Kretzoi 1942 [invalid subgroup], Rostriamynodontinae Wall and Manning 1986 [empty]
G. †Caenolophus Matthew and Granger 1925
+
†Caenolophus magnus Shi 1989
†Caenolophus obliquus Matthew and Granger 1925
†Caenolophus progressus Matthew and Granger 1925
†Caenolophus promissus Matthew and Granger 1925
†Caenolophus suprametalophus Shi 1989
Invalid names: Euryodon Xu et al. 1979 [synonym], Teilhardia Prothero and Schoch 1989 [synonym], Teilhardina Prothero and Schoch 1989 [synonym]
Fm. †Eggysodontidae Breuning 1923
+
G. †Allacerops Wood 1932
G. †Proeggysodon Bai and Wang 2012
Invalid names: Eggysodontinae Breuning 1923 [empty]
Fm. †Hyrachyidae Osborn 1892
+
G. †Hyrachyus Leidy 1871
+
Invalid names: Chasmotherium Rutimeyer [synonym], Colonoceras Marsh 1873 [synonym], Ephyrachyus Wood 1934 [synonym], Metahyrachyus Troxell 1922 [synonym]
G. †Subhyrachyus Gabunia 1999
Invalid names: Hyrachyus douglassi Wood 1934 [nomen dubium]
Fm. †Hyracodontidae Cope 1879
+
Subfm. †Alloceropinae Wood 1932
G. †Ardynia Matthew and Granger 1923
G. †Epitriplopus Wood 1927
G. †Ilianodon Chow 1961
G. †Pataecops Radinsky 1966
G. †Praeaceratherium Abel 1910
G. †Prohyracodon Koch 1897
+
Invalid names: Meninatherium Abel 1910 [synonym], Prothyracodon Scott and Osborn 1887 [synonym]
G. †Rhodopagus Radinsky 1965
G. †Triplopides Radinsky 1967
G. †Ulania Qi 1990
G. †Yimengia Wang 1988
Invalid names: Allaceropinae Wood 1932 [empty], Hyracodontinae Steinmann and Doderlein 1890 [empty], Triplopodinae Cope 1881 [empty]
Fm. †Paraceratheriidae Osborn 1923
+
Subfm. †Forstercooperiinae Wood 1963
Invalid names: Forstercooperia ergiliinensis Gabunia and Dashzeveg 1974 [nomen dubium], Forstercooperia jigniensis Sahni and Khare 1973 [nomen dubium]
G. †Punjabitherium Khan 1971
Fm. Rhinocerotidae Gray 1821 [rhinoceros]
+
Subfm. †Aceratheriinae Dollo 1885
Tr. †Aceratheriini Dollo 1885
G. †Amphicaenopus Wood 1927
G. †Atelodus Pomel 1853
Subfm. †Diceratheriinae Dollo 1885
Subtr. †Elasmotheriina Bonaparte 1850
G. †Gulfoceras Albright 1999
G. †Huangotherium
Subfm. †Iranotheriinae Kretzoi 1942
G. †Penetrigonias Tanner and Martin 1976
Subfm. Rhinocerotinae Gray 1821
G. †Subhyracodon Wood 1927
+
Invalid names: Anchisodon Cope 1879 [synonym], Caenopus Cope 1880 [synonym], Leptaceratherium Osborn 1898 [synonym]
G. †Teletaceras Hanson 1989
G. †Uintaceras Holbrook and Lucas 1997
G. †Woodoceras Prothero 2005
G. †Zalabis
Invalid names: Anchisodon tubifer Cope 1879 [nomen dubium], Dicerorhininae Simpson 1945 [empty], Forstercooperia crudus Gabounia 1977 [nomen dubium], Gigantamynodon akespensis Bayshashov 1993 [nomen dubium], Rhinoceros crassus Leidy 1858 [nomen dubium], Rhinoceros oregonensis Marsh 1873 [nomen dubium], Rhinoceros pacificus Leidy 1872 [nomen dubium], Teleoceratinae [empty]
G. †Toxotherium Wood 1961
+
†Toxotherium hunteri Wood 1961
+
Invalid names: Schizotheriodes jackwilsoni Schiebout 1977 [synonym], Toxotherium woodi Skinner and Gooris 1966 [synonym]
Invalid names: Huananodon You 1977 [nomen dubium]
Superfm. Tapiroidea Gill 1872
+
Fm. †Deperetellidae Radinsky 1965
+
G. †Bahinolophus Tsubamoto 2005
G. †Deperetella Matthew and Granger 1925
+
Invalid names: Cristidentinus Zdansky 1930 [synonym], Diplolophodon Zdansky 1930 [synonym]
G. †Irdinolophus Dashzeveg and Hooker 1997
G. †Euryletes
G. †Haagella Heissig 1978
G. †Helaletes Marsh 1872
+
†Helaletes intermedius Osborn et al. 1878
+
Invalid names: Desmatotherium guyotii Scott 1883 [synonym]
†Helaletes medius Qi 1987
†Helaletes mongoliensis Osborn 1923
Fm. †Helaletidae Osborn 1892
+
G. †Jhagirilophus Thewissen et al. 2001
G. †Paracolodon Matthew and Granger 1925
G. †Vastanolophus Smith et al. 2015
Invalid names: Chasmotheriinae Viret 1858 [empty], Helaletinae Osborn 1892 [empty]
G. †Heptodon Cope 1882
+
†Heptodon minimus Qi 1987
†Heptodon niushanensis Chow and Li 1965
†Heptodon tienshanensis Zhai 1978
G. †Hesperaletes Colbert 2006
+
†Hesperaletes borineyi Colbert 2006
†Hesperaletes walshi Colbert 2006
Fm. Tapiridae Gray 1821 [tapir]
+
G. †Dilophodon Scott 1883
G. †Eotapirus Cerdeno and Ginsburg 1988
G. †Heteraletes Peterson 1931
G. †Miotapirus Schlaikjer 1937
G. †Nexuotapirus Albright 1998
G. †Palaeotapirus Filhol 1888
G. †Paratapirus Depéret and Douxami 1902
G. †Plesiocolopirus Schoch 1989
G. †Plesiotapirus Qiu et al. 1991
G. †Tapiravus Marsh 1877
Subfm. Tapirinae Burnett 1830
G. †Tapiriscus Kretzoi 1951
G. Tapirus Brisson 1762 [tapir]
+
Invalid names: Acrocodia Goldman 1913 [synonym], Cinchacus Gray 1873 [synonym], Pinchacus Hershkovitz 1954 [synonym], Rhinochoerus Wagler 1830 [synonym], Syspotamus Billberg 1827 [synonym], Tapir Blumenbach 1799 [synonym], Tapyra Liais 1872 [synonym]
Invalid names: Heptodontidae Holbrook 1999 [empty]
Invalid names: Deperetellidea Radinsky 1965 [empty], Lophialetidea Matthew and Granger 1925 [empty], Tapiridea Burnett 1830 [empty]
G. †Homogalax Hay 1899
+
†Homogalax aureus Gingerich 1991
†Homogalax namadicus Dashzeveg 1979
†Homogalax primaevus Wortman 1896
†Homogalax protapirinus Wortman 1896
†Homogalax wutuensis Chow and Li 1965
Fm. †Isectolophidae Peterson 1919
+
G. †Chowliia Tong and Wang 2006
+
†Chowliia europea Bronnert et al. 2019
†Chowliia laoshanensis Tong and Wang 2006
G. †Gandheralophus Missiaen and Gingerich 2012
+
†Gandheralophus minor Missiaen and Gingerich 2012
†Gandheralophus robustus Missiaen and Gingerich 2012
Invalid names: Homogalaxinae Peterson 1919 [empty], Systemodontinae Osborn and Wortman 1894 [invalid subgroup]
G. †Isectolophus Scott and Osborn 1887
+
†Isectolophus annectens Scott and Osborn 1887
+
Invalid names: Homogalax uintensis Troxell 1922 [synonym], Isectolophus scotti Peterson 1919 [synonym], Schizolophodon cuspidens Peterson 1919 [synonym]
†Isectolophus latidens Osborn et al. 1878
+
Invalid names: Homogalax bridgerensis Troxell 1922 [synonym], Homogalax reliquius Qi 1987 [synonym], Isectolophus bogdulensis Reshetov 1979 [synonym]
†Isectolophus radinskyi Schoch 1983
Invalid names: Parisectolophus Peterson 1919 [synonym]
G. †Kalakotia Ranga Rao 1972
+
†Kalakotia simplicidentata Ranga Rao 1972
+
Invalid names: Chasmotherium mckennai Sahni and Khare 1973 [synonym], Schlosseria radinskyi Sahni and Khare 1973 [synonym]
Invalid names: Aulaxolophus Ranga Rao 1972 [synonym]
Fm. †Lophialetidae Matthew and Granger 1925
+
G. †Eoletes Biryukov 1974
+
†Eoletes gracilis Biryukov 1974
†Eoletes tianshanicus Averianov and Godinot 2005
†Eoletes tshakpaktasensis Gabunia 1999
†Eoletes xianensis Zhang and Qi 1981
G. †Schlosseria Matthew and Granger 1926
+
†Schlosseria hetaoyuanensis Tong and Lei 1984
†Schlosseria magister Matthew and Granger 1926
+
Invalid names: Lophialetes primus Cheng and Ma 1990 [synonym], Simplaletes sujiensis Qi 1980 [synonym], Simplaletes ulanshirehensis Qi 1980 [synonym]
Invalid names: Simplaletes Qi 1980 [synonym]
Invalid names: Eoletidae Shoch 1989 [invalid subgroup]
G. †Systemodon Cope 1881
+
Invalid names: Breviodontidae [empty], Moropomorpha [synonym], Rhodopagidae [empty]
G. †Teilhardimys Kretzoi and Kretzoi 2000
+
†Teilhardimys brisswalteri Hooker and Russell 2012
†Teilhardimys musculus Kretzoi and Kretzoi 2000
+
Invalid names: Microhyus musculus Teilhard de Chardin 1927 [replaced]
†Teilhardimys reisi Antunes et al. 1987
Invalid names: Microhyus Teilhard de Chardin 1927 [replaced]
Subor. †Titanotheriomorpha
+
Invalid names: Chelodactyla Scott 1913 [empty], Equina Bonaparte 1850 [empty], Hippopotamus iravaticus Noetling 1897 [nomen dubium], Hyracotheriidae Cope 1881 [empty], Mesaxonia Marsh 1884 [empty], Rhinocerotida Owen 1845 [empty], Selenida McKenna and Bell 1997 [invalid subgroup], Stereopterna Ameghino 1889 [empty], Triplopodidae Cope 1881 [empty]
Diagnosis
No diagnoses are available